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Untitled - Primate Research Institute, Kyoto University
ᐕႎ
Vol.42 ⋡ᰴ
Σ. Ꮞ㗡⸒ .................................................................................................................................................................................. 1
Τ. ᭎ⷐ
1. ⚵❱㧔⚵❱࿑㧘ㆇ༡ᆔຬฬ㧘ᚲຬ৻ⷩ㧕 ................................................................................................................... 2
2. ੍▚᭎ᴫ㧔⚻⾌㧘⎇ⓥ⾌㧕 ........................................................................................................................................... 7
3. ࿑ᦠ .................................................................................................................................................................................11
4. ࠨ࡞㘃㘺⢒㗡ᢙ࡮േᘒ ................................................................................................................................................. 13
5. ⾗ᢱ ................................................................................................................................................................................ 14
6. ੱ੐⇣േ......................................................................................................................................................................... 17
7. ᶏᄖᷰ⥶......................................................................................................................................................................... 17
8. 㕖Ᏹൕ⻠Ꮷ..................................................................................................................................................................... 22
9. ࡝ࠨ࡯࠴࡮ࠕࠪࠬ࠲ࡦ࠻㧔R㨯A㧕.............................................................................................................................. 22
10. ࠹ࠖ࡯࠴ࡦࠣ࡮ࠕࠪࠬ࠲ࡦ࠻㧔T㨯A㧕...................................................................................................................... 22
11. ᐕ㑆ࠬࠤࠫࡘ࡯࡞......................................................................................................................................................... 23
Υ. ⎇ⓥᵴേ
1. ⎇ⓥㇱ㐷࡮ነ㒝⎇ⓥㇱ㐷࡮㒝ዻᣉ⸳╬
ㅴൻ♽⛔⎇ⓥㇱ㐷㧔ㅴൻᒻᘒ㧘ࠥࡁࡓᄙ᭽ᕈ㧘♽⛔⊒↢㧕 ................................................................................ 24
␠ળ↢ᘒ⎇ⓥㇱ㐷㧔↢ᘒ଻ో㧘␠ળㅴൻ㧕............................................................................................................ 34
ⴕേ␹⚻⎇ⓥㇱ㐷㧔ᕁ⠨⸒⺆㧘⹺⍮ቇ⠌㧘㜞ᰴ⣖ᯏ⢻㧕 .................................................................................... 41
ಽሶ↢ℂ⎇ⓥㇱ㐷㧔⛔ว⣖ࠪࠬ࠹ࡓ㧘ㆮવሶᖱႎ㧕............................................................................................ 54
ነ㒝⎇ⓥㇱ㐷㧔Ყセ⹺⍮⊒㆐㧔ࡌࡀ࠶࠮ࠦ࡯ࡐ࡟࡯࡚ࠪࡦ㧕㧕 .......................................................................... 61
㒝ዻᣉ⸳㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧘࿖㓙౒หవ┵⎇ⓥ࠮ࡦ࠲࡯㧕 ........................................................ 62
⊕⋲ࡊࡠࠫࠚࠢ࠻ ....................................................................................................................................................... 72
ࡏࡁࡏ㧔ᨋේ㧕࡮ࡅ࠻⑼ 3 ⒳Ყセ⎇ⓥࡊࡠࠫࠚࠢ࠻.............................................................................................. 73
㐳ᦼ㊁ᄖ⎇ⓥࡊࡠࠫࠚࠢ࠻ ....................................................................................................................................... 75
2. ੤ᵹදቯ......................................................................................................................................................................... 76
3. ቇ૏ขᓧ⠪ߣ⺰ᢥ㗴⋡ ................................................................................................................................................. 76
4. ᄖ࿖ੱ⎇ⓥຬ࡮⎇ୃຬ ................................................................................................................................................. 77
5. ᣣᧄੱ⎇ⓥຬ࡮⎇ୃຬ ................................................................................................................................................. 77
6. ⎇ⓥ㓸ળ 㧔ᚲౝ⺣⹤ળ㧕 ........................................................................................................................................... 78
7. 㔤㐳㘃ቇ✚ว࠯ࡒ࠽࡯࡞ 2011 .................................................................................................................................... 78
Φ. ᢎ⢒ᵴേ
ࠣࡠ࡯ࡃ࡞ COE ߣߒߡߩᵴേ ................................................................................................................................... 81
Χ. ᄢဳࡊࡠࠫࠚࠢ࠻
1. ⧯ᚻ⎇ⓥ⠪ࠗࡦ࠲࡯࠽࡚ࠪ࠽࡞࠻࡟࡯࠾ࡦࠣ࡮ࡊࡠࠣ࡜ࡓ (ITP-HOPE) ............................................................ 82
2. ⚵❱⊛ߥ⧯ᚻ⎇ⓥ⠪╬ᶏᄖᵷ㆜ࡊࡠࠣ࡜ࡓ (AS-HOPE)......................................................................................... 83
3. ᦨవ┵⎇ⓥၮ⋚੐ᬺ㧦ᔃߩవ┵⎇ⓥߩߚ߼ߩㅪ៤᜚ὐ㧔WISH㧕᭴▽................................................................ 86
4. ࠕࠫࠕ㨯ࠕࡈ࡝ࠞቇⴚၮ⋚ᒻᚑ੐ᬺ㧦ࡅ࠻⑼㘃ੱ₎ߩⅣႺㆡᔕᯏ᭴ߩᲧセ⎇ⓥ ................................................ 87
5. 㜞ੱญኒᐲ࿾ၞߦ߅ߌࠆቅ┙ߒߚ㔤㐳㘃୘૕⟲ߩᜬ⛯⊛଻⼔▤ℂߦ㑐ߔࠆ⎇ⓥ㧔ⅣႺ⎇ⓥ✚วផㅴ⾌㧕........ 88
