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DBhuiuI).andP、Yonzonz
姉10期プロ・ナトウーラ・ファンド助成旗来報(lril}(2001) SPECIESMAINTENANCEINDYNAMICLANDSCAPEOFTHE CI-IURlYA,EASTERNNEPAL DBhuiuI).andP、Yonzonz’ SUMMARY Th(pChuri>'allillsarcIh(pyoullgeslall(Isoutherllm()slmoul1tainchainoflhellimalaya・Theycon‐ ioinlwocc(〕Iogicalre8imes:IhelropicalinIhesoulhall(llcm1)eraleinlhenorIh」I1Nepal,lheChuriyaHills cover12.7()<,o(thet()mllalldareaandcollI「ibuIeto25、8〔X,o{11MJforestcovero(IhecounIry・A51hehuman encromdlmenIconIil1uestoiIlcrease,lhe(rt18ilelandsCapeo[Churi)'aisil1dallger・IIowever,ecolo8icaliI1‐ formaIi()11()、NCI)alClIuriyilisalIU1ostlloⅣcxisIelll・Thepl・〔psenlsIudycove】・in8IhcChuriyao(e(ISI(Prl1Nepal isalliniIiiIliollofcreMil18cc()I()8imldambas(PC(NepalChu「iyil,TllesIudydweⅡHonsixmajornreas:(i)GIS anal>'sis()(chan8eilllilI1dllsel)ilItcrllbetwcel11958al1dlリリ2,(ii)Horisliccompo5itiol1,forestst「llcIureand regell('『`lIi()、,(iii)trcespcCiesasHociationaIvilriousalIiIlldcs,(iv)loCalkn()wledgeonplalltuSesMIlollgthe residし、、l(DIhnicgroul〕s,(v)(Iislribuliono(blllI〔P『[lies,all。(vi)})msclillcin(()rmaliollonbirds. BACKGROUND ThcChuriy4lc()mpriseIhcyoull8esId1lldIhesouIhcrnnl〔)sIll1oulllminchain()(Ihellim(】1m)'t1I1system (BhatIl977,lla8enlO98)andcollioiIlIwodislil1ctecolo8iCalre8imes:Ihel「o})icalelwironso(Can8elicplail1s inIhcs()ulhalldlheIcmpemlell1()llllmillswl1ichexlendIolllcillpineinI1lenorth(I,i8.1).A1sokllowl1asthe Siwaliks,IlleChuri)'i1col】sisI()「T(Prliar>'1Inc()l1solidaIedilI1dhighl)'erodiblefIlwiaIilesedimcllIsmnging from1℃lMivel)'(ine-8raincdgmywackesilllhcsouIh(LowerSiwalik),Ihrou8hso(lofclay(MiddleSiwalik) lover>'〔()ilrsesmndsM1dcoll81(〕meraIes(U1)I)erSiwaIik)illIhellorlh(CarSonl985).TheChllriyaHills extelld(r〔)lulAfgallisI11allIo八ssamwiIhallnv〔pmgesumnliIoflcssIhanl500maSlalldthewidI1waryin8 from81o50km、 111cChuri>'arocksarcrichilwertcbrMc(ossilcolUIcnIs、Churi)'miHllnearlhedfossilso[laIcrTerIiar)’ havepr(widedmsis(()rmuch(〕fourpresellIknowIed8eo(Iheevoluliono(Asian(loraandfaunm(Ilihara(?! (】ノ.1972).Maior(ossil{allnaincllldesprimaIes,mrI1ivores,un8ulates,elel〕hnnls,rodcllIs,birds,rel〕Iilesand fishTlwl〕rimalecoⅡccIion(r()mtheChuri)'a((〕rmaIionsincludcsanumbero(gclleraofAnIhropoids・A compamIivestudyill〔licmlcslhill(heanimnls〔)IlhaIperi()dwcrelhcimmc(liMe41ncesIorsoflllepr“cI1t-day l)Ro)'alNcpalAcademyofScicnceandTcchI1olo8y,Cl)OBOX3323,Kalhmandu 2)RcsolIrccsllimnlaya,CPOl〕OX2`M8,KM11nlimdu -95‐ species、Fllrlhermore,IllcvarieIylllatexisIedIhenwas8reaIerIhanwhiltexisIsIoday. AN :<lミミご烹奄、 ●や■。