6. 㗡⣖ᓴⅣࡊࡠࠣ࡜ࡓ‫ੱޟ‬㑆ࠄߒߐߩ㔤㐳㘃⊛⿠Ḯࠍߐߋࠆᚢ⇛⊛࿖㓙౒ห⎇ⓥ‫ ޠ‬.......................................... 90
7. ੱ㑆ߩㅴൻ..................................................................................................................................................................... 90
Ψ. ᐢႎᵴേ
1. ౏㐿⻠ᐳ......................................................................................................................................................................... 92
2. Ꮢ᳃౏㐿ᣣ..................................................................................................................................................................... 92
3. ࠝ࡯ࡊࡦࠠࡖࡦࡄࠬ㨯ᄢቇ㒮ࠟࠗ࠳ࡦࠬ ................................................................................................................... 92
Ω. ࠽࡚ࠪ࠽࡞ࡃࠗࠝ࡝࠰࡯ࠬࡊࡠࠫࠚࠢ࠻
1. NBR ................................................................................................................................................................................. 93
2. GAIN ............................................................................................................................................................................... 93
Ϊ. ౒ห೑↪⎇ⓥ
1. ᭎ⷐ ................................................................................................................................................................................ 96
2. ⎇ⓥᚑᨐ㧔(1) ⸘↹⎇ⓥ㧘(2) ৻⥸୘ੱ⎇ⓥ㧘(3)৻⥸ࠣ࡞࡯ࡊ⎇ⓥ㧘(4) 㓐ᤨ൐㓸⎇ⓥ㧘(5) 㔡ἴ㑐ㅪ㧕 ..... 97
3. ᐔᚑ 23 ᐕᐲߢ⚳ੌߒߚ⸘↹೑↪⎇ⓥ ......................................................................................................................126
4. ౒ห೑↪⎇ⓥળ
‫ ╙ޟ‬12 ࿁࠾ࡎࡦࠩ࡞⎇ⓥ࠮ࡒ࠽࡯‫ ޠ‬....................................................................................................................126
‫␠ޟ‬ળⴕേߣ⣖㧙ࠗࡔ࡯ࠫࡦࠣߣಽሶ㧙‫ޠ‬...........................................................................................................127
‫ⴕޟ‬േ․ᕈࠍᡰ㈩ߔࠆࠥࡁࡓၮ⋚ߣ⣖ᯏ⢻ߩ⸃᣿‫ޠ‬...........................................................................................128
ࠕࠫࠕߩ㔤㐳㘃ߩ଻ోߣ␠ળ↢ᘒ⎇ⓥߦ㑐ߔࠆㄭᐕߩᣂߚߥዷ㐿 ...................................................................129
╙ 40 ࿁ࡎࡒ࠾࠯࡯࡚ࠪࡦ⎇ⓥળ‫ߩࡦࡒࡄ࡯࠼ޟ‬ᓎഀ㧦ㆇേᯏ⢻߆ࠄ㜞ᰴᯏ⢻߳‫ޠ‬......................................129
‫߇࡞ࠨޟ‬ੑ⿷ߢᱠߊߣ߈‫ ޠ‬......................................................................................................................................130
‫ ╙ޟ‬7 ࿁›ጊᲧセ␠ળ⹺⍮ࠪࡦࡐࠫ࠙ࡓ‫ޠ‬...........................................................................................................131
Ϋ. ㅌ⡯ߦ޽ߚߞߡ................................................................................................................................................................132
Σ. Ꮞ㗡⸒
ㄭᐕߢ߽ߞߣ߽ᔃ߇⸃᡼ߐࠇߚ੐㗄ߪ‫⎇ޔ‬ⓥᚲߦߣߞߡᦨᄢߩ⺖㗴ߢ޽ߞߚ࠾ࡎࡦࠩ࡞ߩ∛᳇
ߩේ࿃⸃᣿߇ߢ߈ߚߎߣߢߔ‫ޕ‬ੑᐕ೨ߩߎߣߢߒߚ‫ߪ⑳ޕ‬ᒰᤨ㔤㐳㘃ߩ㘺⢒࡮❥ᱺ࡮ஜᐽ▤ℂࠍ
ᜂ߁ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㐳߅ࠃ߮∔∛ኻ╷ᆔຬ㐳ࠍઔߖߟ߆ߞߡ޿ߚߩߢ‫ޔ‬㞲᣿ߥ⸥ᙘ
ߣߒߡᱷߞߡ޿߹ߔ‫ޕ‬2001 ᐕએ᧪ 10 ᐕ㑆⎇ⓥᚲ߇⧰ᖠߒ⛯ߌߚ᩺ઙߢߒߚ‫ޕ‬಴ⴊࠍߣ߽ߥ߁⥌
ᱫᕈߩ∔ᖚߢᴦ≮ᴺ߽ߥ޿㊀◊ߥ∛᳇ߢߒߚ‫ޕ‬ේ࿃ߩ♻ญߔࠄ⷗߃ߕ߽߇߈⧰ߒߺ߹ߒߚ‫ޕ‬10
ᐕ⋡ߦߒߡ૗ࠄ߆ߩᗵᨴ∝ߢ޽ࠆߣ޿߁⛉ࠅㄟߺ߇ߢ߈‫ޔ‬ᚲᄖߩ⎇ⓥᯏ㑐ߦ౒ห⎇ⓥࠍଐ㗬ߒ‫ޔ‬
ߘࠇߙࠇߩᯏ㑐ߩᓧᗧߥ⎇ⓥ㗄⋡ࠍㅴ߼ߚ⚿ᨐ‫ߩࠬ࡞ࠗ࠙ࡠ࠻࡟࡞ࠨޔ‬ᗵᨴߦࠃࠆ‫࡞ࠩࡦࡎ࠾ޟ‬
ⴊዊ᧼ᷫዋ∝‫್ߣޠ‬᣿ߒ߹ߒߚ‫ޕ‬᳿߼ᚻߪ RNA ߿ DNA ࠍ↪޿ࠆಽሶㆮવሶᬌᩏᴺߢߒߚ‫ޕ‬10
ᐕ೨ߪߎߩᣇᴺ߇⏕┙ߒߡ޿ߥ߆ߞߚߩߢߔ‫ޕ‬ᚲຬߩ ๮ߩദജ߇ഞࠍᄼߒߚߎߣߪ޿߁߹ߢ߽
޽ࠅ߹ߖࠎ߇‫ޔ‬ᚲᄖߣߩ౒ห⎇ⓥ߇ᄢ߈ߥ᳿߼ᚻߦߥࠅ߹ߒߚ‫ߦߐ߹ޕ‬ᔟ᜼ߢߒߚ‫ޕ‬
ߎߩ∛᳇ߪ࠾ࡎࡦࠩ࡞ߦߣߞߡᣂ⥝ᗵᨴ∝ߛߞߚߩߢ‫ޔ‬ᣢሽߩ࠺࡯࠲߇߶ߣࠎߤᓎߦ┙ߜ߹ߖ
ࠎߢߒߚ‫౒ޕ‬ห⎇ⓥ࡮ㅪ៤⎇ⓥ߇⸃᳿߳ߩ㆏ࠍ㐿߈߹ߒߚ‫ޕ‬ᚲౝ࡮ᚲᄖߩซᥓߩ⚿㓸߇ේ࿃∛ේ
૕ߩ⊒⷗‫ޔ‬㒐ᓮኻ╷‫ޔ‬ᴦ≮⮎ߩㆬቯߥߤᄙߊߩߎߣࠍ৻᜼ߦㅴ߼ࠆߎߣ߇ߢ߈߹ߒߚ‫ޕ‬
㔤㐳㘃⎇ⓥᚲߪ 1967 ᐕ㧔ᤘ๺ 42 ᐕ㧕ߦో࿖౒ห೑↪⎇ⓥᚲߣߒߡ⸳┙ߐࠇ߹ߒߚ‫ޕ‬2010 ᐕ㧔ᐔ
ᚑ 22 ᐕ㧕ߦߪ೙ᐲࠍ޽ࠄߚ߼‫౒ޔ‬ห೑↪࡮౒ห⎇ⓥ᜚ὐߦᛚ⹺ߐࠇ㔤㐳㘃✚ว⎇ⓥ᜚ὐߣߒߡ
㔤㐳㘃⎇ⓥࠍផㅴߒߡ޿߹ߔ‫⎇ޕ‬ⓥౝኈ߇߹ߔ߹ߔⶄ㔀ߦߥߞߡ߈ߚߎߣ߿㔤㐳㘃೑↪ߩ㊀ⷐᕈ
߇㜞߹ߞߡ޿ࠆ⃻࿷ߢߪ‫౒ޔ‬ห೑↪࡮౒ห⎇ⓥߪ߹ߔ߹ߔ㊀ⷐᕈࠍჇߔߎߣߢߒࠂ߁‫ޕ‬
ߎߎߦ߅ዯߌߔࠆᐕႎߪ⎇ⓥᚲߢታᣉߒߚ⎇ⓥᚑᨐ߿੐ᬺㆀⴕߩㆊ෰৻ᐕ㑆ߩ߹ߣ߼ߢߔ‫ޕ‬ᚲ
ᄖߦᐢߊ㈩Ꮣߔࠆߎߣߢ‫⹏ޔ‬ଔߩኻ⽎ߦߥࠆߢߒࠂ߁ߒ‫ޔ‬ᚲౝߦ߅޿ߡߪ⸥㍳߿෻⋭᧚ᢱߣ߽ߥ
ࠅ߹ߔ‫ᧄޕ‬ዊౠሶߦߪᚲౝߩᢎຬ࡮⎇ⓥ⠪߿ᄢቇ㒮↢߿⡯ຬߩ⎇ⓥᚑᨐߥࠄ߮ߦታᣉౝኈ߇⸥㍳
ߐࠇࠆߛߌߢߥߊ‫౒ޔ‬ห೑↪࡮౒ห⎇ⓥߩታᣉႎ๔ߣᚑᨐ߽ోຬߩ߽ߩ߇⸥タߐࠇߡ޿߹ߔ‫ޕ‬ᤓ
ᐕᐲ߆ࠄ౒ห೑↪࡮౒ห⎇ⓥߩ࿖㓙ൻߦജࠍ޿ࠇߡ߈߹ߒߚ‫ޕ‬ᤓᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕ߪ 4 ઙߩ
ᶏᄖ⎇ⓥᯏ㑐ߩ⸘↹⎇ⓥ߇ታᣉߐࠇ߹ߒߚ‫੹ޕ‬ᐕᐲ㧔ᐔᚑ 24 ᐕᐲ㧕ߪ 9 ઙߣჇടߒߡ޿߹ߔ‫ޕ‬
࿖㓙⊛ߥ౒ห೑↪࡮౒ห⎇ⓥࠍផㅴߔࠆߎߣߢ‫ޔ‬వੱߚߜ߇Ⓧߺ਄ߍߚᚑᨐߣ‫ࠄ߆ࠇߎޔ‬ᣂߚߦ
ᒻᚑߔࠆ⎇ⓥ᜚ὐࠍၮ⋚ߦߒߡ‫ߩߡߒߣࡉࡂߩࠕࠫࠕޔ‬࿖㓙⊛㔤㐳㘃✚ว⎇ⓥ᜚ὐࠍ⋡ᜰߒ߹ߔ‫ޕ‬
߹ߚ‫੹ޔ‬ᓟߩฦ⒳੐ᬺߦ߅޿ߡ߽ቇౝᄖߩㅪ៤߇㊀ⷐߢߔ‫⁜ޕ‬㓖ൻߒߚᣉ⸳ߩߺߥࠄߕ୘೎ߩ⎇
ⓥ૕೙߽ⶄวൻߒߥߌࠇ߫㆏߇㐿ߌߥ޿߆ࠄߢߔ‫ޕ‬୘ੱ⎇ⓥߩ㊀ⷐᕈߪ⥄᣿ߩߎߣߢߪ޽ࠅ߹ߔ
߇‫⊒ߩߡߒߣ❱⚵ޔ‬ዷࠍ⋡ᜰߔߦߪઁ⎇ⓥᯏ㑐ߣߩㅪ៤੐ᬺ߽ⷞ㊁ߦ౉ࠇߡ߅ߊᔅⷐ߇޽ࠅ߹ߔ‫ޕ‬
੹ᓟߣ߽⎇ⓥᚲߦኻߔࠆߏᗧ⷗‫␜ߏޔ‬ໂࠍ߅ነߖ޿ߚߛ߈߹ߔࠃ߁ࠃࠈߒߊ߅㗿޿↳ߒ਄ߍ߹ߔ‫ޕ‬
ᚲ㐳 ᐔ੗໪ਭ
㧙㧙
Τ. ⎇ⓥᚲߩ᭎ⷐ
1. ⚵❱
(1) ⚵❱ߩ᭎ⷐ㧔2012 ᐕ 3 ᦬ 31 ᣣ⃻࿷㧕
ㅴൻᒻᘒಽ㊁
ㅴൻ♽⛔⎇ⓥㇱ㐷
ࠥࡁࡓᄙ᭽ᕈಽ㊁
♽⛔⊒↢ಽ㊁
␠ળ↢ᘒ⎇ⓥㇱ㐷
↢ᘒ଻ోಽ㊁
␠ળㅴൻಽ㊁
ᕁ⠨⸒⺆ಽ㊁
⎇ⓥㇱ㐷
ⴕേ␹⚻⎇ⓥㇱ㐷
⹺⍮ቇ⠌ಽ㊁
㜞ᰴ⣖ᯏ⢻ಽ㊁
ㆇ༡ᆔຬળ
ಽሶ↢ℂ⎇ⓥㇱ㐷
⛔ว⣖ࠪࠬ࠹ࡓಽ㊁
ㆮવሶᖱႎಽ㊁
Ყセ఺∉ᓸ↢‛㗔ၞ
⎇ⓥㇱ
ᚲ㐳
ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ
࠮ࡦ࠲࡯
㒝ዻᣉ⸳
ද⼏ຬળ
㊁↢↢‛ᄙ᭽ᕈ㗔ၞ
⥃ᐥ࡮∛ℂᬌᩏቶ
ᬌᩏ࡮▤ℂㇱ
ㆮવሶ࡮↢ൻቇᬌᩏቶ
❥ᱺ࡮⢒ᚑ࡮⑔␩ቶ
࿖㓙౒หవ┵⎇ⓥ
࠮ࡦ࠲࡯
✚ോដ
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⎇ⓥഥᚑដ
ળ⸘ដ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯੐ോដ
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᧻ ᴛ ື ㇢
⺪⸰ ర
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ᴡ ᧛ ᱜ ੑ
દ૒ ᱜ
౉ ૼ ◊ ผ
ਛ ㆏ ᱜ ਯ
੖⊖ㇱ ⵨
ጊ ᭂ ᄈ ৻
㒙 ᒻ ᷡ ๺
ਛ ോ ⌀ ੱ
ᐔ ੗ ໪ ਭ
㜟 ↰ ᣽ ᒾ
౎ ᧁ ቯ ⴕ
㧔᧲੩ᄢቇ✚ว⎇ⓥඳ‛㙚 ᢎ᝼㧕
㧔᧲੩ᄢቇᄢቇ㒮✚วᢥൻ⎇ⓥ⑼ ᢎ᝼㧕
㧔᧲੩ᄢቇᄢቇ㒮ᣂ㗔ၞഃᚑ⑼ቇ⎇ⓥ⑼ ᢎ᝼㧕
㧔↢ℂቇ⎇ⓥᚲ ᢎ᝼㧕
㧔ℂൻቇ⎇ⓥᚲ ࠴࡯ࡓ࡝࡯࠳࡯㧕
㧔ᄢ㒋ᄢቇᄢቇ㒮ੱ㑆⑼ቇ⎇ⓥ⑼ ᢎ᝼㧕
㧔ᬉጊᅚቇ࿦ᄢቇੱ㑆㑐ଥቇㇱ ᢎ᝼㧕
㧔੩ㇺᄢቇᄢቇ㒮ℂቇ⎇ⓥ⑼ ᢎ᝼㧕
㧔੩ㇺᄢቇᄢቇ㒮ℂቇ⎇ⓥ⑼ ᢎ᝼㧕
㧔੩ㇺᄢቇᄢቇ㒮ℂቇ⎇ⓥ⑼ ᢎ᝼㧕
㧔੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲ ᢎ᝼㧕
㧔੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲ ᢎ᝼㧕
⡯ຬߩౝ⸶
ᢎ᝼
ಎᢎ᝼
ഥᢎ
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੐ോ⡯ຬ
ᛛⴚ⡯ຬ
ዊ⸘
㕖Ᏹൕ㧔ᤨ㑆㧕
ว⸘
14
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6
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10
74
111
185
ᄢቇ㒮↢࡮⎇ⓥ⠪╬ߩౝ⸶
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16
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▸ሶ
▵ሶ
ដ㐳
੐ോ⡯ຬ
੐ോ⵬૒ຬ
੐ോ⵬૒ຬ
੐ോ⵬૒ຬ
੐ോ⵬૒ຬ
੐ോ⡯ຬ
੐ോ⵬૒ຬ
੐ോ⵬૒ຬ
ഭോ⵬૒ຬ
ഭോ⵬૒ຬ
ഭോ⵬૒ຬ
ដ㐳
੐ോ⡯ຬ
੐ോ⵬૒ຬ
੐ോ⵬૒ຬ
ᄙ⋡⊛ࡎ࡯࡞
ᯅᧄ ᕺ⟤
⇌ ⟤ජઍ
ᣣᲧ㊁ ᕺ⟤ሶ
ዊፉ ๺ઍ
ഭോ⵬૒ຬ
ഭോ⵬૒ຬ
ഭോ⵬૒ຬ
ഭോ⵬૒ຬ
ᖱႎᬌ⚝ቶ
⑔ን ᙗม
(3) ᄢቇ㒮↢
2011 ᐕᐲ ↢‛⑼ቇኾ᡹㧔㔤㐳㘃ቇ࡮㊁↢േ‛♽㧕
ቇᐕ
ᜰዉᢎຬ
᳁ฬ
⍹Ꮉ ⋥᮸
D3
Ỉ↰ ⃍ሶ
D3
㡨ᳰ ⩿଻
D3
દ⮮ Პ
D3
Ỉ↰ ᥏ሶ
D3
౗ሶ ፄ᣿
D3
⁚㊁ ᢥᶈ
D3
Christopher Martin
D3
(H23.6.30 ⹺ቯㅌቇ)
દ⮮ ␭ᐽ
D3
⑧භ 㓷ผ
D3
⷏ጟ ૓৻㇢
D2
ጊ᪸ ⵨⟤
D2
ᄢ⼱ ᵗ੺
D1
ዊᎹ ⹞ਫ
D1
ᷡ㐳 ⼾
D1
㋈ᧁ ධ⟤
D1
Yena Kim
D1
⏷᧛ ᦮ሶ
M2
૒⮮ ᧙ᄹ
M2
ਛ᧛
ᄢ⍹
ᱜ㜞
㜟੗
ች࿾
ᄢ⍹
㜟੗
Ძ೑
ඨ⼱
㜟੗
෹᳗
ች࿾
෹᳗
ᯅᧄ
᧻ᴛ
స᮸
㜞↢
ା↵
ᱜᚑ
㊀ᒄ
㜞↢
ᱜᚑ
ବ㓶
๋㇢
ᱜᚑ
㓷Ꮖ
㊀ᒄ
㓷Ꮖ
ජ⛗
ື㇢
MA Huffman
ᱜ㜞 ା↵
⷏᧛ ೰
ች࿾ ㊀ᒄ
ᄢ⍹ 㜞↢
㜟੗ ᱜᚑ
ᐔ㦮 ㍈⍫
᧻ᴛ ື㇢
㋈ᧁ ᮸ℂ
ᨋ ⟤㉿
ඨ⼱ ๋㇢
Ꮉᧄ ⧐
ᱜ㜞 ା↵
⷏᧛ ೰
ᱜ㜞 ା↵
⷏᧛ ೰
੹੗ ໪㓶
ฎ⾐ ┨ᒾ
෹᳗ 㓷Ꮖ
M2
᧛᧻ ᣿Ⓞ
M2
్ ࡝࡜
M2
ේ ᥰ
M1
ỳ↰ 㘧㠽
M1
ᳯፉ ବ
M1
ᓾጊ ᄹᏔሶ
M1
ᩉ ⥝㎽
M1
㖧 ᤘ᥏
M1
R Sayuri
Cicalise Takeshita
M1
ᬀ↰ ᗐ
M1
ᰞᐸ 㓁ሶ
M1
↰ਛ ⟤ㇺ
M1
ጊ↰ ᥓሶ
M1
ᅏ↰ ᵏᒄ
M1
Israt Jahan
M1
(4) ⎇ⓥᡰេផㅴຬ
᳁ฬ
MA Huffman
ᱜ㜞
⣁↰
ᱜ㜞
⷏᧛
ᣧᎹ ථᔒ
ା↵
⌀ᷡ
ା↵
೰
㧙㧙
੹੗
෹᳗
᧻ᴛ
㚅↰
෹᳗
Ძ೑
ฎ⾐
ᐔ੗
ඨ⼱
Ꮉᧄ
ฎᏒ
ጟᧄ
ᯅᧄ
ฎᏒ
੹੗
ฎᏒ
ฎ⾐
ฎᏒ
ฎ⾐
໪㓶
㓷Ꮖ
ື㇢
໪⾆
㓷Ꮖ
ବ㓶
┨ᒾ
໪ਭ
๋㇢
⧐
೰ผ
ቬ⵨
ජ⛗
೰ผ
໪㓶
೰ผ
┨ᒾ
೰ผ
┨ᒾ
MA Huffman
⿷┙ ᐞ⏴
෹᳗ 㓷Ꮖ
ᄢ⍹ 㜞↢
᧻ᴛ ື㇢
㋈ᧁ ᮸ℂ
ᨋ ⟤㉿
ᱜ㜞 ା↵
෹᳗ 㓷Ꮖ
ᱜ㜞 ା↵
⣁↰ ⌀ᷡ
㜟↰ ᣽ᒾ
ᐔ੗ ໪ਭ
ᐔ੗ ໪ਭ
ฎ⾐ ┨ᒾ
ណ↪ᦼ㑆
ⁿ㘺 ⦟ሶ
2011 ᐕ 4 ᦬ 1 ᣣ㨪2012 ᐕ 3 ᦬ 31 ᣣ
⪲ᩙ ๺ᨑ
2011 ᐕ 4 ᦬ 1 ᣣ㨪2012 ᐕ 3 ᦬ 31 ᣣ
ᄢၳ ⟤૫
2011 ᐕ 4 ᦬ 1 ᣣ㨪2012 ᐕ 3 ᦬ 31 ᣣ
ㄭ⮮ ߭ࠈሶ
2011 ᐕ 4 ᦬ 1 ᣣ㨪2012 ᐕ 3 ᦬ 31 ᣣ
ၳౝ ࠁ߆ࠅ
2011 ᐕ 4 ᦬ 1 ᣣ㨪2012 ᐕ 3 ᦬ 31 ᣣ
ฎᯅ ଻ᔒ
2011 ᐕ 4 ᦬ 1 ᣣ㨪2012 ᐕ 3 ᦬ 31 ᣣ
ਛᎹ ජᨑ⟤
2011 ᐕ 4 ᦬ 1 ᣣ㨪2012 ᐕ 3 ᦬ 31 ᣣ
2. ੍▚᭎ᴫ
੍▚᭎ⷐ ㆇ༡⾌੤ઃ㊄
㧔㊄㗵ߩන૏ߪߔߴߡජ౞㧕
ੱઙ⾌
‛ઙ⾌
527,495
159,760
‛ઙ⾌㧔․೎⚻⾌㧕
203,746
ᣉ⸳ᢛ஻⾌⵬ഥ㊄
0
⸘
891,001
ฃ⸤⎇ⓥ⾌㧔7 ઙ㧕
124,659
౒ห⎇ⓥ⾌㧔3 ઙ㧕
12,038
ฃ⸤੐ᬺ⾌㧔2 ઙ㧕
21,400
ᢥㇱ⑼ቇ⋭ᣣᧄቇⴚᝄ⥝ળ⑼ቇ⎇ⓥ⾌ഥᚑ੐ᬺ㧔71 ઙ㧕
ᄖ ㇱ ⾗ ㊄
236,357
ෘ↢ഭ௛⋭⑼ቇ⎇ⓥ⾌⵬ഥ㊄㧔3 ઙ㧕
37,000
ࠣࡠ࡯ࡃ࡞ COE ࡊࡠࠣ࡜ࡓ㧔1 ઙ㧕
18,770
ࠣࡠ࡯ࡃ࡞ 30 ࡊࡠࠣ࡜ࡓ㧔1 ઙ㧕
15,300
࠽࡚ࠪ࠽࡞࡮ࡃࠗࠝ࡝࠰࡯ࠬ࡮ࡊࡠࠫࠚࠢ࠻㧔2 ઙ㧕
ᦨవ┵⎇ⓥ㐿⊒ᚢ⇛⊛ᒝൻ⾌⵬ഥ㊄㧔1 ઙ㧕
⧯ᚻ⎇ⓥ⠪ᚢ⇛⊛ᶏᄖᵷ㆜੐ᬺ⾌⵬ഥ㊄㧔1 ઙ㧕
61,600
500,000
14,330
⑼ቇᛛⴚ✚วផㅴ⾌⵬ഥ㊄㧔1 ઙ㧕
4,094
⎇ⓥ⠪ᶏᄖᵷ㆜ၮ㊄ഥᚑ㊄㧔1 ઙ㧕
25,796
వ┵⎇ⓥഥᚑၮ㊄ഥᚑ㊄㧔2 ઙ㧕
65,495
ነ㒝㊄㧔3 ઙ㧕
2,865
㑆ធ⚻⾌㧘৻⥸▤ℂ⾌╬
66,787
ోቇ⚻⾌
6,145
⸘
1,212,636
ว ⸘
2,103,637
(1) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕ฃ⸤⎇ⓥ⾌ ౝ⸶৻ⷩ
⎇ⓥ⒳೎
⎇ⓥઍ⴫⠪
㊄ 㗵
ฃ⸤⎇ⓥ⾌
ᄢ⍹㜞↢
62,553,847
ฃ⸤⎇ⓥ⾌
㜞↰᣽ᒾ
11,500,000
ฃ⸤⎇ⓥ⾌
㜞↰᣽ᒾ
2,500,000
ฃ⸤⎇ⓥ⾌
ਛ᧛స᮸
1,000,000
ฃ⸤⎇ⓥ⾌
ฎᏒ೰ผ
45,139,231
ฃ⸤⎇ⓥ⾌
ਛ᧛స᮸
385,000
ฃ⸤⎇ⓥ⾌
ฎᏒ೰ผ
1,580,880
ว ⸘
7ઙ
⎇ ⓥ ⺖ 㗴
వ┵⊛ㆮવሶዉ౉࡮ᡷᄌᛛⴚߦࠃࠆ⣖⑼ቇ⎇ⓥߩߚ߼ߩ⁛ഃ⊛㔤㐳㘃ࡕ࠺
࡞ߩ㐿⊒ߣᔕ↪㧔ㆮવሶᡷᄌ㔤㐳㘃ࡕ࠺࡞ߩ㐿⊒ߣ㜞ᰴ⣖ᯏ⢻ߩ⸃ᨆ㧕
ਛᨔ␹⚻㓚ኂᓟߩ␹⚻࿁〝ౣ✬ᚑߣᯏ⢻࿁ᓳߩࡔࠞ࠾࠭ࡓߩ⸃᣿㧔ࠨ
࡞ࡕ࠺࡞ߦࠃࠆ⊹⾰⣄㜑〝ߩน႟ᕈ೙ᓮᯏ᭴ߩᬌ⸛㧕
࠼࡯ࡄࡒࡦߦࠃࠆⴕേߩ⊒㆐ߣ⊒⃻ߩ೙ᓮᯏ᭴㧔࠼࡯ࡄࡒࡦߦࠃࠆⴕ
േߩ⊒㆐ߣ⚵❱ൻߩࡔࠞ࠾࠭ࡓߩࠪࠬ࠹ࡓ⊛⸃ᨆ㧕
ࡑ࡯ࡕ࠮࠶࠻ߩᗵⷡ࡮⹺⍮ᯏ⢻⸘᷹ᴺߩ⏕┙
㜞ੱญኒᐲ࿾ၞߦ߅ߌࠆቅ┙ߒߚ㔤㐳㘃୘૕⟲ߩᜬ⛯⊛଻⼔▤ℂ
⺒ᦠ෸߮ቇ⠌ߩㆊ⒟ߦ߅ߌࠆᐜఽ࡮ఽ┬ߩⷞ✢ᄌൻߦߟ޿ߡߩ⺞ᩏ⎇ⓥ
The Bonobo Conservation Project:Working towards gazettement of the
Iyondji Community Bonobo Reserve in the Democratic Republic of Congo
124,658,958
㊄㗵ߪ‫ޔ‬㑆ធ⚻⾌ࠍ㒰ߊ
(2) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕౒ห⎇ⓥ⾌ ౝ⸶৻ⷩ
⎇ⓥ⒳೎
౒ห⎇ⓥ⾌
⎇ⓥઍ⴫⠪
ਛ᧛స᮸
㊄ 㗵
2,000,000
⎇ ⓥ ⺖ 㗴
㔤㐳㘃ࠍኻ⽎ߣߒߚേ૞ℂ⸃ߩᲧセ⎇ⓥ
౒ห⎇ⓥ⾌
ਛ᧛స᮸
9,129,000
ࡑ࡯ࡕ࠮࠶࠻ߩ⹺⍮ᯏ⢻ߦ㑐ߔࠆⴕേ⮎ℂቇ⊛⸃ᨆ
౒ห⎇ⓥ⾌
᣿㉿ብᢥ
909,000
ว ⸘
3ઙ
ࠨ࡞㘃ߦࠃࠆ㧴㧯㨂ᗵᨴ࡮⊒∝ࡕ࠺࡞േ‛ߩ᭴▽ߦ㑐ߔࠆ⎇ⓥᐔᚑ 23
ᐕᐲಽ
12,038,000
㧙㧙
(3) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕ฃ⸤੐ᬺ⾌ ౝ⸶৻ⷩ
⎇ⓥ⒳೎
⎇ⓥઍ⴫⠪
㊄ 㗵
ฃ⸤੐ᬺ⾌
᧻ᴛື㇢
16,400,000
ฃ⸤੐ᬺ⾌
ฎᏒ೰ผ
5,000,000
ว ⸘
2ઙ
21,400,000
⎇ ⓥ ⺖ 㗴
ੱ㘃ㅴൻߩ㔤㐳㘃⊛⿠Ḯߩ⸃᣿ߦะߌߚ⧯ᚻ⎇ⓥ⠪⢒ᚑ࿖㓙ࡊࡠࠣ࡜ࡓ
HOPE
ࡅ࠻⑼㘃ੱ₎ߩⅣႺㆡᔕᯏ᭴ߩᲧセ⎇ⓥ
(4) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕ᢥㇱ⑼ቇ⋭࡮ᣣᧄቇⴚᝄ⥝ળ⑼ቇ⎇ⓥ⾌ഥᚑ੐ᬺ ౝ⸶৻ⷩ
⎇ⓥ⒳೎
⎇ⓥઍ⴫⠪
㊄ 㗵
ᣂቇⴚ㗔ၞ⎇ⓥ
ਛ᧛స᮸
10,400,000
ᣂቇⴚ㗔ၞ⎇ⓥ
㜞↰᣽ᒾ
4,500,000
ㆇേ㓚ኂߣ⹺⍮㓚ኂࠍಾࠅಽߌࠆࡄ࡯ࠠࡦ࠰ࡦ∛ߩࠨ࡯ࠠ࠶࠻ࡄ࠰ࡠࠫ࡯
ᣂቇⴚ㗔ၞ⎇ⓥ
੗਄⻞৻
2,500,000
㔤㐳㘃ߦ߅ߌࠆ␹⚻〝ㆬᛯ⊛ߥᯏ⢻ಽሶ೙ᓮᛛⴚߩ㐿⊒
ᣂቇⴚ㗔ၞ⎇ⓥ
⿷┙ᐞ⏴
2,600,000
㗻⍮ⷡ᭽ᑼߩ㔤㐳㘃⊛ၮ⋚߳ߩᲧセ⊒㆐ቇ⊛ࠕࡊࡠ࡯࠴
ᣂቇⴚ㗔ၞ⎇ⓥ
દ᧛⍮ሶ
2,300,000
࠴ࡦࡄࡦࠫ࡯ߣࡅ࠻ߦ߅ߌࠆ⾰ᗵᖱႎಣℂߦ㑐ߔࠆታ㛎⊛ᬌ⸛
⎇ⓥಽᜂ
㧔ᣂቇⴚ㗔ၞ㧕
ᴡౝጊ㓉♿
1,000,000
ᣥੱ࡮ᣂੱߩቇ⠌ⴕേߦ㑐ߔࠆ⣖ᯏ⢻ࡑ࠶ࡊߩ૞ᚑ
․೎ផㅴ⎇ⓥ
᧻ᴛື㇢
67,100,000
⹺⍮⊒㆐ߩ㔤㐳㘃⊛ၮ⋚
ၮ⋚⎇ⓥ(S)
㜞↰᣽ᒾ
24,000,000
␹⚻〝ㆬᛯ⊛ߩᵴേᛥ೙ߣ࠻࡟࡯࠾ࡦࠣߦࠃࠆᄢ⣖ࡀ࠶࠻ࡢ࡯ࠢߩ᭴▽ߣ
ᯏ⢻ߩ⸃᣿
ᶏߩߎߎࠈ‫ޔ‬᫪ߩߎߎࠈ㧙㟗㘃ߣ㔤㐳㘃ߩ⍮ᕈߦ㑐ߔࠆᲧセ⹺⍮⑼ቇ㧙
⎇ ⓥ ⺖ 㗴
ઁ⠪ߣߩ⋧੕૞↪ࠍ੺ߒߚᖱႎ₪ᓧࡔࠞ࠾࠭ࡓߩ⸃᣿
ၮ⋚⎇ⓥ(S)
෹᳗㓷Ꮖ
35,500,000
⎇ⓥಽᜂ㧔ၮ⋚ S㧕
ᐔ↰ ⡡
1,000,000
ᗧ⼂࡮ౝ⋭࡮⺒ᔃ⹺⍮⊛ࡔ࠲ࡊࡠ࠮ࠬߩ⊒↢ߣᯏ⢻
ၮ⋚⎇ⓥ(A)
ᐔ੗໪ਭ
7,100,000
ࠥࡁࡓਇᲫ࿾Ꮺ㧔RCRO㧕ߩㅴൻߣᗧ⟵
ၮ⋚⎇ⓥ(A)
ự↰ ⓛ
6,000,000
ࠕࠫࠕᄢ㒽ㇱߦ߅ߌࠆࡑࠞࠢߩㅴൻ࿾ℂቇ㧦⒖૑ߣᄙ᭽ൻߩ⸃᣿
ၮ⋚⎇ⓥ(A)
ฎᏒ೰ผ
6,500,000
ࡅ࠻⑼ߦ߅ߌࠆ᡹᠄ᕈߣᛥ೙ߩࡔࠞ࠾࠭ࡓߩㅴൻ㧦Pan ዻߩ㓸࿅㑆࡮㓸࿅ౝ
੤ᷤߩಽᨆ
⎇ⓥಽᜂ㧔ၮ⋚ A㧕
㋈ᧁ᮸ℂ
1,000,000
㐳ኼߣㆡᔕߩࡎࡒ࠾࠯࡯࡚ࠪࡦ
⎇ⓥಽᜂ㧔ၮ⋚ A㧕
ᯅᧄජ⛗
1,200,000
ࠕࡈ࡝ࠞ↥ࠝ࠽ࠟࠩ࡞⑼㔤㐳㘃ߩ↢ሽᚢ⇛ߣᒻᘒߩ㑐ㅪ
⎇ⓥಽᜂ㧔ၮ⋚ A㧕
ጟᧄቬ⵨
1,000,000
਎⇇ߦ߅ߌࠆࠛࠠࡁࠦ࠶ࠢࠬ‫ߩ⯻᧦ࠕ࠾࠹ޔ‬⒳ಽൻ‫ޔ‬ಽሶ౒ㅴൻ⎇ⓥ
⎇ⓥಽᜂ㧔ၮ⋚ A㧕
੹੗໪㓶
100,000
㔤㐳㘃ߩ⥄ὼ㓸࿅ߦᵈ⋡ߒߚᗵⷡ㑐ㅪㆮવሶߩᄙ᭽ᕈߩត⚝ߣㆡᔕㅴൻߩ
ᬌ⸽
⎇ⓥಽᜂ㧔ၮ⋚ A㧕
Ꮉᧄ
650,000
ᾲᏪ㜞࿾ⅣႺߦ߅ߌࠆኅ⇓ൻ࡮’⇓ᚑ┙ㆊ⒟ߦ㑐ߔࠆቇ㓙⊛⎇ⓥ࡯ࠕࡦ࠺
ࠬࠍਛᔃߦ
⧐
ၮ⋚⎇ⓥ(B)
੹੗໪㓶
2,800,000
ࠥࡁࡓᄙ᭽ᕈࠍၮ⋚ߣߒߚ㔤㐳㘃ߩ⒳ౝ࡮⒳㑆ᗵⷡ․ᕈߩ⸃᣿
ၮ⋚⎇ⓥ(B)
㜞੗ᱜᚑ
3,200,000
᧲࡙࡯࡜ࠪࠕߦ߅ߌࠆᣂ↢ઍᓟඨߩ㔤㐳㘃ㅴൻߦ㑐ߔࠆฎ↢‛ቇ⊛⎇ⓥ
ၮ⋚⎇ⓥ(B)
ᐔ੗໪ਭ
2,500,000
࠹࠽ࠟࠩ࡞㘃ߩᄙ᭽ᕈߣ♽⛔↢‛࿾ℂቇ
ၮ⋚⎇ⓥ(B)
ጟᧄቬ⵨
3,200,000
࠹࠾ࠕ⑼᧦⯻ᐜ⯻ᗵᨴኅ⇓୘૕ߩ⼂೎ߦ᦭↪ߥᣂߒ޿ᛛⴚ㐿⊒ߣ࡝ࠬࠢߩ
⹏ଔ߳ߩᔕ↪
ၮ⋚⎇ⓥ(B)
ᐔ↰ ⡡
1,987,405
ᄢဳ㘃ੱ₎ߩઁ⠪ℂ⸃ߣ⥄Ꮖℂ⸃ߦ㑐ߔࠆᲧセࠕࠗ࠻࡜࠶ࠠࡦࠣ⎇ⓥ
ၮ⋚⎇ⓥ(B)
ᳯᧁ⋥ሶ
3,000,000
⛘Ṍູ੃㘃⡺ᱤ⋡ߩ♽⛔⊛૏⟎ߣ࿾ℂ⊛ಽᏓߦߟ޿ߡߩ⎇ⓥ
⎇ⓥಽᜂ㧔ၮ⋚ B㧕
ᐔ↰ ⡡
100,000
⢝ఽᦼ߆ࠄߩࠦࡒࡘ࠾ࠤ㧙࡚ࠪࡦ㧙⡬ⷡ⚻㛎ߣࠢࡠࠬࡕ࠳࡞⍮ⷡߩᲧセ⊒
㆐⎇ⓥ
⎇ⓥಽᜂ㧔ၮ⋚ B㧕
320,000
ࡏࡁࡏࠍਛᔃߣߔࠆࡅ࠻਄⑼㔤㐳㘃ߩ╭㛽ᩰ᭴ㅧ߆ࠄ⺒ߺ⸃ߊⅣႺㆡᔕ