●■■●o●● Fig.1.Physiogral)11iczonesofNcpal InNepal(are、:小17,181k,12),lheChuriyaHillscover12.7%olthcI()tallaI1dilremandcollIributeto 25.8%oflheforestcoveroflhecounI「>'(LRMl〕1978).Chllriym'shardwoodcomponcllI(S/10庇,(lrol)lイsmand Ter"li1lll1ilIs})p)islhehigheslinIhe(x)unlrywhichcomprises37%o(lh〔'1(北11(11M(iNlO88).Thcexisting proIeclednelworkdocs】1olencoml)assIheChu「iyaHillsbumtlcasMwoproIecledmremsinIhesouth,Cl1iIwall andBardia,arebordercdwilhIhelIills・AIsomep1acestheChuriyalIiⅡscouldals()n1akcbiolo8icalcorridorsuchasbeIwecnllKlrdiaall(IShlIklaPhanIainIhefar-wcslerllNepill・ TI1ereisa8rowilIBconcerllonmpiddeleriomtiol1o(thcChuri)'allillsillNc1〕31.AsIhehumallen‐ croachmenIconIil1llesIoillcredlse,Ihe(ra8ilelandscapeofChuriyaisindilI1Ber」lowever,ecolo8icalinfor‐ maIionollChuri)'i1llillsofNeI)i1lisnlmostllo汁existence,excepta((pwsIudicsc()nducIediIlsul)porto( developmcntacIivⅡics、Tllepres(pntsludy,collducledilleasIcrnNep(】1,c(wcrcdIh(pChuriyalliⅡsbctween MechiandSapmkoSlli.'「hesIudyincludedsixmajorareas,viz.(i)chnngcill1im〔I-llscpnucmill1958,l978 andl992,(ii)(Iorislicc()mposiIi()'1,(olPstslrucIureandre8clleraIioI1,(iii)lruesl〕ccicsi1ssociaIion,(iv)Iocal knowledgcollplantllse,(v)disIribllIionofbuIIcrHies,and(vi)baselill〔pin(ormaIi()、()nbirdspccics. -96- METHODOLOGY Studyarea ThestudyareaisapartofIhcChuriyaineastNepal(26.40,N-2753'N,87.08,E-88.11,E)exlcndin8105 kmfromthcMechiRiVer(Nepal,seasternborder)toIheSaptakoshiRivercoveringatotalareaof779.24 km2(widIh:500m‐20km;elevalioll:115-1300m)(Fig.2).Therewere390villa8es(settlemenls)under37 ViUa8eDevelopmentCommittecsofIhefivedisIricts:]I1apa(32.9kmz),Momng(211.3kmz)andSunsari (89.4km2)disIriclsandllam(443.9km2)andDhaI1kuta(L8km2)disIricIs. 926000 58か000 異O)00 鋤0○00 (⑪刺CO GcogmphicRe庇KelDccI》oinl8(SamplingSiにs)intlneChlIriya,EastNcpal CO-生一・--- ;トー'三コ■一十F’二丁'0二 ▲■_◆ -------◆■-=、=一一・・・-←--●--● ・‐--「‐◆ -------I申。-や 6 --十---.◆‐ 『 マサ---=-◆----- -f--÷---ヤー◆----- (AL」 ミ□」へ’ と ⅥⅡaOeB /V Road5 1. |_ P◆--.---千一 l-JL -,.------ ̄一一一一 屡gjA:Ls厩HpHQ> 、L--け ;鍵詳鵠討U薩劃辱片旨5冨禿、医… 両U7面頭-両毎万万万[T56刀一一一一一一 Y■6600■“」OETCpOObD仇fFUUM。A870>羽 F”、淀SIB■-UWD「F連P時HbdaHO8・■1,1粥9 1001020KⅡomotnr5 戸写再毛一一= ̄専一 92⑨0,0 540心①0 540000 :$0000 C(巾000 17i8.2.Geogmphicre(er(mcepoil1lsforsamp1ingillIheChuri>'aofeasterIlNepal. Samplingmethods l1orCIS(Ceo8raphiclll((〕rmaIiol1Systelll)allalysis,1111℃eIime-scriesmapsof1958(scalcl:63,360; Govt.o(Illdial961),1978(scalel:50,000;IRMP/IlMCN1986)andl992(scalel:25000;HMGN/PINNIDA 1996)weredigitizedugin8Arclllfo352andArcvicw32.Lan(l-usecategorieSwerca8riculture,forest,shrub, gmziI18,barren,andsimd/gravcl」、(ormatiollwerealsoassesscdforurbk1narea,swamp,tea/coffeepIallta‐ tion,orcllardandiI1(rasIructuresllChascanals・ Samplin8pIots(orfielddamcolleclionwereascermillcdusin88eogmphical8ridsonl992mal〕usm8 randomtablcofRohlfandSokal(1969).Inthe(ieldIhe8ridswerelocatedwiththehelpofGlobalPosition‐ in8System、Therewcrealto8elher35geogmphicreference}〕oints(samplill88rids).Fieldsurveyswcrecom‐ -97‐ pleIedbetweenJanuary‐April,2000. Ve8eIationsampIin8sweredonemquadrates,each8ridbasedsiIecomprisedoftwolargequadrates (25mx25m)represenIingtwodifferentslopeswhereappIicableForshrubstratum,eachsuchquadratehad twosub-quadmtesof5mx5m,alldfoursmallquadralesoflmxlmlor8roundve8etation・SampIingpa‐ rametersincludedheight,DBH,coverage,seedlingandsapIingdensityofpIantspeciesBeside,nolingsiIe condiIioll,soilsaml)leswerecollecIedfromlllestudyploIs、Rccordedplantswereidelllifiedatspecieslevel wherepossibleandspecimenwerecolIfirmedaIIheNationallIerbarium,Codavari(KATH).Nomcnclature followsPressel(7/(2000). OccurrenceofbirdandbuIterHywererecordede汁rouIetolhesamplin8pIols・Elhnobolanicalinfor‐ mationwere8alhered(romthcnearestvilla8e/settlementsofthesampIingploIs(orwhichrapidethllo‐ bolanicalappraisal(Marlinl995)wasapp1ied、Ve8eIationdamwerealsocollected(rom64selectedpointsm anelevaIiollaldi(fercnceof100men-routeloIhesamplin88ridswhichmn8edfrom200-800m. RESULTS Landusepattem GISanalysisoflherecelltmap1992showedthatIhemajorlandilltheChuriyail1theeastemNepal wasoccupiedbylhcforestswithanareaof477.28kmzcoverillg61、25%ofIhetotalarea(779.24km2)andthe a8riculIurelandwas221.71kmz(28.45%).Sandand8ravelcoverednearly6、0%indicaIin8Iheprescnceofa lar8enetworkofriversandslreams、Shruband8razil181andoccupicdL5%each,andthercsIlalld-use includingllrbanareilmade1.5(ルl〕oththe(oresIsanda8riculIurelaI1dsvariedinsimilarpaIterllinlheir altitudinaldislribuIionwhiChwe1℃hi8hinlowerelevation(〈300m)(l11ddecreascd8mduallyinlhehigher elevaIionshowinganinlerrelaIionshipbetwccntheforeslsanda8riculIuralpracliCcsinNepal.(Fig.3). AnaIysisshowedIhattheforestcovcrhasdecreasedbyover25(>(,inlesslhan35years、1,1958,the tolalareaoftheforeslsinIheCllllriyawas631.6km2sharil1881.0%ofIheIotalarea・Byl978,ildecrcasedb)’ 10%orallareaequivalcntlo63、5km2.1,1992,itlurIherdeIerioratedandtheforeslsshrunkloaIomlal・eao( 477.2km2.Durill8Ihcsameperiod,thea8ricullurelalldhadincreascdbymorethanl20%,i、e・(romatolal areaoflOl8kmziI11958to221.7km2inl992、Also,incrcmclltinagriclllturelandwashi8hcomparatively belweenl978-l9921hanl958-1978、WhileIhchi8hesI8aillwasfora8riculturewithl271kmzor56%oflhe totalchangedal・ea,andamar8inalchallgeforurbanareawilha8ainlcssIhanO.'別,ofthetolalchallgedarea、 Similarl)',grazin81qndalldshrllblandhadincrcaseddllrillgIhepcriodattheexpenseofforesIs・Onthe otherhaI1d,IherewasaliIlle8ain(489km2)(orlheforesIswhichincludedsomeforms()(})lanIatiolls(Tablel). -98- 兜CpbO S4DOO⑪ 920口40 兜00(。 GpOOOO LRndu6GPnlI「ml”2o「11】eChuriyaoEa81NepaI ④ ; !  ̄1 ; ; ; K瓢i鰹sVil60n既> EATA卜>・DCE 8cU唾ご丁CO■哺已ご■o及TO0z毛一輩c;OmbDDrごOnHra比FGtlOM Lシご-●0V、~▲●●q「いHeABWpCケ。“上■0●クワ■PDi■ゼハ■U2.■O凶LODW 10 0 10 20KIlOmetBrB  ̄ ̄---- ̄ 、Gmo 94bbOO 46○0屯、 S4nOCO OCmOO Fig.3.Li1I1dusepMIcrll()[IheChllri)'ilill]992,c(1s比rIlNepnl Tmblel L【ln(lljsepatterIlinl958,1978(I、(ll992inChurjynDea81ernMlpal LanlIll5e 1992 1978 1958 A「PJu(km2)A「、(兜)Area(kInJ)ATG、(外)A「PMkm2)A「e、(兜) F【)rlNbil (i:1155 H1U 568,2 72.9 477.23 61.2 AgriCIIltll「eLan(I 1()1.M lIM 138」`1 17.7 221.71 28.5 SllIlI1DL【111. 