150,000
ࡏࡁࡏࠍਛᔃߣߔࠆࡅ࠻਄⑼㔤㐳㘃ߩ╭㛽ᩰ᭴ㅧ߆ࠄ⺒ߺ⸃ߊⅣႺㆡᔕ
592,000
ࠠ࠷ࡀࠩ࡞㘃ߩ↢ᵴผߩㅴൻߦ㑐ߔࠆ␠ળ↢ᘒቇ⊛࡮ㆮવቇ⊛⎇ⓥ
⎇ⓥಽᜂ㧔ၮ⋚ B㧕
ᐔ㦮㍈⍫
ᳯᧁ⋥ሶ
Ꮉᧄ ⧐
㚅↰໪⾆
ᱜ㜞ା↵
500,000
⊒㆐㓚ኂߦ߅ߌࠆ⣀ᒙᕈߣ࿁ᓳᕈߩᬌ⸛ߣ‫ߦࠇߘޔ‬ᔕߕࠆ୘೎ᡰេᴺߩ㐿⊒
⎇ⓥಽᜂ㧔ၮ⋚ B㧕
MA Huffman
500,000
᫪ᨋ⊒↢ੱ₞౒ㅢࡑ࡜࡝ࠕߩ↢ᘒቇ␠ળቇ⊛⎇ⓥ
ၮ⋚⎇ⓥ(C)
㜞੗ᱜᚑ
500,000
╙྾♿ߩ࠾ࡎࡦࠩ࡞ߩㅴൻߦ㑐ߔࠆฎ↢‛ቇ⊛⎇ⓥ
ၮ⋚⎇ⓥ(C)
᣿㉿ብᢥ
700,000
ᣂⷙ㔤㐳㘃ࡕ࠺࡞ࠍ↪޿ߚᕆᕈᘟᕈ㧯ဳ⢄Ἳߦ߅ߌࠆ⥄ὼ఺∉ߩᗧ⟵ߦ㑐
ߔࠆ⎇ⓥ
⎇ⓥಽᜂ㧔ၮ⋚ B㧕
⎇ⓥಽᜂ㧔ၮ⋚ B㧕
⎇ⓥಽᜂ㧔ၮ⋚ B㧕
1,700,000
ࡅ࠻ߣࠨ࡞ߦ߅ߌࠆⴕേવᨴߣᮨ୮ߩ⿠Ḯߦ㑐ߔࠆታ㛎⊛⎇ⓥ
㧙㧙
ၮ⋚⎇ⓥ(C)
ች࿾㊀ᒄ
1,000,000
Ꮑ✺ߥㆇേ೙ᓮߩၮ⋚ߣߥࠆㆇേ㑐ㅪ⊹⾰ߩ↢ᓟ⊒㆐ߩ␹⚻⸃೬࡮␹⚻↢
ℂቇ⊛⎇ⓥ
ၮ⋚⎇ⓥ(C)
Ꮉᧄ ⧐
1,200,000
ࡅࡑ࡜ࡗጊ࿾࡮ࠗࡦ࠼੝ᄢ㒽࡮ࠬ࡝࡜ࡦࠞፉᎩߦ߅ߌࠆࡑࠞࠢߩ♽⛔࿾ℂ
ߣಽ㘃
⎇ⓥಽᜂ㧔ၮ⋚ C㧕
ᯅᧄ⵨ሶ
110,000
㖧࿖಴࿯ฎੱ㛽ߩᒻ⾰ੱ㘃ቇ⊛⎇ⓥ
ၮ⋚⎇ⓥ(C)
ᄢ⍹㜞↢
2,000,000
㔤㐳㘃⣖ㇱ૏․⇣⊛ォ౮࿃ሶߩ⊒⃻⸃ᨆ
ၮ⋚⎇ⓥ(C)
ᣧᎹᢅਯ
1,700,000
ࡅ࠻․⇣⊛ߥ⊒⃻࡮ᯏ⢻ߩᄌൻࠍ␜ߔࠪࠕ࡞㉄ฃኈ૕ Siglec-11 ߩㅴൻ
ၮ⋚⎇ⓥ(C)
ศ↰෹ᢎ
2,000,000
㊁↢ᄢဳ㔤㐳㘃ߦ߅ߌࠆੱ₞౒ㅢᗵᨴ∝ߩ․ቯߣ࡝ࠬࠢࠕ࠮ࠬࡔࡦ࠻ߦ㑐
ߔࠆ⎇ⓥ
᜸ᚢ⊛⪚⧘⎇ⓥ
෹᳗㓷Ꮖ
1,300,000
Ყセ⹺⍮ࠥࡁࡓ⑼ቇߩ⏕┙ߦะߌߡᲧセ⹺⍮⑼ቇߣᲧセࠥࡁࡓ⑼ቇߩࠢ
ࡠࠬ࠻࡯ࠢ
᜸ᚢ⊛⪚⧘⎇ⓥ
੗਄⻞৻
1,200,000
ᡷᄌ⁅›∛࠙ࠗ࡞ࠬࡌࠢ࠲࡯ߦࠃࠆᣂߒ޿␹⚻࿁〝࠻࡟࡯ࠪࡦࠣᚻᴺߩ㐿⊒
᜸ᚢ⊛⪚⧘⎇ⓥ
ਛ᧛స᮸
1,400,000
࠮ࡠ࠻࠾ࡦᰳਲᴺߦࠃࠆ㔤㐳㘃߁ߟ∛ࡕ࠺࡞૞಴ߩ⹜ߺ
ጟᧄቬ⵨
200,000
ᄙ൮⯻∝ߦ㑐ߔࠆ㕖㐿⣻⊛ᴦ≮ᴺ߳ߩ᜸ᚢߣ⹏ଔᴺߩ㐿⊒
⊒㆐㓚ኂߩᒝߺ㧔stregth㧕ߩታ㛎⊛ᬌ⸽
⎇ⓥಽᜂ
㧔᜸ᚢ⊛⪚⧘⎇ⓥ㧕
᜸ᚢ⊛⪚⧘⎇ⓥ
ᱜ㜞ା↵
700,000
᜸ᚢ⊛⪚⧘⎇ⓥ
ඨ⼱๋㇢
2,400,000
࠾ࡎࡦࠩ࡞ߩࠠࡁࠦ㘩ࠍㅢߓߚ⩶㘃ߩ⢩ሶᢔᏓᯏ᭴ߩ⸃᣿
᜸ᚢ⊛⪚⧘⎇ⓥ
ฎ⾐┨ᒾ
1,700,000
ࡅ࠻ߢߩ᭴ᚑ⊛ࡋ࠹ࡠࠢࡠࡑ࠴ࡦߩᶖᄬ㧦ᯏ᭴⸃᣿߳ะߌߚ∥〔ㇱಽߩ᭴
ㅧ⸃ᨆ
⧯ᚻ⎇ⓥ(A)
ඨ⼱๋㇢
2,000,000
᧲ධࠕࠫࠕᾲᏪᨋߢߩ৻ᢧ⚿ታߦኻߔࠆᄢဳേ‛ߩ෻ᔕ
⧯ᚻ⎇ⓥ(B)
᧛੗഼⵨
446,451
ήੱፉߦ⒖౉ߐࠇߚ࠹ࡦࠣࠩ࡞୘૕⟲ߣ㊁↢୘૕⟲ߩ଻ోߦ㑐ߔࠆ⎇ⓥ
⧯ᚻ⎇ⓥ(B)
⿷┙ᐞ⏴
800,000
␠ળ⊛⹺⍮⢻ജ߳ߩᲧセ⹺⍮⑼ቇ⊛ࠕࡊࡠ࡯࠴
⧯ᚻ⎇ⓥ(B)
ᴰ ᣿ብ
900,000
㔤㐳㘃ߩ㖸ჿ੤឵ߦ߅ߌࠆᤨ㑆⊛ⷙೣᕈߩ⎇ⓥ
⧯ᚻ⎇ⓥ(B)
᧲Ớ◊ᓼ
900,000
㔤㐳㘃ߦ߅ߌࠆࠬ࠻࡟ࠬⰮ⊕⾰ߩ⸃ᨆ
⧯ᚻ⎇ⓥ(B)
ㇹ ᐽᐢ
1,700,000
ࡅ࠻ߣ࠴ࡦࡄࡦࠫ࡯ߩᲧセ࠻࡜ࡦࠬࠢ࡝ࡊ࠻࡯ࡓ࡮ࡔ࠴ࡠ࡯ࡓ⎇ⓥ
⧯ᚻ⎇ⓥ(B)
ㄞ ᄢ๺
1,300,000
ᄢဳູ੃㘃ߦࠃࠆ⒳ሶᢔᏓ․ᕈߩ࿾ၞ㑆Ყセ㧦ࡑࠞࠢዻߩ㔤㐳㘃ࠍኻ⽎ߦ
⧯ᚻ⎇ⓥ(B)
⧯ᚻ⎇ⓥ(B)
દ᧛⍮ሶ
1,000,000
㔤㐳㘃ߦ߅ߌࠆⷞⷡᖱႎߩᤨ㑆⊛⛔วㆊ⒟ߦ㑐ߔࠆᲧセ⹺⍮⊒㆐⊛ᬌ⸛
ᨋ ⟤㉿
1,000,000
‛ߩᠲ૞߆ࠄ⷗ߚࡅ࠻ࠍ฽߻㔤㐳㘃ߩᲧセ⹺⍮⊒㆐
⧯ᚻ⎇ⓥ(B)
᧻੗ ᷕ
1,800,000
㔤㐳㘃ߦ߅ߌࠆᗵⷡฃኈ૕ㆮવሶߩ⿠ḮߣⅣႺᔕ╵ߩㅴൻ
⎇ⓥᵴേࠬ࠲࡯࠻ᡰេ
ጊᧄ⌀਽
1,140,000
࠴ࡦࡄࡦࠫ࡯ߣࡏࡁࡏߦ߅ߌࠆ೑ઁᕈ࡮੕ᕺᕈ࡮ઁ⠪ℂ⸃ߩᬌ⸛
⎇ⓥᵴേࠬ࠲࡯࠻ᡰេ
ୖጟᐽᴦ
1,060,000
ࡅ࠻ࠍ฽߻㔤㐳㘃ߦ߅ߌࠆ␠ળ⊛ࡄ࡯࠻࠽࡯ㆬᛯߦߟ޿ߡߩ⒳㑆Ყセߣߘ
ߩ⣖␹⚻ၮ⋚
․೎⎇ⓥຬᅑബ⾌
⁚㊁ᢥᶈ
700,000
࠴ࡦࡄࡦࠫ࡯ߩᖱേⴕേߦ㑐ߔࠆᲧセ⹺⍮࡮ⴕേቇ⊛⎇ⓥ
․೎⎇ⓥຬᅑബ⾌
౗ሶፄ᣿
700,000
࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ‫ⴕޟ‬ὑߔࠆ⥄Ꮖ‫⥄⼂⹺ߩޠ‬Ꮖ⹺⼂ߩᲧセ⹺⍮⑼ቇ
⊛⎇ⓥ
․೎⎇ⓥຬᅑബ⾌
દ⮮␭ᐽ
700,000
⺒ߺᦠ߈ߦ㑐ߔࠆቇ⠌࿎㔍ߦኻᔕߒߚ㨑࡜࡯࠾ࡦࠣߦࠃࠆ≮⢒ߣߘߩታ⸽
⊛⹏ଔᴺߩ᭴▽
․೎⎇ⓥຬᅑബ⾌
㚅↰㧔ᧁ႐㧕␞ሶ
․೎⎇ⓥຬᅑബ⾌
㡨ᳰ⩿଻
301,544
㔤㐳㘃ߩ࡝࠭ࡓ೙ᓮߦ߅ߌࠆ␹⚻ࡔࠞ࠾࠭ࡓߩ⸃᣿
․೎⎇ⓥຬᅑബ⾌
ጊ᪸⵨⟤
700,000
࠴ࡦࡄࡦࠫ࡯ߩⴕേߦၮߠ޿ߚᕈᩰ⹏ቯᣇᴺߩ㐿⊒ߣㆮવ⊛ၮ⋚ߩᬌ⸛
․೎⎇ⓥຬᅑബ⾌
Ỉ↰᥏ሶ
700,000
ደਭፉߦ߅ߌࠆࠠࡁࠦߩᄙ᭽ᕈߣ࠾ࡎࡦࠩ࡞ߩ㘩༵ᅢᕈߦߟ޿ߡߩㅴൻ↢
ᘒቇ⊛⎇ⓥ
․೎⎇ⓥຬᅑബ⾌
દ⮮ Პ
700,000
㔤㐳㘃ߦ߅ߌࠆ㗡㛽ᒻᘒߩ᳇୥ⅣႺㆡᔕ
․೎⎇ⓥຬᅑബ⾌
᧻↰৻Ꮧ
800,000
㔤㐳㘃␠ળߩ㊀ጀ᭴ㅧߩ⸃᣿㧦࠹ࡦࠣࠩ࡞ߩ⒳ౝᄌ⇣
․೎⎇ⓥຬᅑബ⾌
ᄢ⼱ᵗ੺
700,000
࠾ࡎࡦࠩ࡞㓶ߣ⟲ࠇߩⓨ㑆㈩⟎ߣណ㘩‫❥ޔ‬ᱺᚢ⇛
․೎⎇ⓥຬᅑബ⾌
ዊᎹ⹞ਫ
700,000
⊒㆐㓚ኂߩ⹺⍮⊛․ᕈߩ⸃᣿ߣᡰេᣇᴺߩᬌ⸛
․೎⎇ⓥຬᅑബ⾌
ᩊፒోᒄ
600,000
ࡅ࠻ߩਇ቟㓚ኂߩࡔࠞ࠾࠭ࡓ⸃᣿ߦะߌߚേ‛ࡕ࠺࡞ߩ᭴▽
․೎⎇ⓥຬᅑബ⾌
CD Dahl
900,000
࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ㗻⹺⼂߅ࠃ߮ࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦାภߩಣℂߩ⸃᣿
ߦะߌߡ
ว ⸘
71 ઙ
1,000,000
㔤㐳㘃ߦ߅ߌࠆᕈߩ⹺⍮ߣᕈࡎ࡞ࡕࡦߩᓇ㗀
236,357,400
̪㊄㗵ߪ‫ޔ‬㑆ធ⚻⾌ࠍ㒰ߊ
㧙㧙
(5) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕ෘ↢ഭ௛⋭⑼ቇ⎇ⓥ⾌⵬ഥ㊄ ౝ⸶৻ⷩ
⎇ⓥ⒳೎
⎇ⓥઍ⴫⠪
㔍ᴦ࡮৻⥸
㜞↰᣽ᒾ
㧔ಽᜂ㧕
㊄ 㗵
⎇ ⓥ ⺖ 㗴
1,000,000
⢄Ἳ࡮৻⥸
᣿㉿ብᢥ
㧔ಽᜂ㧕
12,000,000
⢄Ἳ࠙ࠗ࡞ࠬࡢࠢ࠴ࡦታ↪ൻߩߚ߼ߩၮ␆⊛⎇ⓥ
᡽╷ഃ⮎࡮৻⥸
᣿㉿ብᢥ
24,000,000
HIV-1 ᗵᨴ࡮⊒∝㔤㐳㘃ࡕ࠺࡞⎇ⓥ㧦ኋਥౝ࿃ᕈ෸߮₪ᓧ఺∉⸃ᨆ
ߦၮߠߊ೨⥃ᐥ⹏ଔࠪࠬ࠹ࡓߩᦨㆡൻ
ว ⸘
3ઙ
37,000,000
ࠕࠗࠞ࡞࠺ࠖ-ࠧ࡯࠹ࠖࠛ∝୥⟲╬ߩࡆࠝࡊ࠹࡝ࡦઍ⻢⇣Ᏹࠍ઻߁
∔ᖚߩ⸻ᢿᣇᴺ⏕┙߅ࠃ߮ᴦ≮ᴺ㐿⊒ߩߚ߼ߩᮮᢿ⊛⎇ⓥ
̪㊄㗵ߪ‫ޔ‬㑆ធ⚻⾌ࠍ㒰ߊ
(6) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕⎇ⓥ᜚ὐᒻᚑ⾌╬⵬ഥ㊄‫ ࡞ࡃ࡯ࡠࠣޝ‬COE ࡊࡠࠣ࡜ࡓ‫ޞ‬ౝ⸶৻ⷩ
⎇ⓥ⒳೎
⎇ⓥઍ⴫⠪
ࠣࡠ࡯ࡃ࡞ COE ࡊࡠࠣ࡜ࡓ
ᱜ㜞ା↵
㊄ 㗵
18,770,000
ว ⸘
1ઙ
18,770,000
⎇ ⓥ ⺖ 㗴
↢‛ᄙ᭽ᕈߣㅴൻ⎇ⓥߩߚ߼ߩ᜚ὐᒻᚑ
̪⋥ធ⚻⾌ߩߺ
(7) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕࿖㓙ൻ᜚ὐᢛ஻੐ᬺ⾌⵬ഥ㊄‫ ࡞ࡃ࡯ࡠࠣޝ‬30 ࡊࡠࠣ࡜ࡓ‫ޞ‬ౝ⸶৻ⷩ
੐ᬺฬ⒓
ࠦ࡯ࠬ㐳
㊄ 㗵
੩ㇺᄢቇᰴ਎ઍ࿾⃿␠ળ࡝࡯࠳࡯
⢒ᚑࡊࡠࠣ࡜ࡓ㧔K.U.PROFILE㧕
᧻ᴛື㇢
15,300,000
ว ⸘
1ઙ
15,300,000
ࠦ ࡯ ࠬ ฬ
࿖㓙㔤㐳㘃ቇ࡮㊁↢േ‛ࠦ࡯ࠬ
̪⋥ធ⚻⾌ߩߺ
(8) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕
⎇ⓥ㐿⊒ᣉ⸳౒↪╬ଦㅴ⾌⵬ഥ㊄‫ޞ࠻ࠢࠚࠫࡠࡊ࡮ࠬ࡯࠰࡝ࠝࠗࡃ࡮࡞࠽࡚ࠪ࠽ޝ‬ౝ⸶৻ⷩ
⎇ⓥ⒳೎
NBR
GAIN㧔ᄢဳ㘃ੱ₎ᖱႎࡀ࠶࠻ࡢ࡯ࠢ㧕
ߩᵴേ
ว ⸘
⎇ⓥઍ⴫⠪
ጟᧄቬ⵨
㊄ 㗵
52,000,000
᧻ᴛື㇢
9,600,000
2ઙ
61,600,000
⎇ ⓥ ⺖ 㗴
ᄢဳ㘺⢒ᣉ⸳ߢߩ࠾ࡎࡦࠩ࡞ߩ❥ᱺ࡮⢒ᚑ੐ᬺ
ᄢဳ㘃ੱ₎ߩᖱႎᢛ஻ߣࡀ࠶࠻ࡢ࡯ࠢߠߊࠅ
̪⋥ធ⚻⾌ߩߺ
(9) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕ᦨవ┵⎇ⓥ㐿⊒ᚢ⇛⊛ᒝൻ⾌⵬ഥ㊄ ౝ⸶৻ⷩ
⎇ⓥ⒳೎
⎇ⓥઍ⴫⠪
㊄ 㗵
ᦨవ┵⎇ⓥၮ⋚੐ᬺ
᧻ᴛື㇢
500,000,000
ว ⸘
1ઙ
500,000,000
⎇ ⓥ ⺖ 㗴
ᔃߩవ┵⎇ⓥߩߚ߼ߩㅪ៤᜚ὐ㧔WISH㧕᭴▽
̪⋥ធ⚻⾌ߩߺ
(10) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕⧯ᚻ⎇ⓥ⠪ᚢ⇛⊛ᶏᄖᵷ㆜੐ᬺ⾌⵬ഥ㊄ ౝ⸶৻ⷩ
⎇ⓥ⒳೎
⎇ⓥઍ⴫⠪
㊄ 㗵
㗡⣖ᓴⅣࠍടㅦߔࠆ⧯ᚻ⎇ⓥ⠪
ᶏᄖᵷ㆜ࡊࡠࠣ࡜ࡓ
ᐔ੗໪ਭ
14,330,000
ว ⸘
1ઙ
14,330,000
⎇ ⓥ ⺖ 㗴
ੱ㑆ࠄߒߐߩ㔤㐳㘃⊛⿠Ḯࠍߐߋࠆᚢ⇛⊛࿖㓙
౒หవ┵⎇ⓥ
̪㊄㗵ߪ‫ޔ‬੐ᬺ▤ℂ⚻⾌ࠍ㒰ߊ
(11) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕⑼ቇᛛⴚ✚วផㅴ⾌⵬ഥ㊄ ౝ⸶৻ⷩ
⎇ⓥ⒳೎
⎇ⓥઍ⴫⠪
࿖㓙౒ห⎇ⓥߩផㅴ
ጟᧄቬ⵨
㊄ 㗵
4,093,780
ว ⸘
1ઙ
4,093,780
⎇ ⓥ ⺖ 㗴
㔍ᴦᕈነ↢⯻∛ߦ㑐ߔࠆㆮવሶ⸻ᢿᴺߩ㐿⊒
̪㊄㗵ߪ‫ޔ‬ⅣႺᡷༀ⾌ࠍ㒰ߊ
㧙㧙
(12) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕⎇ⓥ⠪ᶏᄖᵷ㆜ၮ㊄ഥᚑ㊄ ౝ⸶৻ⷩ
⎇ⓥ⒳೎
⎇ⓥઍ⴫⠪
㊄ 㗵
⎇ ⓥ ⺖ 㗴
⚵❱⊛ߥ⧯ᚻ⎇ⓥ⠪╬ᶏᄖ
ᵷ㆜ࡊࡠࠣ࡜ࡓ
᧻ᴛື㇢
25,795,787
ว ⸘
1ઙ
25,795,787
ੱ㑆ߩᧄᕈߩㅴൻ⊛⿠Ḯߦ㑐ߔࠆవ┵⎇ⓥ
㧔AS-HOPE㧕
̪㊄㗵ߪ‫▤⥸৻ޔ‬ℂ⾌ࠍ㒰ߊ
(13) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕వ┵⎇ⓥഥᚑၮ㊄ഥᚑ㊄ ౝ⸶৻ⷩ
⎇ⓥ⒳೎
⎇ⓥઍ⴫⠪
ᦨవ┵࡮ᰴ਎ઍ⎇ⓥ㐿⊒ᡰេ
ࡊࡠࠣ࡜ࡓ
᧻ᧄᱜᐘ
42,745,000
ᗧ᰼ࠍ↢ߺ಴ߔ␹⚻ࡔࠞ࠾࠭ࡓߩ⸃᣿㧦೨㗡೨㊁
߳ߩਛ⣖࠼࡯ࡄࡒࡦ౉ജߩᓎഀ
ᦨవ┵࡮ᰴ਎ઍ⎇ⓥ㐿⊒ᡰេ
ࡊࡠࠣ࡜ࡓ
૒⮮ ᒎ
22,750,000
ᐢ᳢ᕈ⊒㆐㓚ኂߦ߅ߌࠆኻੱ⋧੕૞↪㓚ኂߩᔃ
ℂ␹⚻ၮ⋚ߩ⛔ว⊛⸃᣿
ว ⸘
2ઙ
65,495,000
㊄ 㗵
⎇ ⓥ ⺖ 㗴
̪㊄㗵ߪ‫ޔ‬㑆ធ⚻⾌ࠍ㒰ߊ
(14) 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕ነ㒝㊄ ౝ⸶৻ⷩ
ነ㒝㊄ฬ⒓╬
⎇ⓥઍ⴫⠪
㊄ 㗵
ነ 㒝 ߩ ⋡ ⊛
ᣩ⎣ሶ⽷࿅
⷏᧛ ೰
2,000,000
㜟᪸ቇⴚᅑബၮ㊄
ᯅᧄ⵨ሶ
285,000
㖧࿖࡮㊄ቭૄ⡍ㆮ〔⟲ߩⴊ✼㑐ଥࠍਅ㗶㛽ߣᱤ
߆ࠄផ᷹ߔࠆ
⮮ේ࠽࠴ࡘ࡜࡞ࡅࠬ࠻࡝࡯⽷࿅
᧻↰৻Ꮧ
580,092
ࡏ࡞ࡀࠝ↥㔤㐳㘃ߩ↢ᵴผ⎇ⓥ㧦৻ᢧ㐿⧎߇❥
ᱺߦਈ߃ࠆᓇ㗀
ว ⸘
3ઙ
ੱ㘃ߩ๭ๆᯏ⢻ߩㅴൻߣ╙৻ᰴ಴ࠕࡈ࡝ࠞߦ㑐
ߔࠆ⎇ⓥ
2,865,092
̪ነ㒝㊄㗵ߪ‫ోޔ‬ቇ౒ㅢ⚻⾌㧔2㧑㧕߅ࠃ߮ㇱዪਛᄩ▤ℂ⾌㧔3㧑㧕ࠍប㒰ߒߚ㊄㗵
3. ࿑ᦠ
㔤㐳㘃ቇߩ⎇ⓥᚑᨐࠍ✂⟜ߔࠆᣇ㊎ߢ࿑ᦠࠍ෼㓸ߒߡ޿߹ߔ‫ߦ․ޕ‬㔤㐳㘃ቇ㑐ㅪ⺰ᢥߩ೎೚ߪ 85,000 ὐߦ㆐
ߒ‫ޡޔ‬㔤㐳㘃ቇ೎೚ࠦ࡟࡚ࠢࠪࡦ‫ߡߒߣޢ‬㑛ⷩߦଏߒߡ޿߹ߔ‫ోߪߡ޿ߟߦ☋ᦠޕ‬ᚲຬ߆ࠄߩផ⮈ࠍฃߌઃߌ‫ޔ‬
ㆬቯߩෳ⠨ߦߒߡ޿߹ߔ‫ޕ‬
(1) ⬿ᦠᢙ㧔2012 ᐕ 3 ᦬ᧃ⃻࿷㧕
๺ᦠ㧦 7,835 ౠ㧔⵾ᧄ㔀⹹߽฽߻㧕
ᵗᦠ㧦17,746 ౠ㧔⵾ᧄ㔀⹹߽฽߻㧕
๺㔀⹹࡮ਛ࿖㔀⹹㧦147 ⹹
ᵗ㔀⹹㧦372 ⹹
♿ⷐ㘃㧦650 ⹹
㔤㐳㘃ቇ㑐ㅪ೎೚㧔㔤㐳㘃ቇ೎೚ࠦ࡟࡚ࠢࠪࡦ㧕㧦85,000 ὐ
(2) ⾗ᢱߩᚲ⬿ᬌ⚝
࿑ᦠቶߢᚲ⬿ߒߡ޿ࠆ࿑ᦠ࡮㔀⹹ߪߔߴߡ‫ޣ‬੩ㇺᄢቇ⬿ᦠᬌ⚝ KULINE‫ߢޤ‬ᬌ⚝ߢ߈߹ߔ‫ޕ‬
‫ޣ‬੩ㇺᄢቇ⬿ᦠᬌ⚝ KULINE‫ޔߒࠬ࠮ࠢࠕߦޤ‬
㨇⹦⚦ᬌ⚝↹㕙㨉㧙㨇ᚲ⬿㙚㨉ߩᰣߢ㨇(82)㔤㐳⎇㨉ࠍㆬᛯߔࠆ
ߣ‫ޔ‬㔤㐳㘃⎇ⓥᚲߩ⬿ᦠߩߺࡅ࠶࠻ߒ߹ߔ‫ޕ‬
⹦ߒߊߪ੩ㇺᄢቇ࿑ᦠ㙚ᯏ᭴ߩࡎ࡯ࡓࡍ࡯ࠫࠍߏⷩਅߐ޿‫ޕ‬http://www.kulib.kyoto-u.ac.jp/
㔤㐳㘃ቇ㑐ㅪ೎೚㧔㔤㐳㘃ቇ೎೚ࠦ࡟࡚ࠢࠪࡦ㧕ߪ‫ޣ‬㔤㐳㘃ቇᢥ₂⚝ᒁ࠺࡯࠲ࡌ࡯ࠬ‫ߢޤ‬ᬌ⚝ߢ߈߹ߔ‫ޕ‬
㔤㐳㘃⎇ⓥᚲࡎ࡯ࡓࡍ࡯ࠫߩ topics‫ޣ‬㔤㐳㘃ቇᢥ₂⚝ᒁ࠺࡯࠲ࡌ࡯ࠬ‫ⷩߏࠍޤ‬ਅߐ޿‫ޕ‬
http://www.pri.kyoto-u.ac.jp/cgi-bin/library/books.cgi
㧙㧙 (3) 㔤㐳㘃⎇ⓥᚲ࿑ᦠቶ೑↪ⷙ⒟
㧵㧚㐿ቶᤨ㑆ߣભቶ
1㧚㐿ቶᤨ㑆㧦ᐔᣣ 9 ᤨ߆ࠄ 17 ᤨ߹ߢ‫ޕ‬
2㧚ભቶ㧦࿯ᦐᣣ‫ޔ‬ᣣᦐᣣ‫ޔ‬࿖᳃ߩ␸⑂ᣣ‫ޔ‬ᐕᧃ࡮ᐕᆎ‫ᤨ⥃ߩઁߩߘޕ‬ભቶߪ‫ߩߘޔ‬ㇺᐲឝ␜ߔࠆ‫ޕ‬
Τ㧚㑛ⷩ
1㧚㑛ⷩ⠪ߩ⾗ᩰ
1) ᧄ⎇ⓥᚲߩᚲຬ‫ޕ‬
2) ᧄ⎇ⓥᚲߩ౒ห೑↪⎇ⓥຬ‫ޕ‬
3) 1)‫ޔ‬2)એᄖߩ‫ޔ‬੩ㇺᄢቇߦᚲዻߔࠆ⠪ߢ‫ޔ‬ᚲዻㇱዪߩ࿑ᦠᣉ⸳߽ߒߊߪ㒝ዻ࿑ᦠ㙚ߩ⚫੺ߩ޽ࠆ⠪‫ޕ‬
4) ߘߩઁ৻⥸೑↪⠪‫ޕ‬
2㧚㑛ⷩ
1) 㑛ⷩߪᚲቯߩ႐ᚲߢⴕࠊߥߌࠇ߫ߥࠄߥ޿‫ޕ‬
2) ᰴߩฦภߦឝߍࠆ႐วߦ߅޿ߡߪ㑛ⷩࠍ೙㒢ߔࠆߎߣ߇ߢ߈ࠆ‫ޕ‬
(1) ᒰ⹥⾗ᢱߦ⁛┙ⴕ᡽ᴺੱ╬ߩ଻᦭ߔࠆᖱႎߩ౏㐿ߦ㑐ߔࠆᴺᓞ㧔ᐔᚑ 13 ᐕᴺᓞ╙ 140 ภ‫ޔ‬એਅ㨬
ᖱႎ౏㐿ᴺ㨭ߣ޿߁‫ޕ‬㧕╙ 5 ᧦╙ 1 ภ‫ ╙ޔ‬2 ภ෸߮╙ 4 ภࠗߦឝߍࠆᖱႎ߇⸥㍳ߐࠇߡ޿ࠆߣ⹺߼
ࠄࠇࠆ႐วߦ߅ߌࠆᒰ⹥ᖱႎ߇⸥㍳ߐࠇߡ޿ࠆㇱಽ‫ޕ‬
(2) ᒰ⹥⾗ᢱߩోㇱ෶ߪ৻ㇱࠍ৻ቯߩᦼ㑆౏ߦߒߥ޿ߎߣࠍ᧦ઙߦ୘ੱ෶ߪᖱႎ౏㐿ᴺ╙ 5 ᧦╙ 2 ภ
ߦⷙቯߔࠆᴺੱ╬߆ࠄነ⿅෶ߪነ⸤ࠍฃߌߡ޿ࠆ႐วߦ߅ߌࠆᒰ⹥ᦼ㑆߇⚻ㆊߔࠆ߹ߢߩ㑆‫ޕ‬
(3) ᒰ⹥⾗ᢱߩේᧄࠍ೑↪ߐߖࠆߎߣߦࠃࠅᒰ⹥ේᧄߩ⎕៊⧯ߒߊߪߘߩᳪ៊ࠍ↢ߓࠆ߅ߘࠇ߇޽ࠆ
႐ว෶ߪᒰ⹥⾗ᢱ߇⃻ߦ૶↪ߐࠇߡ޿ࠆ႐ว‫ޕ‬
Υ㧚⾉಴෸߮㄰ළ
1㧚II ߩ 1 ߩ 1㧕ߩ⹥ᒰ⠪෸߮‫ޔ‬2㧕ߩ߁ߜ੍߼೑↪⠪ࠞ࡯࠼ࠍឭ಴ߒߚ⠪ߪ‫ޔ‬ਅ⸥ߦᓥ޿࿑ᦠࠍ୫↪ߢ߈ࠆ‫ޕ‬
ߔߴߡߩ୫↪⾗ᢱߪ‫ޔ‬ේೣߣߒߡᚲᄖߦᜬߜ಴ߔߎߣߪߢ߈ߥ޿‫ޕ‬
1) ୫↪⾗ᢱߩ⒳㘃ߣ୫↪ᣇᴺ
a㧚නⴕᧄ
(i) නⴕᧄߪ 1 㩀᦬㑆୫↪ߢ߈ࠆ‫ޕ‬
(ii) ୫↪ᤨߦߪ‫࠼࡯ࠞࠢ࠶ࡉޔ‬෸߮ઍᧄ᧼↪⚕ߦᔅⷐ੐㗄ࠍ⸥౉ߔࠆ‫ߪ࠼࡯ࠞࠢ࠶ࡉޕ‬ᚲቯߩ▫ߦ౉
ࠇ‫ޔ‬ઍᧄ᧼↪⚕ߪઍᧄ᧼ߩ⢛ߦᝌ౉ߒߡ‫ᦠޔ‬᨞਄ߩᧄߩ޽ߞߚ૏⟎ߦ⟎ߊ‫ޕ‬
b㧚⵾ᧄ㔀⹹
(i) ⵾ᧄ㔀⹹ߪ 3 ᣣ㑆୫↪ߢ߈ࠆ‫ޕ‬
(ii) ୫↪ᣇᴺߪනⴕᧄߦḰߓࠆ‫ޕ‬
c㧚ᧂ⵾ᧄ㔀⹹
(i) ᧂ⵾ᧄߩ㔀⹹ߪ 15 ᤨ߆ࠄ⠉ᦺ 10 ᤨ߹ߢߩ㑆ߦ㒢ࠅ୫↪ߢ߈ࠆ‫ޕ‬
(ii) ୫↪ᤨߦߪ⾉಴ࠞ࡯࠼ߦᔅⷐ੐㗄ࠍ⸥౉ߔࠆ‫ޕ‬
d㧚೎೚
(i) ೎೚ߪ㐿ቶᤨ㑆ਛߦ࿑ᦠቶౝߢߩߺ೑↪ߢ߈ࠆ‫ޕ‬
(ii) ೑↪ᓟߪ‫ޔ‬㄰ළบߩ▫ߦ㄰ළߔࠆ‫ޕ‬
e㧚ઁᯏ㑐߆ࠄߩ୫↪⾗ᢱ
(i) ઁᯏ㑐߆ࠄߩ୫↪⾗ᢱߪ‫ޔ‬㐿ቶᤨ㑆ਛߩ࿑ᦠቶౝߢߩ೑↪ߦ㒢ࠆ‫ޕ‬
(ii) ೑↪ᓟߪ࿑ᦠଥຬߦ㄰ළߔࠆ‫ޕ‬
2) ෳ⠨࿑ᦠߘߩઁ⑌Ꮺ಴ᛒ޿ߩ࿑ᦠߪ⾉಴ߐߥ޿‫ޕ‬
3) ୫↪ਛߩ⾗ᢱࠍォ⾉ߒߡߪߥࠄߥ޿‫ޕ‬
4) ౣᚻ⛯߈ࠍߔࠆߎߣߦࠃࠅ⾉಴ᦼ㒢ߩᑧ㐳߇ߢ߈ࠆ‫ޕ‬
ߚߛߒ‫ߦઁޔ‬୫↪Ꮧᦸ⠪߇޽ࠆᤨߪ‫ࠍઁޔ‬ఝవߔࠆ‫ޕ‬
5) ୫↪ᓟߩ࿑ᦠߪ㄰ළบߦ㄰ළߔࠆ‫ޕ‬
2㧚II ߩ 1 ߩ 3) ߩ⹥ᒰ⠪ߪ‫ޔ‬ᚲዻㇱዪߩ࿑ᦠᣉ⸳߽ߒߊߪ㒝ዻ࿑ᦠ㙚ࠍㅢߓߡ୫↪ࠍଐ㗬ߔࠆߎߣ߇ߢ߈ࠆ‫ޕ‬
1) ୫↪⾗ᢱߪනⴕᧄߩߺߢ‫ޔ‬ᚲዻㇱዪ࿑ᦠᣉ⸳ౝ߽ߒߊߪ㒝ዻ࿑ᦠ㙚ౝߢߩ೑↪ߦ㒢ࠆ‫ޕ‬
2) ୫↪ᦼ㒢ߪ 2 ㅳ㑆ߣߔࠆ߇‫⎇ᧄޔ‬ⓥᚲຬ߆ࠄߩⷐ⺧߇޽ߞߚ႐วߦߪ‫ޔ‬୫↪ᦼ㒢ౝߢ޽ߞߡ߽‫ޔ‬ㅦ߿
㧙㧙
߆ߦ㄰ළߔࠆߎߣߣߔࠆ‫ޕ‬
Φ㧚✚ὐᬌ෸߮㐳ᦼ⾉಴
1㧚ቯᦼ⊛ߦ࿑ᦠߩ✚ὐᬌࠍⴕ߁‫⾉ޔߪᤨߩߎޕ‬಴ᦼ㒢ౝᄖࠍ໧ࠊߕ‫ߩߡߴߔޔ‬࿑ᦠࠍ㄰ළߔࠆ‫ޕ‬
2㧚✚ὐᬌᦼ㑆ਛ‫ޔ‬࿑ᦠቶࠍભቶߣߔࠆߎߣ߇޽ࠆ‫ޕ‬
3㧚࿑ᦠᆔຬળߦࠃࠅ⎇ⓥቶ╬߳ߩ஻߃ઃߌ߇⹺߼ࠄࠇߚᤨߪ‫ޔ‬㐳ᦼ⾉಴ᛒ޿ߣߔࠆ‫ޕ‬㐳ᦼ⾉಴ᦼ㑆ߪ 1 ᐕ
ߢ‫ޔ‬㐳ᦼ⾉಴ᛒ޿ߩᦝᣂߪ✚ὐᬌᤨߦⴕ߁‫ޕ‬
Χ㧚୘ੱᖱႎṳ߃޿㒐ᱛߩߚ߼ߦᔅⷐߥភ⟎
1㧚࿑ᦠቶߪ‫ޔ‬࿑ᦠቶ⾗ᢱߦ୘ੱᖱႎ㧔↢ሽߔࠆ୘ੱߦ㑐ߔࠆᖱႎߢ޽ߞߡ‫ޔ‬ᒰ⹥ᖱႎߦ฽߹ࠇࠆ᳁ฬ‫↢ޔ‬
ᐕ᦬ᣣߘߩઁߩ⸥ㅀ╬ߦࠃࠅ․ቯߩ୘ੱࠍ⼂೎ߔࠆߎߣ߇ߢ߈ࠆ߽ߩ㧔ઁߩᖱႎߣኈᤃߦᾖวߔࠆߎߣ
߇ߢ߈‫․ࠅࠃߦࠇߘޔ‬ቯߩ୘ੱࠍ⼂೎ߔࠆߎߣ߇ߢ߈ࠆߎߣߣߥࠆ߽ߩࠍ฽߻㧕ࠍ޿߁㧕߇⸥㍳ߐࠇߡ
޿ࠆ႐วߦߪ‫ޔ‬ᒰ⹥୘ੱᖱႎߩṳ߃޿ߩ㒐ᱛߩߚ߼ߦᰴߩฦภߦឝߍࠆភ⟎ࠍ⻠ߓࠆ߽ߩߣߔࠆ‫ޕ‬
1) ᦠᐶߩᣉ㍤ߘߩઁߩ‛ℂ⊛ߥធ⸅ߩ೙㒢
2) ࿑ᦠቶ⾗ᢱߦ⸥㍳ߐࠇߡ޿ࠆ୘ੱᖱႎߦኻߔࠆਇᱜࠕࠢ࠮ࠬ㧔ਇᱜࠕࠢ࠮ࠬⴕὑߩ⑌ᱛ╬ߦ㑐ߔࠆᴺ
ᓞ㧔ᐔᚑ 11 ᐕᴺᓞ╙ 128 ภ㧕╙ 3 ᧦╙ 2 㗄ߦⷙቯߔࠆਇᱜࠕࠢ࠮ࠬࠍ޿߁㧕ࠍ㒐ᱛߔࠆߚ߼ߦᔅⷐߥ
ភ⟎
3) ࿑ᦠቶߩ⡯ຬߦኻߔࠆᢎ⢒࡮⎇ୃߩታᣉ
4) ߘߩઁᒰ⹥୘ੱᖱႎߩṳ߃޿ߩ㒐ᱛߩߚ߼ߦᔅⷐߥភ⟎
Ψ㧚ߘߩઁ
1㧚࿑ᦠቶ⾗ᢱߩ⋡㍳෸߮ߎߩ࿑ᦠቶ೑↪ⷙ⒟ߦߟ޿ߡߪᏱᤨ࿑ᦠቶߦ஻߃ઃߌࠆ‫ޕ‬
2㧚⾗ᢱࠍ⚗ᄬߒߚࠅᳪ៊ߒߚ႐วߪ‫ޔ‬ઍᧄ߹ߚߪ⋧ᒰߩઍ㊄ߢ⵬ࠊߥߌࠇ߫ߥࠄߥ޿‫ޕ‬
3㧚୫↪⾗ᢱࠍᦼᣣ߹ߢߦ㄰ළߒߥ߆ߞߚ႐ว‫ޔ‬એᓟߩ⾉಴ࠍ৻ቯᦼ㑆஗ᱛߐࠇࠆߎߣ߇޽ࠆ‫ޕ‬
4㧚࿑ᦠቶౝ㧔ᦠᐶࠍ฽߻㧕ߪ⑌ᾍߣߔࠆ‫ޕ‬
㒝ೣ
ߎߩⷙ⒟ߪ‫ޔ‬ᐔᚑ 16 ᐕ 4 ᦬ 1 ᣣ߆ࠄᣉⴕߔࠆ‫ޕ‬
㒝ೣ
ߎߩⷙ⒟ߪ‫ޔ‬ᐔᚑ 23 ᐕ 4 ᦬ 1 ᣣ߆ࠄᣉⴕߔࠆ‫ޕ‬
4. ࠨ࡞㘃㘺⢒㗡ᢙ࡮േᘒ
2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕ᧃ 㘺⢒㗡ᢙ
⒳ ฬ
㗡 ᢙ
⒳ ฬ
㗡 ᢙ
ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻
131
ࡢ࠲ࡏ࠙ࠪ࠲ࡑ࡝ࡦ
21
ࠞ ࠾ ࠢ ࠗ ࠩ ࡞
࡛
࡞
15
ࡑ
ࡈࠨࠝࡑࠠࠩ࡞
9
ࠕࠫ࡞࠹࠽ࠟࠩ࡞
3
1
࠴ ࡦ ࡄ ࡦ ࠫ ࡯
14
ว ⸘
1147
ࠩ
ࠤ࠽ࠟࠢࡕࠩ࡞
࠾
ࡎ ࡦ
ࠩ
࡞
࠾
ࡎ ࡦ
ࠩ
࡞ (NBR)*
345
ࠕ
ࠞ ࠥ
ࠩ
࡞
219
࠲ ࠗ ࡢ ࡦ ࠩ ࡞
ࡏࡦࡀ࠶࠻ࠩ࡞
ࡦ
࠻ ࡅ
3
23
ࡅ
3
356
4
*NBR㧔‫ࠬ࡯࠰࡝ࠝࠗࡃޠ࡞ࠩࡦࡎ࠾ޟ‬㧕ࡊࡠࠫࠚࠢ࠻ߢ㘺⢒ߒߡ޿ࠆ߽ߩ
㧙㧙 2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕ࠨ࡞㘃േᘒ⴫
Ⴧ‫ޓ‬ട
඙‫ޓ‬ಽ
಴
ዉ
ᷫ‫ޓޓޓޓޓ‬ዋ㧔ᱫ੢ߥߤ㧕
ዊ
ታ ੐ ᄖ ๭ ᶖ ᗵ ᴲ ⴮ ߘ ೬ ᚲ
ๆ ൻ
Ვ ᱫ ᱫ ∔ ∔ ∝ ེ ᒙ
46
52
71
35
4
11
42
5
22
7
4
2
1
10 208 91
208
3
ᖚ ᖚ
ࠦࡕࡦࡑ㧙ࡕ࠮࠶࠻
࠾ࡎࡦࠩ࡞
࠾ࡎࡦࠩ࡞(NBR㧕
ࠕࠞࠥࠩ࡞
ࠞ࠾ࠢࠗࠩ࡞
ࡏࡦࡀ࠶࠻ࠩ࡞
46
42
71
35
4
ዊ‫⸘ޓޓޓޓޓޓ‬
ว‫⸘ޓޓޓޓޓޓ‬
198
10
2
8
1
1
1
1
4
2
1
4
2
7
ᷫ
⢻ ⛎
⸘
౉
⸘
ઁ ਇ ଏ
♽ ♽
↥
Ⴧ
ߩ ᬌ ᄖ
㛎 ᡿ ் ེ ེ ᨴ ዩ
⒳ฬ
ዊ
1
1
3
0
219
11
*
2
14
24
3
1
10
46
44
56
25
72
82
28
8
4
21
-20
-11
7
-4
-4
219
-11
-11
ⴊዊ᧼ᷫዋ∝㑐ㅪߩᱫ੢࡮቟ᭉᲕࠍ฽߻
5. ⾗ᢱ
ᚲ⬿ߔࠆ⾗ᢱߪ‫ޔ‬㛽ᩰᮡᧄ‫ޔ‬ᶧᶐᮡᧄ‫ޔ‬ൻ⍹ᮨဳ‫ޔ‬ㆮવሶ⹜ᢱ‫ޔ‬CT ↹௝ߥߤ߆ࠄߥࠅ‫ޔ‬ᄖㇱߩ⎇ⓥ⠪ߦ߽ၮ
ᧄ⊛ߦߔߴߡ౏㐿ߐࠇߡ޿ࠆ‫⹜⾗ޕ‬ᢱߩᄙߊߪ࠺࡯࠲ࡌ࡯ࠬൻߐࠇߡ߅ࠅ‫⾗ޔ‬ᢱᆔຬળߩ⸵นߦ߽ߣߠ޿ߡ೑
↪Ꮧᦸ⠪ߦឭଏߐࠇ‫⎇ޔ‬ⓥㆀⴕ਄ߩᔅⷐߦᔕߓߡ⾉ߒ಴ߒ߽߅ߎߥߞߡ޿ࠆ‫ޕ‬
(1) 㛽ᩰᮡᧄ㧔⴫ 1‫ޔ‬2㧕
⾗ᢱᆔຬળߩ࠺࡯࠲ࡌ࡯ࠬ㧔PRISK㧕ߦ⊓㍳ߐࠇߚ㔤㐳㘃㛽ᩰᮡᧄߪ 8,600 ὐࠍ⿥߃ࠆ㧔⴫ 1㧕‫ࠩࡦࡎ࠾ߦ․ޕ‬
࡞ߩᮡᧄߪᚲౝ㘺⢒࡮㊁↢↱᧪୘૕ࠍ޽ࠊߖߡ 3,000 ὐએ਄ࠍᢙ߃‫ޔ‬਎⇇⊛ߦ⷗ߡ߽⒘ߥࠦ࡟࡚ࠢࠪࡦߢ޽ࠆ‫ޕ‬
㔤㐳㘃એᄖߦ߽‫ࠬ࡯ࡌ࠲࡯࠺ޔ‬㧔PRISK-Z㧕ߦߪ⚂ 1,700 ὐߩ₞㛽ᮡᧄ߇⊓㍳ߐࠇߡ޿ࠆ㧔⴫ 3㧕‫ޔߦ․ޕ‬ᣣᧄ↥
࠲࠿ࠠ߿࠹ࡦ‫ߩࡑࠣࡢࡁࠠ࠷ޔ‬ᮡᧄᢙߪ਎⇇᦭ᢙߢ޽ࠅ‫ޔ‬ᣣᧄ↥㊁↢ູ੃㘃߇ᷫߞߡ޿ࠆ⃻࿷‫⾆ߪࠄࠇߎޔ‬㊀ߥ
⾗ᢱߣ⸒߃ࠆ‫ޕ‬ᮡᧄߪ⒳ߏߣߦಽ㘃ߐࠇ‫ޔ‬⒳ౝߢߪᮡᧄ⇟ภߦߒߚ߇ߞߡ㈩೉ߐࠇߡ޿ࠆ‫ޕ‬೑↪Ꮧᦸ⠪ߪ‫ޔ‬ᣂ᫟
4 㓏⾗ᢱቶߩࠦࡦࡇࡘ࡯࠲࡯਄ߦ⟎߆ࠇߚᮡᧄ࠺࡯࠲ࡌ࡯ࠬ߆ࠄᮡᧄ⇟ภ‫ޔ‬ዻฬ‫ޔ‬⒳ฬ‫⊓ޔ‬㍳ᣣ‫ޔ‬ᕈ೎‫ޔ‬૕㊀‫ޔ‬
ᐳ㜞‫ޔ‬೨⢵㐳ߥߤߩᖱႎࠍᬌ⚝ߔࠆߎߣ߇ߢ߈ࠆ‫ޕ‬
(2) ᶧᶐᮡᧄ㧔⴫ 1‫ޔ‬3㧕
ᧄ᫟࿾ਅ෸߮ᩙᩘ࿾඙ߩᶧᶐ⾗ᢱቶߦฦ⒳㔤㐳㘃ߩࡎ࡞ࡑ࡝ࡦ߽ߒߊߪࠕ࡞ࠦ࡯࡞ߢ࿕ቯߐࠇߚᶧᶐᮡᧄ߇
⚂ 1,000 ὐ଻▤ߐࠇߡ޿ࠆ㧔⴫ 1㧕
‫ޕ‬㔤㐳㘃એᄖߩᶧᶐᮡᧄ߽⚂ 200 ὐ޽ࠆ㧔⴫ 3㧕‫╭ޕ‬㛽ᩰ♽ߩഀว߇㜞޿߇‫ޔ‬⣖
߿⤳ེߩᮡᧄ߽฽߻‫ޕ‬ᐔᚑ 21 ᐕᐲ߆ࠄᶧᶐᮡᧄߩ࠺࡯࠲ࡌ࡯ࠬൻࠍⴕ޿‫ޔ‬PRISK ߅ࠃ߮ PRISK-Z ߢߩᬌ⚝߇น
⢻ߦߥߞߚ‫ߦ߁ࠃߩߎޕ‬ᄢⷙᮨߥᶧᶐᮡᧄ⾗ᢱߪ਎⇇⊛ߦ⷗ߡ߽⒘᦭ߢ޽ࠅ‫⾆ޔ‬㊀ߥࠦ࡟࡚ࠢࠪࡦߢ޽ࠆ‫ޕ‬
(3) ൻ⍹ᮨဳ
ੱ㘃߅ࠃ߮ਛᣂ਎ࡎࡒࡁࠗ࠼ࠍਛᔃߦ 495 ὐߩൻ⍹ᮨဳ߇࠺࡯࠲ࡌ࡯ࠬ㧔PRICAST㧕ߦ⊓㍳ߐࠇߡ޿ࠆ‫ޕ‬
(4) 㔤㐳㘃ಽሶ↢‛ቇ↪⹜ᢱ㧔⴫ 4㧕
ᐔᚑ 21 ᐕᐲࠃࠅᄢဳ㘃ੱ₎ࡀ࠶࠻ࡢ࡯ࠢ㧔GAIN㧕ߩᖱႎࠍㅢߓߡ⼑ᷰࠍฃߌߚ㘃ੱ₎⤳ེ⹜ᢱ߿‫⎇ޔ‬ⓥᚲౝ
ߢᄙ㊀೑↪ߩኻ⽎ߣߥߞߚ⹜ᢱߩ৻ㇱ㧔ᣥ਎⇇ࠩ࡞‫ޔ‬ᣂ਎⇇ࠩ࡞╬㧕ࠍ⾗ᢱᆔຬળ߇଻▤▤ℂߒߡ޿ࠆ‫ࠄࠇߎޕ‬
ߪ RNAlater ಣℂ⹜ᢱ߿ಓ⚿⹜ᢱ߇ਥߢ޽ࠆ‫⼑ޕ‬ᷰᄾ⚂╬ߩ㑐ଥ߆ࠄ‫ޔ‬ේೣߣߒߡ೑↪ߪᚲຬߣ౒ห೑↪⎇ⓥຬ
ߦ㒢ቯߒߡ޿ࠆߩߢ‫ޔ‬೑↪Ꮧᦸ⠪ߪ㑐ଥᚲຬߦ໧޿วࠊߖߡ޿ߚߛ߈ߚ޿‫⃻ޕ‬࿷࠺࡯࠲ࡌ࡯ࠬൻࠍㅴ߼ߡ޿ࠆ‫ޕ‬
㧙㧙
(5) CT ↹௝
ᚲ⬿ᮡᧄߩ CT ↹௝࠺࡯࠲ߩ࠺࡯࠲ࡌ࡯ࠬൻࠍㅴ߼‫ޔ‬Web ਄ߩ Digital Morphology Museum (http://www.pri.