4.96 Il6 1870 2.4 1152 15 (HnwinHland 5.27 0.7 3.56 05 1087 1.4 35.62 ‘1.6 5(1.82 6.5 53.12 6.8 ()lllID「日 rroIIlI 779.241()0.0771124IOU()779.24】00.0 FIoristiccompositionandforestsIructu「e Ow「265vtlsclIIarplantHpccics,belol1llin8Io85(ilmiliesandl788enera,wcrerecordedfromIhe studysitcs、Famil>'L(D8uminosacincludcdlhchi8hestllumbcro(sl)(J(jcs(25SPP.)whichincludedrare/ Ihremtene(1t1℃esIxjDcicHlikeD(lルノXhlj(l/llljβ()i11iIlldA(T7dll(wl(,「Alイ,and(m(lemicspecicsOr"J()Smgノロllr1l・Olher familicsc()mprisiIl8hi8hnumb(Pr()(specieHwereCmmineae(l6Hl〕P.),Composime(l2spp.)and Euph()rbiacekle(l()sl)1).).Byhabil,11℃eandsub-Ireespeciesc()mprisedIhehi8hesIcoml〕osiIioI1wiIh45%of thet(〕mlsl1ccies,(()Ⅱ()wedbyllcrbs(22%),Shrllbs(20%),LinIm(10(X))imdI7ern(3%). Outofl61)Ii1nIso(Nepalincludedi11ClTljS,threespccics(CM11/IMC(,(]s/】.,Cyals1)Pclj"(11(landR(IWC伽 serノノピ""11(1)were(()llndinthestudicsarca(〕「Chllriya・AIOいIC(36individualsofCノ)(,clj)l(lmwer(Dobserved -99- whichwerefoulld8rowin8mosl1yinlheS〔ノli"川and/orSノノo'wJforesIsillassocialionwithPノIOF"/Xsp・and/ orPnl1(mlIlls/ilscic1(/nrjs,Similarly,l9individualsofCl/"lハclJsp・wcrealsolloIed・Therecordedindividualsof bolhspecicswere8rowillgnearslrcamsandinIheeasIernsitesinlhcalIiIlldinalmn8eof300-750m・Seven rareand/orendal18ercdspecicsidclllifiedb)'IUCN,andSeveralspecicsofspeci[icsciellti(icinlerestwere alsonoIed・ TI1eforesIs()[ChllriyaillcilSIerllN(J1)alwereesIimMedwiIhilbi1st】la1℃a(1)A)(〕f37.28m211(T1anda densityof78651111a-1.Tlle1〕【℃(1ominallttreespeciesatcanol〕ysIl・aIum(hei8ht〉10m)wereSノノ(〕'(?(1)UDI(Sm (relativebasalarea:`#1.0`X,),Tl,r"lij11l1nlRMl1(RBA:6.8%),L(ISU’rSlroC"ln1/)'1パ)(/11()m(5.0`X,),imdAdi11l1cordl/bノノ(l (5.0%).MeanDBH(diameIeraIbrcaslhei8hl)ofIhesespecieswas26、4cm,Ihou8ha(CWS・rol)!(smhada maximllmDBHo(124.()cm・ThcdomillanlspeciespresellIedaspomdicIweofsizc-classdislribulionindi‐ catin8IhevariaIiollinsnmp1epIoIsand/orpresenceofspecicswiIhdifkrel1la8e8rolll)sBesideIhesespe‐ cies,IhcmoslfrequcnIlyoccurril18(Freq.〉40%)treeswereS(wlFalrノノl(s(1)1,1(w(/jl("1,ハルizilllFl)l)(,ck,Syzl/Rjll"I Cノイ"lノノlj,(1)!(lScノノノ"l(、()(1"j(ソlノノ.SMIノノicノノノノwas()bscrvedupIo300m,alowcslrecordo[iIsoccurrcl1cewhich couldbeaIIribuledIoIhemoislerclwirollso(eastcmNepal.(Fig.4,Tablc2). TheundcrslorysImlum(111.〈4m),hadsl〕ecieslikeCoMjloolW()ノ)”jli/b/i(](I;『(x1.329%),Elリノl110rM〃 SPP.(Freq.>22%),l似,(ISP.(Frcq・'8.6%),MII(?s(J1"())11(1)川(Freq、17.1%),al1dlll)(,ノl"I(,ノブ111)/(lllノノ)ノリノノハ】(F「(Jq、15.7%). Thehi8hcsldensiI)',h()wcver,wnsllli1IofCノビrolノ1W,lrM"lu1scosl""wiIh29lOI1M1、EIIノ】111()rnl"ISP.`COノ)/)()sノリbノ、, andノリloCllなsyノ【!(PSlrjs,nlreespeciesbutoccurringinthellllderstor)'illIhesludysite(Avem8cll1.1.3m), sharedcompamlivcl)'ahi8hbasalarea,eaChhavill8RBAover3、0%・TlwI;roundslmIllmwaspoorl)'devel‐ Table2 Structuralpmrameter8o1.,1lljortreespecie8intheChuTiya,ea8terllNcpaI SNSpecies 1 652208250404350 1 222221212121112 1 ●●巳●●け。