kyoto-u.ac.jp/dmm/WebGallery/Index.html) ࠍ੺ߒߡ‫↹ޔ‬௝࠺࡯࠲ࠍ࿖ౝᄖߩᄙߊߩ⎇ⓥ⠪ߦឭଏߒߡ޿ࠆ‫ޕ‬
(6) ߘߩઁ
㔤㐳㘃⎇ⓥᚲ⾗ᢱᆔຬળߢߪ‫ޔ‬࿖ౝᄖߩᄙߊߩ⎇ⓥ⠪߇ߎࠇࠄߩ⾗⹜ᢱࠍ೑↪ߒߡ⎇ⓥࠍㅴ߼ࠆߎߣࠍᏗᦸߒ
ߡ߅ࠅ‫ޔߦ߼ߚߩߘޔ‬Ფᐕ 200 ὐએ਄ߩᣂߚߥᮡᧄߩฃߌ౉ࠇߣ૞⵾ࠍⴕ޿‫⹜⾗ޔ‬ᢱߩలታߦദ߼ߡ޿ࠆ‫ޔߚ߹ޕ‬
ᚲ⬿⾗⹜ᢱߩ࠺࡯࠲ࡌ࡯ࠬൻߥߤ೑↪ⅣႺߩᢛ஻ࠍ߽ⴕߞߡ޿ࠆ‫ޕ‬㛽ᩰ࡮ᶧᶐᮡᧄߩ೑↪ᚻ⛯߈ߪ‫ޔ‬㕖⎕უ⊛ߥ
૶↪⋡⊛ߩ႐วߪᲧセ⊛◲ଢߢ‫ޔ‬ᮡᧄߩ⎕უ࡮⎕៊ࠍ઻߁⎇ⓥߢ޽ߞߡ߽‫⾗ޔ‬ᢱᆔຬળ߳ߩలಽߥ⺑᣿ߣߘࠇߦ
ၮߠߊክᩏࠍ⚻ߚ਄ߢน⢻ߥ႐ว߽޽ࠆ‫⾗ߪߕ߹ޕ‬ᢱᆔຬળ߹ߢㅪ⛊ߐࠇߚ޿‫⹜⾗ޕ‬ᢱߦߪቇᄖᣉ⸳߆ࠄ⼑ᷰߐ
ࠇߚ⾆㊀ߥ߽ߩ߽฽߹ࠇࠆ‫ޕ‬೑↪ⷙ⚂߿ᄾ⚂╬ߩㆩ቞߽දജߒߡ޿ߚߛߊࠃ߁߅㗿޿ߔࠆ‫ޕ‬
஻⠨㧦⾗ᢱᆔຬળߢߪ⾗⹜ᢱߩ৻ጀߩలታࠍ࿑ࠆߚ߼‫ޔ‬㊁ᄖ⺞ᩏߥߤࠍⴕ߁ᣇ‫ߦޘ‬ᮡᧄ⾗ᢱណ㓸߳ߩදജࠍ߅
㗿޿ߒߡ߅ࠅ߹ߔ‫⻉ޔߚ߹ޕ‬੐ᖱ߆ࠄ▤ℂ࿎㔍ߣߥߞߚᮡᧄߩขࠅᛒ޿߿‫ઁޔ‬ᯏ㑐ᚲ⬿ߩ⾗ᢱߣߩ੤឵ߦߟ޿ߡ
ߩ⋧⺣߽ฃߌ߹ߔ‫ߦࠄࠇߎޕ‬㑐ߔࠆߏㅪ⛊ߪ‫⾗ޔ‬ᢱᆔຬળ߹ߢ߅㗿޿޿ߚߒ߹ߔ‫ޕ‬
㧔ᐔᚑ 24 ᐕᐲㅪ⛊వ㧦੹੗໪㓶 siryou [at] pri.kyoto-u.ac.jp㧕
㧔ᢥ⽿㧦ᐔፒ㍈⍫㧕
2011 ᐕᐲ㧔ᐔᚑ 23 ᐕᐲ㧕ᚲ⬿⾗⹜ᢱ
⴫ 1 㔤㐳㘃㛽ᩰ߅ࠃ߮ᶧᶐᮡᧄ
๺ฬ
ቇฬ
ࡎࡒࡁࠗ࠼
Hominoidea
Hylobates spp.
࠹࠽ࠟࠩ࡞ዻ
Pan troglodytes
࠴ࡦࡄࡦࠫ࡯ዻ
Gorilla gorilla
ࠧ࡝࡜ዻ
Pongo pygmaeus
ࠝ࡜ࡦ࠙࡯࠲ࡦዻ
ᣥ਎⇇ࠩ࡞
Cercopithecoidea
Macaca spp.
ࡑࠞࠢዻ
Presbytis spp.
ࠦࡁࡂࠩ࡞ዻ
Trachypithecus cristata
࡜ࡦࠣ࡯࡞ዻ
Colobus spp.
ࠦࡠࡉࠬዻ
ࡔࡦ࠲ࡢࠗࠪࠪࡃ࠽ࠩ࡞ዻ Simias concolor
Nasalis larvatus
࠹ࡦࠣࠩ࡞ዻ
Cercopithecus spp.
ࠣࠛࡁࡦዻ
Erythrocebus patas
ࡄ࠲ࠬዻ
Cercocebus spp.
ࡑࡦࠟࡌࠗዻ
Theropithecus gelada
ࠥ࡜࠳ࡅࡅዻ
Papio spp.
ࡅࡅዻ
Mandrillus spp.
ࡑࡦ࠼࡝࡞ዻ
Colobinae indet.
ࠦࡠࡉࠬ੝⑼ዻਇ᣿
6917 599
5191 452
177 25
8
366 9
132 1
573 66
18 7
16 4
3
4
415 27
18 3
1
ᣂ਎⇇ࠩ࡞
࡝ࠬࠩ࡞ዻ
࡛ࠩ࡞ዻ
࠹ࠖ࠹ࠖዻ
ࡎࠛࠩ࡞ዻ
ࠢࡕࠩ࡞ዻ
࠙࡯࡝࡯ࠢࡕࠩ࡞ዻ
࠙࡯࡝࡯ࡕࡦࠠ࡯ዻ
ࠝࡑࠠࠩ࡞ዻ
ࠨࠠዻ
1479 227
994 63
62 21
52 3
51 4
14 12
3
17 2
93 30
16 2
Ceboidea
Saimiri sciureus
Aotus trivirgatus
Callicebus spp.
Alouatta spp.
Ateles spp.
Brachyteles spp.
Lagothrix spp.
Cebus spp.
Pithecia spp.
㛽ᩰ ᶧᶐ
102 100
61 48
33 46
7
3
1
3
࠙ࠕࠞ࡝ዻ
ࠥ࡞࠺ࠖࡕࡦࠠ࡯ዻ
ࡇࠣࡒ࡯ࡑ࡯ࡕ࠮࠶࠻ዻ
ࡑ࡯ࡕ࠮࠶࠻ዻ
࠲ࡑ࡝ࡦዻ
࡜ࠗࠝࡦ࠲ࡑ࡝ࡦዻ
ࡑ࡯ࡕ࠮࠶࠻⑼ዻਇ᣿
Cacajao calvus
1
Callimico goeldi
1
Cebuella pygmaeus
7
Callithrix spp.
64
Saguinus spp.
98
Leontopithecus rosalia 6
Callitrichidae indet.
-
ේ₎㘃߅ࠃ߮࠷ࡄࠗ㘃 Prosimii + Scandentia
Tupaia spp.
࠷ࡄࠗዻ
Lyonogale spp.
ࠝࠝ࠷ࡄࠗዻ
Lemur spp.
ࠠ࠷ࡀࠩ࡞ዻ
ࠛ࡝ࡑࠠࠠ࠷ࡀࠩ࡞ዻ
8
19
62
1
105 101
47 12
1
16
Varecia spp.
11
ࠪࡈࠔࠞዻ
4
Microcebus spp.
Lepilemur mustelinus 1
Propithecus verreauxi 2
ࠬࡠ࡯ࡠ࡝ࠬዻ
Nycticebus spp.
13
21
ࡐ࠶࠻࡯ዻ
Perodicticus spp.
1
1
ࡠ࡝ࠬዻ
ࠟ࡜ࠧዻ
ࡔࠟࡀࠩ࡞ዻ
Loris spp.
Galago spp.
1
16
2
41
Tarsius spp.
ࠠ࠷ࡀࠩ࡞਄⑼ዻਇ᣿ Lemuroidea indet.
1
-
1
5
ᦛ㥦₎㘃ዻਇ᣿
Strepsirrhini indet.
-
3
⒳ਇ᣿
unidentifiable
-
9
✚⸘
Total
8603 1036
ࡀ࠭ࡒࠠ࠷ࡀࠩ࡞ዻ
ࠗ࠲࠴ࠠ࠷ࡀࠩ࡞ዻ
3
1
-
⴫ 2 㔤㐳㘃એᄖߩ㛽ᩰᮡᧄ
๺ฬ[⋡࡮⑼]
ᮡᧄᢙ
Taxa [order/genus]
㘩⡺⋡
Carnivora
1003
࡟࠶ࠨ࡯ࡄࡦ࠳⑼(1) Ailurus; ࠗ࠿⑼(547) Canis‫ޔ‬
Nyctereutes‫ޔ‬Vulpes; ࠗ࠲࠴⑼(229) Martes‫ޔ‬Meles‫ޔ‬Mustela‫ޔ‬
Pteronura; ࠕࠪࠞ⑼(16) Callorhinus‫ޔ‬Eumetopias‫ޔ‬
㧙㧙
Zalophus; ࠕ࡜ࠗࠣࡑ⑼(7) Procyon; ࠢࡑ⑼(137)
Helarctos‫ޔ‬Melursus‫ޔ‬Selenarctos‫ޔ‬Ursus; ࡀࠦ⑼(40) Felis‫ޔ‬
Neofelis‫ޔ‬Panthera; ࡑࡦࠣ࡯ࠬ⑼(4) Crossarchus‫ޔ‬Mungos‫ޔ‬
Suricata; ࠫࡖࠦ࠙ࡀࠦ⑼(23) Artictis‫ޔ‬Paguma‫ޔ‬Viverricula
ᄸか⋡
Perissodactyla
10
࠙ࡑ⑼(8) Equus; ࡃࠢ⑼(2) Tapirus
㟗஧か⋡
Cetartiodactyla
457
ࠗࡁࠪࠪ⑼(346) Sus; ࡍ࠶ࠞ࡝࡯⑼(3) Tayassus; ࠙ࠪ⑼
(53) Ammotragus‫ޔ‬Antilope‫ޔ‬Bos‫ޔ‬Buvalus‫ޔ‬Capra‫ޔ‬
Capricornis‫ޔ‬Cephalophus‫ޔ‬Ovis; ࠪࠞ⑼(43) Cervus‫ޔ‬
Hydropotes‫ޔ‬Muntiacus; ࡑࠗ࡞ࠞ⑼(9) Delphinidae indet.;
⑼ਇ᣿/Family indet. (3)
࠻ࠟ࡝ࡀ࠭ࡒ⋡
Soricomorpha
48
࠻ࠟ࡝ࡀ࠭ࡒ⑼(39) Crocidura‫ޔ‬Suncus; ࡕࠣ࡜⑼(9)
Euscaptor‫ޔ‬Mogera‫ޔ‬Urotrichus
⠢ᚻ⋡
Chiroptera
4
ࠝࠝࠦ࠙ࡕ࡝⑼(1) Pteropus; ࠠࠢࠟࠪ࡜ࠦ࠙ࡕ࡝⑼(1)
Rhinolophus; ⑼ਇ᣿/Family indet. (2)
⊹⠢⋡
Dermoptera
2
ࡅ࡛ࠤࠩ࡞⑼(2) Cynocephalus
㦅ᱤ⋡
Rodentia
130
ࡗࡑࡀ⑼(1) Glirulus; ࡝ࠬ⑼(45) Callosciurus‫ޔ‬Eutamias‫ޔ‬
Petaurista‫ޔ‬Sciurus‫ޔ‬Spermophilus; ࡀ࠭ࡒ⑼(65) Apodemus‫ޔ‬
Cricetomys‫ޔ‬Microtus‫ޔ‬Mus‫ޔ‬Rattus; ࡄࠞ⑼(2) Agouti; ࠹
ࡦࠫࠢࡀ࠭ࡒ⑼(3) Cavis‫ޔ‬Dolichotis; ࠝࡑࠠࡗࡑࠕ࡜ࠪ
⑼(1) Coendou; ࠞࡇࡃ࡜⑼(1) Hydrochoerus; ࡗࡑࠕ࡜ࠪ
⑼(2) Atherurus‫ޔ‬Hystrix; ࠿࡯࠻࡝ࠕ⑼(6) Myocastor; ⑼
ਇ᣿/Family indet. (4)
࠙ࠨࠡ⋡
Lagomorpha
15
࠙ࠨࠡ⑼(13) Lepus; ࠽ࠠ࠙ࠨࠡ⑼(2) Ochotona
⇣▵⋡
Xenarthra
2
ࡈ࠲࡙ࡆ࠽ࡑࠤࡕࡁ⑼(2) Choloepus
ࠕࡈ࡝ࠞ࠻ࠟ࡝ࡀ࠭ࡒ⋡Afrosoricida
1
࠹ࡦ࡟࠶ࠢ⑼(1) Echinops
ጤ⁠⋡
Hyracoidea
1
ࠗࡢ࠳࠿ࠠ⑼(1) Procavia
㐳㥦⋡
Proboscidea
2
࠱࠙⑼(2) Elephas‫ޔ‬Loxodonta
᦭ⴼ⋡
Marsupialia
10
ࠝࡐ࠶ࠨࡓ⑼(3) Didelphis; ࠞࡦࠟ࡞࡯⑼(2) Macropos;
ࠢࠬࠢࠬ⑼(4) Phalanger‫ޔ‬Trichosurus; ࠙ࠜࡦࡃ࠶࠻⑼(1)
Vombatus
ູ੃㘃࡮⸘
Mammalia total
1685
㠽㘃
Aves
15
Phoenicopteriformes [ ࡈ ࡜ ࡒ ࡦ ࠧ 㘃 ] (3) indet.;
Columbiformes [ࡂ࠻㘃] (2) Columba; Falconiformes [࠲ࠞ
㘃 ] (2) Butastur ‫ ޔ‬Milvus; Galliformes [ ࠠ ࠫ 㘃 ] (4)
Bumbusicola‫ޔ‬Gallus;
Passeniformes [ࠬ࠭ࡔ㘃] (4) Zoothera‫ޔ‬Passer‫ޔ‬Strunus
Ῥ⯻㘃
Reptilia
Cheloniidae [࠙ࡒࠟࡔ㘃] (3) Caretta‫ޔ‬Chelonia;
Alligatoridae [ࠕ࡝ࠥ࡯࠲࡯㘃] (1) indet.; Boidae [ࡏ
ࠕ㘃] (2) Eunectes
6
㝼㘃
Pisces
1
Perciformes [ࠬ࠭ࠠ㘃] (1) Lateolabrax
✚⸘
Total
1707
⴫ 3 㔤㐳㘃એᄖߩᶧᶐ⾗ᢱ
๺ฬ[⋡࡮⑼]
ᮡᧄᢙ
Taxa [order/genus]
㘩⡺⋡
Carnivora
68
࡟࠶ࠨ࡯ࡄࡦ࠳⑼(1) Ailurus; ࠗ࠿⑼(4) Canis‫ޔ‬
Urocyon; ࠗ࠲࠴⑼(33) Martes‫ޔ‬Mustela; ࠕࠪࠞ⑼(1)
Otariidae indet.; ࠢࡑ⑼(10) Selenarctos; ࡀࠦ⑼(3)
Felis; ࠫࡖࠦ࠙ࡀࠦ⑼(4) Artictis‫ޔ‬Paguma; ⑼ਇ᣿
/Family indet.(12)
஧か⋡
Artiodactyla
1
࠙ࠪ⑼(1) Capricorni
࠻ࠟ࡝ࡀ࠭ࡒ⋡
Soricomorpha
9
࠻ࠟ࡝ࡀ࠭ࡒ⑼(7) Sorex‫ޔ‬Suncus; ࡕࠣ࡜⑼(2) Mogera‫ޔ‬
Urotrichus
⠢ᚻ⋡
Chiroptera
4
ࠝࠝࠦ࠙ࡕ࡝⑼(4) Roussetus
㦅ᱤ⋡
Rodentia
108
࡝ࠬ⑼(1) Sciuridae indet.; ࡀ࠭ࡒ⑼(106) Clethrionomys‫ޔ‬
Rattus; ࠿࡯࠻࡝ࠕ⑼(1) Myocastor
࠙ࠨࠡ⋡
Lagomorpha
1
࠙ࠨࠡ⑼(1) Leporidae indet.
ࠕࡈ࡝ࠞ࠻ࠟ࡝ࡀ࠭ࡒ⋡ Afrosoricida
1
࠹ࡦ࡟࠶ࠢ⑼(1) Tenrecidae indet.
᦭ⴼ⋡
Marsupialia
4
ࠞࡦࠟ࡞࡯⑼(1) Macropodidae indet.; ࡈࠢࡠࡕࡕࡦࠟ⑼
(1) Petaurus; ࠢࠬࠢࠬ⑼(2) Trichosurus
ູ੃㘃࡮⸘
Mammalia total
196
㠽㘃 Aves (1) Galliformes - Gallus [࠾ࡢ࠻࡝]
Ῥ⯻㘃 Reptilia (1)
✚⸘
Squamata indet. [࠻ࠞࠥ]
Total
198
⴫ 4 㔤㐳㘃ಽሶ↢‛ቇ↪⹜ᢱ
๺ฬ
ࡎࡒࡁࠗ࠼
࠴ࡦࡄࡦࠫ࡯ዻ
ࠧ࡝࡜ዻ
ࠝ࡜ࡦ࠙࡯࠲ࡦዻ
࠹࠽ࠟࠩ࡞ዻ
ࡈࠢࡠ࠹࠽ࠟࠩ࡞ዻ
ᣥ਎⇇ࠩ࡞
ࡑࠞࠢዻ
ࠣࠛࡁࡦዻ
ࡅࡅዻ
ᣂ਎⇇ࠩ࡞
ࡑ࡯ࡕ࠮࠶࠻ዻ
࠲ࡑ࡝ࡦዻ
࡝ࠬࠩ࡞ዻ
㧙㧙
ቇฬ
Hominoidea
Pan troglodytes
Gorilla gorilla
Pongo pygmaeus
Hylobates lar
Symphalangus syndactylus
୘૕ᢙ
27
17
4
4
1
1
Cercopithecoidea
Macaca spp.
Cercopithecus spp.
Papio spp.
9
7
1
1
Ceboidea
Callithrix spp.
Saguinus spp.