□凸の■■■e□ 196860733365202 980868858715871 1 5.6 11 7.7 1 8423.4 28605.8 7.2 116.7 436027036477655 15744.9 8.2 289787050374140 TOTAIJ 9204.1 8.2 ●●●●●●■●●■●●。p● 15”〃"/"イノノノ〃DPノ〃が(?〃 36715.8 14.0 068850860841125 141DutlIliKMM 8860.3 12.1 5 l3此"。()“〃"〕"Sルβノフノノル"ノノ 56638.2 19.8 809151617516651 12MWo/usノルルノノjnPw7s/s 9569.3 19.1 10.7 22221221213113 llL"""“(W切刃”此〃〔Pa 79181.5 16.1 604926520551034 lOGrelrjDQ/Ww 239444 37.3 24.3 611 9/ldma〔WIメノハWap 29240.9 162.4 960684856333032 86Ver〔Wノノ〃17"OS〃 366922 ●●●CG■●■●。●。●●● 7SCA/"】HDWタノ〃〔.A〃、 108941.8 008385065636058 6SKI'2J<9Wノ"jclノ、/"ノ 80047.0 外/baNlnoallmaxl11eaumax 456211503020101 5′1ノハノzノィノノC66eCA 702871.1 RI)ADensStcII】I)l)11(cn】)Ileight(、) 4 4&v"e(?/jl7wJw7"JJ(wYmノ") F●の。●■●●■DB●●白● 3九mノノ”ノ、aノ"/a‘ 704399073444639 21,agWSlmCmla”「I'ノノル、‘ 501422054111842 875444433333222 1SAO八DJJm6"SlJJ' BA Freq91cm2 1597312.0100.0786.5337017.81247.475 巾donlinnntsI)ecic3byIlIlA#Loco11Mlme -100- Hoi9hIClassDisIribution DBHC1assDisI「ibuIion 62840626284062 Sho「eambusIa 。 } 2860 050100 22111 C LageノsI/O0m/a pa/vノソノOノa 0102030 1■■■■■ 284 211 062 1 ・ ・ 。 二 ■ 二 ■ 。■ ■■ |■■■ ■■■ ■■■■ ■■■ ■□■■ ■■■ ■■■■■ 211 05050 ロロロロ■四四四m四四m 25 22111 l[|lOlm1MllllIlL- 髻 二 7,7)/、aノノョa旧la 020406080100 22 201 Scノリ/mawaノノルノリノノ 一 120 051015 22 84062 (【上)の②⑩一○五m|の工 111 DBHclass(c、) 111 1L1111,,M,繊癬 84062 1[''1.1ⅡLhⅧ- = Somecamus anacaU7um 05101520 Slemnumber(N) Fig.4.DiamIcratbrcasthcight(DBIl)alldheightcl&ISSdisIribulion ofmajortrecsl)cciesinthcC11uriya,castemNepal ‐101‐ opedwilhacovem8eof15.4%inavem8ewhichcanbeatlributedtoprevailingdryseasondurin8Iheinves‐ ti8aIionandwidespreadover-grazin8.Themostabundantspeciesbyfrequencyofoccurrencewere OP1js"IC"llssp.(Freq、23.46%),C(lノノノルe(/jIllll(lssi"l庇(Fre〔]、16.54%),andSi(ノロcor(ノlJm(Freq、10.38%).Byrelativedominance,Cq5sj)"i化,Arll)!dj"eノノロノノビノ)(l1cllsjs,O/)ノノs"IC'11イSSP.,A8〃(111"〃CO'11/zoi(lcsandCI1rcxcrlにjMJ weremostprevalelllinlhearea. TreeSpeciesassociation Adendro8mm(romsimilarityanal)'sisbasedonRBAandHorisliccomposiIioncate8orizedIhesludy siteintoseven8roupswithvar)'in8diversiI)'index・T11emajor8roul)swerelhatofdominaledbyShoグピ(l rol)IIS、,Tcr"!j"ロノノm/lllm,L"gersfluf1"j(1ノwM()r(1,A(/j"17corllリt)ノ、,Scノji"l(JM1nノノノcノlijalldBol"l)llxcc肋11.Thepre‐ dominalltspecieswasS、roblイsminIhestudiedsiteindicaIin8ilshi8her8erminaIiollpercellla8e,hi8her adaplabilityandhi8herrateofsurvivorship・ThespecieswasreCordedasdomil1antspeciesin24siIes(375% oftheloIal),andco-dominantillatleastllsiles、However,asalsoshownbythedendrogmm,S、rol)Msm appearedalon8wilhotherdomiI1antand/orco-dominantlreesinthosesites(Table3). Tablc3 A8sociatinnfrequency(Wi)ofmajortreesI]eciesintheChuriya,ea8ternNePal A8sociated AssocintionFreq,9【1 ,SルoreamOUsfa 7.日ノaraL./wW/701as.、"“"畝"mawaノノノcノW4.CuJYノノノbノノaTotal3pp. 42.18 ルrm/"且ノノaJTノdwla 35.93 32.81 21.88 20.31 28.13 17.19 938 17.19 2031 7.81 17.19 9.38 7.81 La“rszJUcm/βノww〃om 8bme“"【ノsD"acam/""J 7.81 SbAjmaII'βノノノcノケガ A磁血“J1"/b"a 624209 444323 species OutofGlsaml〕lill8sites,S、rol)l(smwas「oulldassocialedwiIhoneormores}〕ccicsin49siles,andwas 8rowingwith46t〉'pcsoftreespcciesinIotalWhilcL・ノノ(1パ1帆omand7.、MhadanassociationwiIh44and 42t>'pesofspecies.S・roblイsmshowedthehi811cstassociaIiollwiIhalldominanIspecies;with7.llノ(J1q42.2% frequency,withLノ)llW7or"35.リルwithS.("1(lmrlノノM1"32.8%,withS.【()(リノノic/lii21リルandwithA.coll/(/b/jn 203〔ルAhi811associaIionwasalsoseenbelwceI1T.(Jノロノ(IandL./)(1M/11(〕m(28.1`X,).Onlheotherhand,S・ 『(MJノノノcMshowedlesserassociaIionfrequency(<10%)exceptwiththeS、roblィ5,.Amoll8Ihedominantspe‐ cies,S、rol)lイsl(JhadIhewidestdisIribuIiollmn8eandoccurredfrom200-800malliludes,whileST()(lノノノcノJノノ wasbasicallyamidllillelementmostlydistribuIedabove500m. Localknowledgeandethnicgroups ThepreselltsIudydocumellted82sI〕eciesofmedicinalpIallIsand76wildedibleplanIs・A1Io8ether ‐102- 301ypesofdiseaseswere(oulldIobeIreaIedbyphyIo-therapyusil1gIhereported82plantspeciesinlhe Churi)'aofeastNepaLAz(7(ノノmcノノW"(ノノclJ(Neem)wasreporIedtobeusedforlhehi8hestnumber(10)of diseasesasanIiseptic,anal8esic,aIlIispasmodicalldanIipyreIic・Thespecieswasfoulldalsoeffectivetocure diabetes,gastric,sllakebiteand化verduetotyphoid,malaria,pIleumollia・Threespecies,viz.P/lyl!(1)11ノllls (リ"lノノノc、(Amala),AqRノe"、r"|ビノ0s(I〕Cl),andノリ1ノO8mcMfA11sl/iyrsリ70111s(Chuwa)werereportedtobeusedforsix differenldiseaSes、Similarly,RMU01/1ims(】'7)c"lj1l1l(CI1andmaruwa),B1llfl1"1i"or(Bllolelro)andTell"j"(1ノノロ”んrkW (Barro)werealsofoul1dformulIi-usesIreatin8atleastsixdi([erellIIypeso(coml〕lainIsanddisorders・ Amongthe{ourmajoreIhnic8roupsinterviewed,Limbureporledalolalof64medicinalp1anls (781%ofthetotalpIanIsreportcdf()rmedicinalpurposes)whileBahun-ChheIriin(ormed55suchspecies (67.W,ofthetolal).Similarly,RaiandMagarinformed43and37spccicsrespectively(Fig.5).Inavera8e, cachindividualbelon8il18toLimbu,RaiandMagarwasfamiliarwithlcssthanfourspeciesofmcdicillal p1anIs,whileeachindividualbelon8in8IoBahun-ChheIriwas(amiliarwithmorethamfivesuChplanls・The LimbuswerethepiolleeringalldsIilldominalltin11abilanIso(theChuriyaiIlI〕aslNepal、BahlmandChhelris, dominaiingsocialHillduclasshtwe8ivenIhemaccesstobeltcrIhcircdllcalion・TheyhadklIowled8eon AyurvedandotherHilldulileralu1℃whichcontainaweallhofkllowled8eonplanIsalldplantuses. 00000000 7654321 ,の⑩.⑩←仁⑩一Q一ロに一○一つのE一○・oz DUB2 トいiMaC Chhetri EIhnicGroups Fig,5.Nllmbcrofmedicinalplantsusedbyvarious ctbnicgroupslivillgintheChuriyaofl〕asINcI)al Distributionofbutterilies A1to8ether35speciesbelon8in8Io288enemand8{amilieswcreellumemtedfromthepresenIsludy siIeoftheChuriyaofeastemNepal(Table4).Thedocumentedspeciescom1)rised5.3%ofthetotalspeciesof butterfliesknowntoNcpal・Cellusl)r(,cjSwasfoundIobethemostcommonwilhiIsfourspecies,while /Mノノyノノノ(1,,,)M1イs,Dピノノl7sandEノ!