Saimiri sciureus
12
10
1
1
6. ੱ੐⇣േ
ᚲዻಽ㊁╬
⡯ฬ
ㆮવሶᖱႎಽ㊁
࿖㓙౒หవ┵
⎇ⓥ࠮ࡦ࠲࡯
࿖㓙౒หవ┵
⎇ⓥ࠮ࡦ࠲࡯
࿖㓙౒หవ┵
⎇ⓥ࠮ࡦ࠲࡯
ੱ㘃ㅴൻࡕ࠺࡞
⎇ⓥ࠮ࡦ࠲࡯
̪ࡏࡁࡏ
⇣േ
ౝኈ
஻⠨
2011/4/1
૬છ
છᦼߪ 2012/3/31 ߹ߢ
ਤ⠀ ⟤Ꮤ
2011/4/1
ណ↪
․ቯഥᢎ㧔G30㧕
⿷┙ ᐞ⏴
2011/7/31
ㄉ⡯
ഥᢎ
⿷┙ ᐞ⏴
2011/8/1
ណ↪
૒⮮ ⧷ᰴ
2011/8/1
ណ↪
᳁ฬ
ᐕ᦬ᣣ
೽ᚲ㐳
ᐔ੗ ໪ਭ
․ቯ⡯ຬ
․ቯഥᢎ
㧔⎇ⓥ㐿⊒ᣉ⸳౒↪㧕
2011/8/31
ቴຬಎᢎ᝼
ᐔ↰ ⡡
․ቯഥᢎ
2011/8/31
ጊᧄ ⌀਽
̪ࡏࡁࡏ
㧔ነ㒝⎇ⓥㇱ㐷㧕
․ቯಎᢎ᝼
2011/9/1
̪ࡅ࠻⑼ 3 ⒳
ᐔ↰ ⡡
㧔․೎ᢎ⢒⎇ⓥ㧕
․ቯഥᢎ
2011/9/1
̪ࡅ࠻⑼ 3 ⒳
ጊᧄ ⌀਽
㧔․೎ᢎ⢒⎇ⓥ㧕
ੱ㘃ㅴൻࡕ࠺࡞⎇
2011/9/30
ᐔ੗ ໪ਭ
ㆮવሶᖱႎಽ㊁
ⓥ࠮ࡦ࠲࡯㐳
․ቯഥᢎ
2011/9/30
̪Ყセ⹺⍮
દ᧛ ⍮ሶ
㧔ነ㒝⎇ⓥㇱ㐷㧕
ੱ㘃ㅴൻࡕ࠺࡞
ੱ㘃ㅴൻࡕ࠺࡞⎇
2011/10/1
ጟᧄ ቬ⵨
⎇ⓥ࠮ࡦ࠲࡯
ⓥ࠮ࡦ࠲࡯㐳
․ቯഥᢎ
2011/10/1
ᕁ⠨⸒⺆ಽ㊁
દ᧛ ⍮ሶ
㧔․೎ផㅴ⎇ⓥ㧕
㊁ᄖ㐳ᦼ⎇ⓥ
․ቯഥᢎ
2011/11/1
᧻↰ ৻Ꮧ
ࡊࡠࠫࠚࠢ࠻
㧔․೎ᢎ⢒⎇ⓥ㧕
2012/3/31
↢ᘒ଻ోಽ㊁
ᢎ᝼
ᷰㆻ ㇌ᄦ
․ቯഥᢎ
2012/3/31
ᕁ⠨⸒⺆ಽ㊁
દ᧛ ⍮ሶ
㧔․೎ផㅴ⎇ⓥ㧕
․ቯഥᢎ
2012/3/31
ୖጟ ᐽᴦ
㜞ᰴ⣖ᯏ⢻ಽ㊁
㧔ᣂቇⴚ㗔ၞ⎇ⓥ㧕
̪Ყセ⹺⍮㧦Ყセ⹺⍮⊒㆐㧔ࡌࡀ࠶࠮ࠦ࡯ࡐ࡟࡯࡚ࠪࡦ㧕⎇ⓥㇱ㐷
̪ࡏࡁࡏ㧦ࡏࡁࡏ㧔ᨋේ㧕⎇ⓥㇱ㐷
̪ࡅ࠻⑼ 3 ⒳㧦ࡅ࠻⑼ 3 ⒳Ყセ⎇ⓥࡊࡠࠫࠚࠢ࠻
࿖㓙౒หవ┵⎇ⓥ࠮ࡦ࠲࡯࡮․ቯഥᢎ
(G30)ࠃࠅ
࠙ࠗ࡞ࠬ⎇ⓥᚲ࡮⎇ⓥຬ㧔↥ቭቇㅪ៤㧕
ࠃࠅ
ㄉ⡯
ㄉ⡯
ណ↪
ណ↪
ࡏࡁࡏ㧔ᨋේ㧕⎇ⓥㇱ㐷࡮ቴຬಎᢎ᝼
ࠃࠅ
ࡏࡁࡏ㧔ᨋේ㧕⎇ⓥㇱ㐷࡮․ቯഥᢎ㧔ነ
㒝⎇ⓥㇱ㐷㧕ࠃࠅ
૬છ⚳ੌ
ㄉ⡯
૬છ
ណ↪
ណ↪
છᦼߪ 2013/9/30 ߹ߢ
Ყセ⹺⍮⊒㆐㧔ࡌࡀ࠶࠮ࠦ࡯ࡐ࡟࡯࡚ࠪࡦ㧕
⎇ⓥㇱ㐷࡮․ቯഥᢎ㧔ነ㒝⎇ⓥㇱ㐷㧕ࠃࠅ
ᣣᧄቇⴚᝄ⥝ળ࡮․೎⎇ⓥຬ(PD)ࠃࠅ
ቯᐕㅌ⡯
ㄉ⡯
ㄉ⡯
ᣂẟ࿖㓙ᖱႎᄢቇ⻠Ꮷ߳
ㄭ⇰ᄢቇකቇㇱഥᢎ߳
7. ᶏᄖᷰ⥶
(1) ᢎ⡯ຬ
ᚲዻ
᳁ฬ
ᕁ⠨⸒⺆
ᨋ ⟤㉿
2011/4/5㨪4/11
ࡑ࡟࡯ࠪࠕ
⋡⊛࿖
ࠝ࡜ࡦ࠙࡯࠲ࡦߩⴕേ⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
ᕁ⠨⸒⺆
෹᳗㓷Ꮖ
2011/4/10㨪4/14
ਛ࿖
2011 ␠ળ⹺⍮␹⚻⑼ቇ࿖㓙ળ⼏ෳട㨯⊒⴫㨯⾗ᢱ෼㓸
♽⛔⊒↢
⷏᧛ ೰
2011/4/14㨪4/18
ࠕࡔ࡝ࠞ
╙ 80 ࿁ࠕࡔ࡝ࠞᒻ⾰ੱ㘃ቇળෳട࡮⊒⴫࡮⾗ᢱ෼㓸
ࠥࡁࡓᄙ᭽ᕈ
Ꮉᧄ ⧐
2011/4/23㨪5/8
ࡀࡄ࡯࡞㨯ࡉ࡯࠲ࡦ
ᕁ⠨⸒⺆
᧻ᴛື㇢
2011/4/25㨪5/4
ࠕࡔ࡝ࠞ
␠ળㅴൻ
MA Huffman
2011/4/25㨪5/4
ࠕࡔ࡝ࠞ࡮ࠞ࠽࠳
㊁↢ࠨ࡞ⷰኤ࡮⹜ᢱ෼㓸
⎇ⓥᛂߜวࠊߖ࡮࠾ࡘ࡯࡛࡯ࠢ㔤㐳㘃ࠦࡠࠠࠕࡓෳ
ട࡮⾗ᢱ෼㓸
౒ห⎇ⓥߦ㑐ߔࠆ⎇ⓥㅪ⛊࡮ᖱႎ෼㓸࡮࠮ࡒ࠽࡯ෳട
↢ᘒ଻ో
ᯅᧄජ⛗
2011/4/26㨪8/19
ㆮવሶᖱႎ
ᐔ੗໪ਭ
2011/4/27㨪4/30
ᕁ⠨⸒⺆
᧻ᴛື㇢
2011/5/15㨪5/20
ᦼ㑆
⋡ ⊛
࠙ࠟࡦ࠳࡮ࠦࡦࠧ ㊁↢࠴ࡦࡄࡦࠫ࡯ߩ↢ᘒቇ⊛⺞ᩏ࡮⎇ⓥળෳട࡮⎇ⓥ
ㅪ⛊
᳃ਥ౒๺࿖
บḧ
࿖㓙ࡢ࡯࡚ࠢࠪ࠶ࡊ಴Ꮸ࡮⊒⴫࡮⎇ⓥㅪ⛊
ੱ㑆ߩᧄᕈߩㅴൻ⊛⿠Ḯߦ㑐ߔࠆవ┵⎇ⓥߦଥࠆ
⧷࿖
⎇ⓥㅪ⛊
㧙㧙
࠮ࡦ࠲࡯
౗ሶ᣿ਭ
2011/5/16㨪5/22
࠲ࠗ
ࡢ࡯࡚ࠢࠪ࠶ࡊෳട࡮⎇ⓥᛂߜวࠊߖ࡮ᣉ⸳⷗ቇ
࠮ࡦ࠲࡯
೨↰ౖᒾ
2011/5/16㨪5/22
࠲ࠗ
ࡢ࡯࡚ࠢࠪ࠶ࡊෳട࡮⎇ⓥᛂߜวࠊߖ࡮ᣉ⸳⷗ቇ
੐ോቶ
਄၂ᵏᶈ
2011/5/16㨪5/22
࠲ࠗ
ࡢ࡯࡚ࠢࠪ࠶ࡊෳട࡮ㅪ៤ᯏ㑐ߣߩㅪ⛊⺞ᢛ࡮ᣉ⸳⷗ቇ
ㅴൻᒻᘒ
ự↰ ⓛ
2011/5/16㨪5/23
࠲ࠗ
ࡢ࡯࡚ࠢࠪ࠶ࡊෳട࡮⎇ⓥᛂߜวࠊߖ࡮ᣉ⸳⷗ቇ
࠮ࡦ࠲࡯
ጟᧄቬ⵨
2011/5/18㨪5/21
࠲ࠗ
ࡢ࡯࡚ࠢࠪ࠶ࡊෳട࡮⎇ⓥᛂߜวࠊߖ
⹺⍮ቇ⠌
ᱜ㜞ା↵
2011/5/28㨪6/3
ࠣ࡝࡯ࡦ࡜ࡦ࠼
࿖㓙࠮ࡦ࠲࡯
DA Hill
2011/6/5㨪6/10
ࠗࡦ࠼ࡀࠪࠕ
␠ળㅴൻ
ࡏࡁࡏ
㧔ᨋේ㧕
MA Huffman
2011/6/13㨪6/29
ጊᧄ⌀਽
2011/6/13㨪8/23
౒ห⎇ⓥᛂߜวࠊߖ࡮⾗ᢱ࡮ᖱႎ෼㓸
╙ 2 ࿁᧲ධࠕࠫࠕ࿖㓙ࠦ࠙ࡕ࡝ቇળ಴Ꮸ࡮⾗ᢱ෼
㓸࡮⎇ⓥᛂߜวࠊߖ࡮G30 ࿖㓙ࠦ࡯ࠬᐢႎᵴേ
࿖㓙ࠪࡦࡐࠫ࠙ࡓਥ௅࡮⊒⴫࡮㊁ᄖ↢ᘒⷰኤ࡮⎇ⓥㅪ⛊
㊁↢ࡏࡁࡏߩⴕേ⺞ᩏ࡮ࡏࡁࡏ㘺⢒ߦ㑐ߔࠆ⾗ᢱ෼
㓸
␠ળㅴൻ
ㄞ ᄢ๺
2011/6/15㨪8/20
ᕁ⠨⸒⺆
᧻ᴛື㇢
2011/6/16㨪7/3
␠ળㅴൻ
ฎᏒ೰ผ
2011/6/16㨪7/5
ࠬ࡝࡜ࡦࠞ
ࠦࡦࠧ᳃ਥ౒๺
࿖࡮ࡌ࡞ࠡ࡯
ࡌ࠻࠽ࡓ࡮ࠗࡦ࠼
⎇ⓥᛂߜวࠊߖ࡮ណ㘩↢ᘒ⺞ᩏ࡮ࠨࡦࡊ࡞ಽᨆ
ࡀࠪࠕ
㊁↢ࠝ࡜ࡦ࠙࡯࠲ࡦߩ↢ᘒⅣႺ⺞ᩏ࡮⎇ⓥㅪ⛊࡮⾗
ࡑ࡟࡯ࠪࠕ
ᢱ෼㓸
࠙ࠟࡦ࠳
࠴ࡦࡄࡦࠫ࡯ߩ↢ᘒ⺞ᩏ
♽⛔⊒↢
㜟੗ᱜᚑ
2011/6/18㨪6/29
ࡒࡖࡦࡑ࡯
㜞ᰴ⣖ᯏ⢻
ਛ᧛స᮸
2011/6/18㨪6/23
࠼ࠗ࠷
ࠥࡁࡓᄙ᭽ᕈ
Ꮉᧄ ⧐
2011/6/19㨪6/29
ࠬ࡝࡜ࡦࠞ
↢ᘒ଻ో
ඨ⼱๋㇢
2011/6/22㨪6/30
ࡑ࡟࡯ࠪࠕ
࿖㓙࠮ࡦ࠲࡯
FB Bercovitch
2011/7/2㨪7/17
࠽ࡒࡆࠕ
੐ോቶ
ዊ㊁৻ઍ
2011/7/5㨪7/13
ࡉ࡯࠲ࡦ
ㅴൻᒻᘒ
ự↰ ⓛ
2011/7/8㨪7/12
࠲ࠗ
ࡏࡁࡏ
㧔ᨋේ㧕
ᐔ↰ ⡡
2011/7/18㨪7/22
ࠕࡔ࡝ࠞ
⛔ว⣖ࠪࠬ࠹ࡓ
㜟↰᣽ᒾ
2011/7/13㨪7/22
ࠗ࠲࡝ࠕ࡮
ࡈ࡜ࡦࠬ
ᕁ⠨⸒⺆
᧻ᴛື㇢
2011/8/4㨪8/19
ᐔ↰ ⡡
2011/8/4㨪8/14
ᨋ ⟤㉿
2011/8/5㨪9/6
ࡏࡁࡏ
㧔ᨋේ㧕
ᕁ⠨⸒⺆
ࠤ࠾ࠕ࡮࡞ࡢࡦ
࠳࡮⧷࿖
ࠤ࠾ࠕ࡮
࡞ࡢࡦ࠳
ࠦࡦࠧ᳃ਥ౒๺࿖
ൻ⍹ᮡᧄⷰኤ
࿖㓙ࡑ࡯ࡕ࠮࠶࠻⎇ⓥ⃰ળ⼏ෳട࡮⊒⴫෸߮⣖ᯏ⢻
ࡕ࠺࡞േ‛⎇ⓥ࠮ࡦ࠲࡯⸳┙Ḱ஻ߩ⎇ⓥേะ⺞ᩏ
࿖㓙ࠪࡦࡐࠫ࠙ࡓ಴Ꮸ࡮⾗ᢱ෼㓸࡮⎇ⓥᛂߜวࠊߖ
㊁ᄖ⺞ᩏ࡮⾗ᢱ෼㓸࡮⎇ⓥᛂߜวࠊߖ
╙ 1 ࿁ Wild Giraffe Indaba ࠞࡦࡈࠔ࡟ࡦࠬෳട࡮࿖
㓙ࠦ࡯ࠬᐢႎᵴേ࡮⾗ᢱ෼㓸
੩ㇺᄢቇࡉ࡯࠲ࡦ෹ᅢࡊࡠࠣ࡜ࡓዷ㐿ߦଥࠆⷞ
ኤ࡮㑐ଥᯏ㑐ߣߩᛂߜวࠊߖ
Use of non-human primates for biomedical Research ෳ
ട࡮⻠Ṷ࡮⎇ⓥᛂߜวࠊߖ
ࠕࡔ࡝ࠞേ‛࿦᳓ᣖ㙚දળࡏࡁࡏ SSP ળ⼏಴Ꮸ࡮⾗
ᢱ෼㓸
8th International Brain Research Organization World
Congress of Neuroscience ෳട࡮⊒⴫࡮⾗ᢱ෼㓸࡮࠮
ࡒ࠽࡯ෳട
ࡢ࡯࡚ࠢࠪ࠶ࡊෳട࡮⾗ᢱ෼㓸෸߮㊁↢േ‛ߩⴕ
േ࡮↢ᘒⅣႺ⺞ᩏ
ࡢ࡯࡚ࠢࠪ࠶ࡊෳട࡮⾗ᢱ෼㓸෸߮㊁↢േ‛ߩⴕ
േ࡮↢ᘒⅣႺ⺞ᩏ
ࠪࡦࡐࠫ࠙ࡓෳട࡮⾗ᢱ෼㓸෸߮ࡏࡁࡏߩ↢ᘒ⺞ᩏ
␠ળㅴൻ
ฎᏒ೰ผ
2011/8/5㨪9/27
ࠦࡦࠧ᳃ⴐ౒๺࿖
ࠪࡦࡐࠫ࠙ࡓෳട࡮⾗ᢱ෼㓸෸߮㔤㐳㘃⃻࿾⺞ᩏ
ࠥࡁࡓᄙ᭽ᕈ
Ꮉᧄ ⧐
2011/8/6㨪8/21
ࡍ࡞࡯
࠮ࡦ࠲࡯
ᯅᧄ⋥ሶ
2011/8/15㨪8/22
ࠕࡔ࡝ࠞ
ㅴൻᒻᘒ
ự↰ ⓛ
2011/8/15㨪8/31
ࡌ࠻࠽ࡓ࡮࠲ࠗ
ࠕࡦ࠺ࠬߩ࡜ࠢ࠳⑼േ‛ߩ㓸࿅ㆮવቇ⺞ᩏ
࿖㓙ࠛࡦ࡝࠶࠴ࡔࡦ࠻ળ⼏಴Ꮸ࡮⊒⴫࡮⾗ᢱ෼㓸෸
߮േ‛㘺⢒ᣉ⸳⷗ቇ
⎇ⓥᛂߜวࠊߖ࡮㔤㐳㘃ߩᄙ᭽ᕈ⺞ᩏ
࿖㓙࠮ࡦ࠲࡯
DA Hill
2011/8/19㨪8/28
㖧࿖
࿖㓙ࠦ࡯ࠬᐢႎᵴേ࡮ቇ↢൐㓸ᵴേ
࿖㓙࠮ࡦ࠲࡯
ኋベࡑࡒ
2011/8/17㨪8/26
࠲ࡦࠩ࠾ࠕ
࠮ࡦ࠲࡯
㋈ᧁ᮸ℂ
2011/8/25㨪9/4
บḧ
Ყセ⹺⍮⊒㆐
દ᧛⍮ሶ
2011/8/27㨪9/3
ࡈ࡜ࡦࠬ
࿖㓙ࠦ࡯ࠬߩ๟⍮࡮ቇ↢⺃⥌ᵴേ
ࠟࡦࡑࡋ࡞ࡍࠬ࠙ࠗ࡞ࠬ㑐ਈ࡝ࡦࡄ⣲ߦ㑐ߔࠆ౒
ห⎇ⓥ࡮⎇ⓥᛂߜวࠊߖ
࡛࡯ࡠ࠶ࡄⷞⷡቇળෳട࡮⊒⴫࡮⾗ᢱ෼㓸
࿖㓙࠮ࡦ࠲࡯
FB Bercovitch
2011/8/29㨪9/30
ࠩࡦࡆࠕ
࿖㓙ࠦ࡯ࠬᐢႎᵴേ࡮ቇ↢൐㓸ᵴേ
ࠥࡁࡓᄙ᭽ᕈ
Ꮉᧄ ⧐
2011/8/27㨪9/7
ࡑ࠳ࠟࠬࠞ࡞
ࡢࠝࠠ࠷ࡀࠩ࡞ߩ↢ᘒ⺞ᩏ
࿖㓙࠮ࡦ࠲࡯
⿷┙ᐞ⏴
2011/8/30㨪8/31
㖧࿖
␠ળㅴൻ
MA Huffman
2011/9/1㨪9/22
ࠗ࠲࡝ࠕ࡮⧷࿖
⹺⍮ቇ⠌
ᱜ㜞ା↵
2011/9/2㨪9/8
࠽ࡒࡆࠕ
౒ห⎇ⓥᛂߜวࠊߖ
㔤㐳㘃ߩ⥄Ꮖᴦ≮⎇ⓥߦ㑐ߔࠆ౏㐿⻠Ṷ࡮⺰ᢥ૞
ᚑ࡮⎇ⓥᛂߜวࠊߖ
⎇ⓥᛂߜวࠊߖ࡮⾗ᢱ෼㓸
㧙㧙
♽⛔⊒↢
㜟੗ᱜᚑ
2011/9/4㨪9/24
⧷࿖࡮࠼ࠗ࠷
ൻ⍹ᮡᧄⷰኤ
ㅴൻᒻᘒ
ự↰ ⓛ
2011/9/5㨪9/13
࠲ࠗ
ᕁ⠨⸒⺆
᧻ᴛື㇢
2011/9/5㨪9/19
ࡑ࡟࡯ࠪࠕ
࿖㓙࠮ࡦ࠲࡯
DA Hill
2011/9/6㨪9/23
⧷࿖
ࠪࡦࡐࠫ࠙ࡓෳട࡮⊒⴫࡮⾗ᢱ෼㓸෸߮ࡈࠖ࡯࡞࠼⺞ᩏ
࿖㓙ࡢ࡯࡚ࠢࠪ࠶ࡊෳട࡮⾗ᢱ෼㓸෸߮േ‛⋧ߩ⺞
ᩏ࡮ࠝ࡜ࡦ࠙࡯࠲ࡦߩⴕേⷰኤ
ࡄ࡯࠻࠽࡯ᯏ㑐ߣߩㅪ៤ᛂߜวࠊߖ
ࠥࡁࡓᄙ᭽ᕈ
Ꮉᧄ ⧐
2011/9/8㨪9/29
ࡉ࡯࠲ࡦ₺࿖
㔤㐳㘃ߣኅ⇓ߩ㓸࿅ㆮવቇ⺞ᩏ
ࠥࡁࡓᄙ᭽ᕈ
↰ਛᵗਯ
2011/9/15㨪9/26
࠲ࠗ
࠲ࠗධㇱߩ㔤㐳㘃ߩಽᏓߣ↢ᕷ⁁ᴫ⺞ᩏ
ㅴൻᒻᘒ
ự↰ ⓛ
2011/9/15㨪9/26
࠲ࠗ
࠲ࠗධㇱߩ㔤㐳㘃ߩಽᏓߣ↢ᕷ⁁ᴫ⺞ᩏ
࠮ࡦ࠲࡯
ጟᧄቬ⵨