/"lノノノ(lswerealsocommollinIhispmrI(hercstweresinglespecies8enus・ Fi8ure2showsthatthcmajoriIyofrccordcdspeciescome[r(〕mNymphalidaefamil)'whichcomprised 34、3%fouowedb)'Lycaenidae(20.()!X)),Pieridae(14.3%)andSatyridae(11.4%). Consideringseasonalchan8c,thespeciesricImessofbuIlcr(lywas[olmdtoincreasefromwinterlo -103- sprin8season、Outof35speciescollecIed,15species(42.85%)werefoundinwinterseasonandIherestin sprin8・Asi811i(icantcorrelalionwasfoundbetweenthespeciesriclmessandthemonthsilldicalin8thatlhe ablmdallceofbullerflywasposilivelyaffectedbyapproachin8warmerdaysandavailabiliIyofmorefood thatislheneclar・Similarly,lhespecicsrichllessofbutterflyilllheChuriyaofeastNepal8raduallyincreased upto500m,lhel1itdecreased、Thistcndenc>'correspondswilhlheareaofshrub-landil1Ihestudiedarea indicaIin81hehabilatpreferencesofthcbulterfIies、However,IhehabiIatlossisadverselya(feCtill8Ihebut‐ terflydiversiI)'illNepal. Table4 Di8triIjUtionOfbutterflyspecieBiIldifroren0altitudes(、)and8oaBonBintbo Churi 8.eaBtcrnNepal A11iludG(【Ⅱ FdllliIvADclfics Scdsoll 2503504505506507508509501050 1150 WillIcrSprinFK lDdIiliollidoP 十十 十十 十十 十十 十十十 十十十 十十十 十十 十十 十十 十十 十十 十十十 + 十十 十十 十十 十十 十十 十十 + 十十 ノjセツノゾノわけ“ノロノc"s 十十 十十 ノブhjm"/q/raaJ7s(。/bにv'@ lDiPridkIc 十十 十十十 十十 十十 十十 十十 十十 十十十 十十十十 十十 十十十 十十 十十十 十十十 十十十 十十十 十十 十十 十十十 十十十 十 十 + +++++十 + 十十 十 十・十 + 十十十 + (姫sDmbU"〃〃ノo〔/Us 十 十十 卜 十十十 ハノcノヨノノノ〃sノ[Yノa 十十 LP"Ip〔w7ハノ&/『 + 十十卜 + 十 十十十十十 s`Ilyri(LMP 〃.wノノ"ノルラノ""/pノピノs aw""/“/Ww")"jFfra + 十十十十十十 /牝C/Sルノ〃んn 十十十十十十十 + 十十 ノ対"〔q/wブイlAoJMD'7曲 Z物""“/aノG/放/“ Aiw"Drp"/〃〃〃ノ” 十十十 ノケU『?/sノc"1,W/"s P'℃c店〃mlz"Ja 十十 A/bノWSルリゾィリs 〃、/Sウカノ/a 十十十十十十十十十 Jlノ"o"/〃イノノ"ノコノノロ 十 十十 Ab/sww/)"汀 NymuPIMIi(IdHc ノlrZr.l'"j〃ノノ” '1ノムリw'""jspノM〔WIom 十十 + NPuuwolDiilIolC 十十 L"ノノリ”Cs“c『/〔ws + 十十 JIT"ノ"ハノbsnノ“/O Z7zec〃ロ〃JMa 十十 十十 十十 + ↓、十 バノ、qノリWa"curjrG"fa"、ノさ ccAJs〃/"ロバノs/rT ルノノロノカoバパノSc〆c/c 十十 LWHG11i(1M、 ’1"J647rrノ、〃「Csル 十十十 Gtハイ'G/〕feJ:jwノメリ"")"ノ ノヴeJ7盃6mg副“D 十十 十十 戯"℃"、AC“6G + 十十 〃c〃“。“CO"ノbe.sノ 〃c〃"sAj7wcfc DdlMilLlc 十十 十十 m"`"s圃創“ ノルノノ""s`M''2}“mJs EfVy〔,“【Porp IIQPspcri(l`IIP 7℃匂"(/“ノノ〔広れ】SYI + ToIdI71625292423191311101521 Note:+dcnolespreseIlce -104‐ Baselineinformationonbirds Atotalofl50birdspecieswererecordedtotaling2,374birdsdurin8thefieldstudy、Fromtheob‐ serveddaIa,ilwasinferredthataspecies/timecurveisofinterestbecauseitnotonlyindicatesspecies richllessperullittimeilwesImentbutalsoascertainswhethertimeilwestmenIinthefieldwasadequate、Of the255observation,birdswerepredominantlyobservedeitheril1foresls(38%)orinfarmlands(31%).Ad‐ ditional24observationswerenoIinchldedintheanalysisasbirdswereseenHyingoutofa8ivenhabiIat type・MajoriIyofthebirdswerefOundina8riculturefields,especiallyattheedgeofcultivationordegraded shrubberies・ Allobservedbirdspecieswere8roupedmIofourcategoriestodeterminetheirstatus:endan8ered(E), vuh1erable(V),rareandconmlon、n1efirstIwocate8ories(EandV)constihlte4%ofthetotalspecies(Table5). Oftheobservedbirdspecies,gianthombill(Bllc〃osl)jcorlljs),paleblueHycatcher(M1イscjc(IノノHlll1ico[or),and loll8-tailedbroadbill(PSαノブso"J1/sdnl/ICI(sme)areendan8ered・SpeciesIhatarevulnerablewerecrow-billed drongo(Djc)wrlイsn'111〔Pcm"s),pintailgreenpi8eon(Trero〃npic(M、)andlargenecklacedlau8hing-thrush (Clmwノnxpuclol"ノノs).Theoran8e-breastedgreenpi8eon(Trero'11)jcjlJcl(7)wasconsideredrare、Somesixhabi‐ Iatsiteswhereallendangeredandvulnerablespecieswerefound,werecol1sideredimportantforbirdsand lhesesitesneedtobemoniIoredtocheckifthesespeciescontinuetoexistinsuchadversesettings. Table5 Bird8tatusandob8ervationfrequencyintheChuriya,ea8temNepal No.ofslDeriP5 No.ofobscrvjlioms 3313 51.(us 6 Endangered Vulnerable Rare 4 1 Commom 8 1 2359 ACKNOWLEDGMENT ThestudywassupportedbyProNaturaFundtolhefirstauthor・GISfacilityandCPSweremade availablebytheResourcesHimalaya,Kathmalldu・SincerestlhanksaretoDrT・NakamuraofNahlralHistory Museumalldlnslitute,ChibaandProf,DrMOhsawaofUniversityofTokyofOrtheirkmdCooperation・ AllGISworkwasconductedbyBidurBaidya・CiriJoshialldBhesholididHoristicsurveyDaya BhusalconducledbutterHysurvey.GokamaKhanalconductedbirdsurvey・WethankthemaU. REFERENCES Bhatt,DB、1977.NaturalHistoryalldEconomicBolanyofNepal・OrientLongmanLtd・NewDelhi Carson,B、1985.ErosionandsedimelltationprocessesinlheNepalHimalaya・ICIMODOccns/01JロノI〕αperL IntemationalCentreforlnte8ratedMolmtainDevelopment,Kathmandu、 CovI・ofIndial958.IMml1SMrWyMH/)I958SurveyofIndiaOf(ices,Covt、oflndia・ Hagen,T、1908.NCI)Hノ.HimalBooks,POBox42,Lalilpur,NepaLForthed・ IlMGNl988M(vslcrlリlJ'j/brPoreslrySecforMWm/MnjjjRqj0〃.MinistryofForesIsandSoilConservation,His maiestyosGovt・ofNepal/Covt、ofFinland,Kathmalldu・ Itihara,M,Tatsuo,SandNoboru,M、1972.Photo8eologicalsurveyoftheSiwalikrangesandTeraiplains, southeasternNepalJ、Ccoscje'ICCS,OsakaUniv、15:77-98. -105- HMGN/PINNIDA1996・LRMllsFM")1992.HisMajesty,sGovt・ofNepal/Covt・ofFinland,KaIhmalldu LRMP/HMGN1986、LanduseMapl978・LandResourcesMappingProiect、Dep1.o(Forests,HisMajesty0s Govt、ofNepal/Govt.o(Canada/KeatingEarthSciences,Kalhmandu、 Martin,cル1915.Elhllol)ohllll/:八M(?'ノlodsMnlllイロノ,ChapmanandHalls,Londoll、 Press,I.R,Shrestha,KK、andSutlon,、A、2000八"'loledCノ1“Mislq/ノノl(,F1ol()crill8ノリmllsq/Ncl"M・Natural HistoryMuseum,LoI1don・ RohlfJ.F・andSokal,R、R、1969.SmlMal17MノノCs.W、H・FreemanandCo.. 鞆l01Olプ【ルナトウーラ・ファンド助成成果報告il}(2001) 』曰 要 ネパールシワリク山地の植生の生態学的研究 動的景相域(ランドスケープ)における生物極管理 ブジュ・ディネス、ヨンゾン・プロラド シワリク地区は、ヒマラヤ''1脈の肢南城に位置 ル、シワリク地区の生態学的Iilj報は極めて少な する。地史的には縦NM1rい111域である。ここは、南 かった。この訓イ削り1究は、束ネパールのシワリク地 から熱帯、北からiliAイ|ザと二つの生態域の結節域で 区の生態学的ノlL礎情報の収集を主たる目的としつ もある。 つ、以下の6つのテーマについて詳しく研究され シワリク高地はネパールのIEI上の12.7%、森林面積 た。①()ISによる1958年と1992年の土地利用状況の の25.8%を占めている。nM;、土地|;;1発が増加し続 比較、②植物jlllおよび森林の構造と更新、③様々な ける状況の1'1で、シワリク地区の自然環境の分断 標高域における樹i1剛成、①地域民族の植物利用、 化が深刻化している。しかし、これまで、ネパー ⑤チョウ類の分布、⑥』洲|'1のベースライン調査。 -106-