2011/9/25㨪10/2
ࠗࡦ࠼ࡀࠪࠕ
ᖱႎ੤឵࡮∉ቇ⺞ᩏ
ㅴൻᒻᘒ
ự↰ ⓛ
2011/10/4㨪10/8
࠲ࠗ
㔤㐳㘃⺞ᩏߩߚ߼ߩ੐೨⃻࿾⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
࿖㓙࠮ࡦ࠲࡯
DA Hill
2011/10/9㨪10/27
ࡑ࡟࡯ࠪࠕ
࿖㓙ࠦ࡯ࠬߩ๟⍮࡮ቇ↢⺃⥌ᵴേ෸߮⎇ⓥᛂߜวࠊߖ
࿖㓙࠮ࡦ࠲࡯
⿷┙ᐞ⏴
2011/10/12㨪10/16
ࠕࡔ࡝ࠞ
ࡢ࡯࡚ࠢࠪ࠶ࡊෳട࡮⾗ᢱ෼㓸
ㆮવሶᖱႎ
ㇹ ᐽᐢ
2011/10/12㨪10/16
ਛ࿖
ࠪࡦࡐࠫ࠙ࡓෳട࡮⊒⴫
↢ᘒ଻ో
ᷰㆻ㇌ᄦ
2011/10/18㨪10/30
ਛ࿖
੐ോቶ
౎ᧁቯⴕ
2011/10/20㨪10/22
㖧࿖
࿖㓙࠮ࡦ࠲࡯
ਤ⠀⟤Ꮤ
2011/10/20㨪10/22
㖧࿖
ㆮવሶᖱႎ
ᐔ੗໪ਭ
2011/10/20㨪10/22
㖧࿖
㔤㐳㘃㊁ᄖ⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
⧯ᚻ⎇ⓥ⠪ࠗࡦ࠲࡯࠽࡚ࠪ࠽࡞࡮࠻࡟࡯࠾ࡦࠣ࡮ࡊ
ࡠࠣ࡜ࡓߦ㑐ߔࠆㅪ៤ㅪ⛊⺞ᢛ
⧯ᚻ⎇ⓥ⠪ࠗࡦ࠲࡯࠽࡚ࠪ࠽࡞࡮࠻࡟࡯࠾ࡦࠣ࡮ࡊ
ࡠࠣ࡜ࡓߦ㑐ߔࠆㅪ៤ㅪ⛊⺞ᢛ
࿖㓙ࠪࡦࡐࠫ࠙ࡓෳട࡮⻠Ṷ࡮⎇ⓥᛂߜวࠊߖ
࠮ࡦ࠲࡯
ጟᧄቬ⵨
2011/10/23㨪11/1
ਛ࿖
࠮ࡦ࠲࡯
᣿㉿ብᢥ
2011/10/24㨪10/29
ࠕࡔ࡝ࠞ
♽⛔⊒↢
⷏᧛ ೰
2011/11/1㨪11/14
ࡒࡖࡦࡑ࡯
♽⛔⊒↢
㜟੗ᱜᚑ
2011/11/1㨪12/9
ࡒࡖࡦࡑ࡯
⹺⍮ቇ⠌
㚅↰໪⾆
2011/11/9㨪11/17
␠ળㅴൻ
ㄞ ᄢ๺
2011/11/10㨪
2012/1/9
㜞ᰴ⣖ᯏ⢻
ୖጟᐽᴦ
2011/11/11㨪11/18
ࡈ࡜ࡦࠬ
ࠗࡦ࠼ࡀࠪࠕ࡮
ࡌ࠻࠽ࡓ
ࠕࡔ࡝ࠞ
⛔ว⣖ࠪࠬ࠹ࡓ
᧻ᧄᱜᐘ
2011/11/11㨪11/18
ࠕࡔ࡝ࠞ
╙ 41 ࿁ർ☨␹⚻⑼ቇળᄢળෳട࡮⊒⴫࡮⾗ᢱ෼㓸
ᖱႎ੤឵࡮∉ቇ⺞ᩏ
╙ 29 ࿁㔤㐳㘃ࠛࠗ࠭ࡕ࠺࡞ࠪࡦࡐࠫ࠙ࡓ಴Ꮸ࡮⾗
ᢱ෼㓸࡮⎇ⓥᛂߜวࠊߖ
ൻ⍹⊒ជ⺞ᩏ
ൻ⍹⊒ជ⺞ᩏ෸߮࿖㓙ࠪࡦࡐࠫ࠙ࡓ಴Ꮸ࡮⊒⴫࡮⾗
ᢱ෼㓸
⎇ⓥ࡮ታ㛎ᛂߜวࠊߖ
⥄ὼ଻⼔඙ߢߩ↢ᘒ⺞ᩏ࡮⎇ⓥㅪ⛊
╙ 41 ࿁ർ☨␹⚻⑼ቇળᄢળෳട࡮⊒⴫࡮⾗ᢱ෼㓸
㜞ᰴ⣖ᯏ⢻
ች࿾㊀ᒄ
2011/11/11㨪11/18
ࠕࡔ࡝ࠞ
╙ 41 ࿁ർ☨␹⚻⑼ቇળᄢળෳട࡮⊒⴫࡮⾗ᢱ෼㓸
⛔ว⣖ࠪࠬ࠹ࡓ
㜟↰᣽ᒾ
2011/11/12㨪11/18
ࠕࡔ࡝ࠞ
╙ 41 ࿁ർ☨␹⚻⑼ቇળᄢળෳട࡮⊒⴫࡮⾗ᢱ෼㓸
␠ળㅴൻ
MA Huffman
2011/11/11㨪11/18
บḧ
♽⛔⊒↢
ᳯᧁ⋥ሶ
2011/11/13㨪11/21
࠼ࠗ࠷
࿖㓙࠮ࡦ࠲࡯
FB Bercovitch
2011/11/16㨪12/1
ࠕࡔ࡝ࠞ
⻠⟵࡮⎇ⓥᛂߜวࠊߖ
╙ 22 ࿁࠮ࡦࠤࡦࡌ࡞ࠣ࿖㓙ળ⼏಴Ꮸ࡮⊒⴫࡮⾗ᢱ
෼㓸෸߮ᮡᧄⷰኤ
࿖㓙ࠦ࡯ࠬᐢႎᵴേ࡮ቇ↢൐㓸ᵴേ
␠ળㅴൻ
MA Huffman
2011/11/20㨪12/6
ࠬ࡝࡜ࡦࠞ
㊁↢㔤㐳㘃࿾ၞ⟲ߦ㑐ߔࠆᦨᣂᖱႎ࡮⹜ᢱ෼㓸
ᕁ⠨⸒⺆
෹᳗㓷Ꮖ
2011/11/21㨪11/29
ࡑ࡟࡯ࠪࠕ
㊁↢ࠝ࡜ࡦ࠙࡯࠲ࡦߩⷰኤ
ࡅ࠻⑼ 3 ⒳
ጊᧄ⌀਽
2011/11/22㨪12/9
ࡉ࡯࠲ࡦ
㊁ᄖ⺞ᩏ࡮⾗ᢱ෼㓸
࿖㓙࠮ࡦ࠲࡯
DA Hill
2011/11/23㨪12/1
⧷࿖
࿖㓙ࠦ࡯ࠬᐢႎᵴേ࡮ቇ↢൐㓸ᵴേ෸߮ᣣᧄ⇐ቇࡈ
ࠚࠕෳട
࿖㓙࠮ࡦ࠲࡯
ኋベࡑࡒ
2011/11/23㨪11/27
⧷࿖
࠮ࡦ࠲࡯
ጟᧄቬ⵨
2011/12/1㨪12/3
࠲ࠗ
♽⛔⊒↢
ᳯᧁ⋥ሶ
2011/12/3㨪12/10
ࡒࡖࡦࡑ࡯
࿖㓙ࠦ࡯ࠬᐢႎᵴേ࡮ቇ↢൐㓸ᵴേ෸߮ᣣᧄ⇐ቇࡈ
ࠚࠕෳട
Joint International Tropical Medicine Meeting 2011 ಴
Ꮸ࡮⾗ᢱ෼㓸
ൻ⍹ᮡᧄ࠺࡯࠲෼㓸࡮ᮡᧄᢛℂ
࠮ࡦ࠲࡯
ችㇱ⾆ሶ
2011/12/7㨪2013/3/7
⧷࿖࡮ࠝ࡜ࡦ࠳
㤗㈮ߣᗧ⼂ߦ㑐ߔࠆ⎇ⓥ
㐳ᦼ㊁ᄖ
᧻↰৻Ꮧ
2011/12/10㨪12/16
ࠬࠗࠬ
⎇ⓥᛂߜวࠊߖ࡮⻠Ṷ
ᕁ⠨⸒⺆
᧻ᴛື㇢
2011/12/11㨪2012/1/6
ࠡ࠾ࠕ࡮ࡈ࡜ࡦࠬ ㊁↢࠴ࡦࡄࡦࠫ࡯ߩ↢ᘒ⺞ᩏ࡮⎇ⓥㅪ⛊
ࠥࡁࡓᄙ᭽ᕈ
Ꮉᧄ ⧐
2011/12/12㨪12/22
ࡉ࡯࠲ࡦ࡮ࡀࡄ࡯࡞
㔤㐳㘃ߣኅ⇓ߩ⹜ᢱಽᨆߦ㑐ߔࠆ⎇ⓥᛂߜวࠊߖ
↢ᘒ଻ో
ᷰㆻ㇌ᄦ
2011/12/14㨪12/21
ࠗࡦ࠼ࡀࠪࠕ
ࠪ࡞ࡃ࡯࡞࠻ࡦ࡮ࠞ࠾ࠢࠗࠩ࡞ߩㅊ〔⺞ᩏ
㧙㧙
␠ળㅴൻ
ฎᏒ೰ผ
2011/12/17㨪2012/1/7 ࠙ࠟࡦ࠳
࠴ࡦࡄࡦࠫ࡯ߩᗵᨴ∝╬ߦ㑐ߔࠆ⃻࿾⺞ᩏ
␠ળㅴൻ
MA Huffman
2011/12/14㨪12/21
࠴ࠚࠦ
౒ห⎇ⓥߦ㑐ߔࠆ⎇ⓥㅪ⛊࡮⻠Ṷ
ࠥࡁࡓᄙ᭽ᕈ
ฎ⾐┨ᒾ
2012/1/4㨪1/8
࠲ࠗ
࠹࠽ࠟࠩ࡞ߩ DNA ࠨࡦࡊ࡞ߩ⺞ᢛ෸߮៝౉
ㆮવሶᖱႎ
ᐔ੗໪ਭ
2012/1/4㨪1/8
࠲ࠗ
࿖㓙࠮ࡦ࠲࡯
⿷┙ᐞ⏴
2012/1/4㨪1/7
ࠗࡦ࠼
ㅴൻᒻᘒ
ự↰ ⓛ
2012/1/14㨪1/18
ࠗࡦ࠼ࡀࠪࠕ
࠹࠽ࠟࠩ࡞ߩ DNA ࠨࡦࡊ࡞ߩ⺞ᢛ෸߮៝౉
Looking Within:Interdisciplinary Approaches to
Consciousness ෳട࡮⊒⴫
⺰ᢥክᩏ࡮⺞ᩏ⎇ⓥᛂߜวࠊߖ
ࠥࡁࡓᄙ᭽ᕈ
↰ਛᵗਯ
2012/1/18㨪2/18
࠲ࠗ
㐳ᦼ㊁ᄖ
᧻↰৻Ꮧ
2012/1/19㨪3/1
ࡑ࡟࡯ࠪࠕ
ࡅ࠻⑼ 3 ⒳
ᐔ↰ ⡡
2012/1/24㨪1/30
ࠕࡔ࡝ࠞ
ࡉ࠲ࠝࠩ࡞⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
㔤㐳㘃ߩ❥ᱺⴕേߦ㑐ߔࠆࡈࠖ࡯࡞࠼࠺࡯࠲ߩ෼
㓸࡮⎇ⓥᛂߜวࠊߖ
ࡏࡁࡏ SSP/EEP ળ⼏಴Ꮸ࡮⾗ᢱ෼㓸
␠ળㅴൻ
ฎᏒ೰ผ
2012/1/27㨪2/6
࠙ࠟࡦ࠳
࠴ࡦࡄࡦࠫ࡯ߩⷰኤ࡮⎇ⓥᛂߜวࠊߖ
ࡅ࠻⑼ 3 ⒳
ጊᧄ⌀਽
2012/2/4㨪3/16
⹺⍮ቇ⠌
ᱜ㜞ା↵
2012/2/5㨪2/10
♽⛔⊒↢
ᳯᧁ⋥ሶ
2012/2/12㨪2/26
ࠡ࠾ࠕ࡮ࡈ࡜ࡦࠬ ㊁↢࠴ࡦࡄࡦࠫ࡯ߩⴕേ⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
㖸ᭉߣ⸒⺆ߩ⿠Ḯߦ㑐ߔࠆ౒ห⎇ⓥᚑᨐߩߣࠅ߹
ࠕࡔ࡝ࠞ
ߣ߼࡮⾗ᢱ෼㓸
ࡈ࡜ࡦࠬ
╙ਃ♿ູ੃㘃ൻ⍹࠺࡯࠲෼㓸
ㅴൻᒻᘒ
ự↰ ⓛ
2012/2/16㨪2/26
࠲ࠗ
ࠕ࠶ࠨࡓࡕࡦࠠ࡯ߩ㊁ᄖ⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
♽⛔⊒↢
㜟੗ᱜᚑ
2012/2/19㨪2/21
บḧ
⎇ⓥᛂߜวࠊߖ
␠ળㅴൻ
MA Huffman
2012/2/25㨪3/9
ࡌ࠻࠽ࡓ
㊁ᄖ⺞ᩏ࡮⎇ⓥㅪ⛊෸߮ቇⴚࠪࡦࡐࠫ࠙ࡓෳട࡮⊒⴫
ᕁ⠨⸒⺆
ᨋ ⟤㉿
2012/2/28㨪3/8
ࡑ࡟࡯ࠪࠕ
ࠝ࡜ࡦ࠙࡯࠲ࡦߩⴕേ⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
ࠥࡁࡓᄙ᭽ᕈ
ฎ⾐┨ᒾ
2012/3/7㨪3/12
ࡑ࡟࡯ࠪࠕ
࠹࠽ࠟࠩ࡞ߩ૕⦡ᄌ⇣ߦ㑐ߔࠆ⺞ᩏ
࿖㓙࠮ࡦ࠲࡯
FB Bercovitch
2012/3/8㨪3/24
ࠕࡔ࡝ࠞ
࿖㓙ࠦ࡯ࠬᐢႎᵴേ࡮ቇ↢൐㓸ᵴേ
♽⛔⊒↢
㜟੗ᱜᚑ
2012/3/12㨪3/15
ਛ࿖
⹺⍮ቇ⠌
㚅↰໪⾆
2012/3/12㨪
2013/3/22
⧷࿖
ㆮવሶᖱႎ
ᐔ੗໪ਭ
2012/3/15㨪3/22
⧷࿖
ᮡᧄ⸘᷹࡮⎇ⓥᛂߜวࠊߖ
㔤㐳㘃ߩ⇣⒳㑆ߢߩࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦ෸߮␠ળ
⹺⍮ߩㅴൻߦ㑐ߔࠆ࿖㓙౒ห⎇ⓥ
㗡⣖ᓴⅣࡊࡠࠣ࡜ࡓߩ౒ห⎇ⓥᛂߜวࠊߖ
ㆮવሶᖱႎ
੹੗໪㓶
2012/3/17㨪3/24
ਛ࿖
㊁↢ࠨ࡞ߩⷰኤ࡮⹜ᢱ෼㓸࡮⎇ⓥᛂߜวࠊߖ
ㅴൻᒻᘒ
ự↰ ⓛ
2012/3/19㨪3/30
ࡒࡖࡦࡑ࡯
㔤㐳㘃ಽᏓߣ↢ᕷ⁁ᴫ⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
㜞ᰴ⣖ᯏ⢻
ਛ᧛స᮸
2012/3/21㨪3/26
ࠕࡔ࡝ࠞ
ᣉ⸳⷗ቇ࡮⎇ⓥㅪ⛊࡮ᖱႎ෼㓸
⛔ว⣖ࠪࠬ࠹ࡓ
ᄢ⍹㜞↢
2012/3/21㨪3/27
ࠕࡔ࡝ࠞ
ᣉ⸳⷗ቇ࡮⎇ⓥㅪ⛊࡮ᖱႎ෼㓸
(2) ᄢቇ㒮↢
ᚲዻ
᳁ฬ
↢ᘒ଻ో
Ỉ↰᥏ሶ
2011/5/9㨪6/5
ࠬࠗࠬ
⎇ⓥᛂߜวࠊߖ࡮⾗ᢱ෼㓸࡮⺰ᢥ૞ᚑ
↢ᘒ଻ో
ᄢ⼱ᵗ੺
2011/5/25㨪6/7
ࡑ࡟࡯ࠪࠕ
ࡉ࠲ࠝࠩ࡞ߩㆆേߣⴕേߦ㑐ߔࠆ੍஻⺞ᩏ࡮⎇ⓥㅪ⛊
♽⛔⊒↢
⷏ጟ૓৻㇢
2011/6/18㨪6/29
ࡒࡖࡦࡑ࡯
ൻ⍹ᮡᧄⷰኤ
↢ᘒ଻ో
ỳ↰㘧㠽
2011/6/22㨪6/30
ࡑ࡟࡯ࠪࠕ
㊁ᄖ⺞ᩏ࡮⾗ᢱ෼㓸࡮⎇ⓥᛂߜวࠊߖ
↢ᘒ଻ో
ᳯፉ ବ
2011/7/1㨪7/21
࠙ࠟࡦ࠳
ᕁ⠨⸒⺆
C Martin
2011/7/5㨪8/17
ࠕࡔ࡝ࠞ
ᕁ⠨⸒⺆
ጊ᪸⵨⟤
2011/7/8㨪7/14
ࡑ࡟࡯ࠪࠕ
㔤㐳㘃ߩⷰኤ࡮♮෼㓸࡮⎇ⓥᛂߜวࠊߖ
࠴ࡦࡄࡦࠫ࡯ߩⴕേ߆ࠄ⷗ࠆࠥ࡯ࡓℂ⺰ߩ౒ห⎇
ⓥ࡮⎇ⓥᛂߜวࠊߖ
ࠝ࡜ࡦ࠙࡯࠲ࡦߩⴕേ⺞ᩏ࡮⾗ᢱ෼㓸
ᕁ⠨⸒⺆
⁚㊁ᢥᶈ
2011/7/25㨪8/5
ࠕࡔ࡝ࠞ
Behavior 2011 ෳട࡮⊒⴫࡮⾗ᢱ෼㓸࡮⎇ⓥᛂߜวࠊߖ
␠ળㅴൻ
ᓾጊᄹᏔሶ
2011/8/5㨪9/27
ࠦࡦࠧ᳃ਥ౒๺࿖
ࡏࡁࡏߩ↢ᘒߦ㑐ߔࠆࠪࡦࡐࠫ࠙ࡓෳട෸߮⃻࿾⺞ᩏ
␠ળㅴൻ
ᩉ ⥝㎽
2011/8/5㨪2012/1/30
ࠦࡦࠧ᳃ਥ౒๺࿖
ᕁ⠨⸒⺆
ጊ᪸⵨⟤
2011/8/13㨪8/26
ࠕࡔ࡝ࠞ
ᕁ⠨⸒⺆
ᬀ↰ ᗐ
2011/8/17㨪8/26
࠲ࡦࠩ࠾ࠕ
ࡏࡁࡏߩ↢ᘒߦ㑐ߔࠆࠪࡦࡐࠫ࠙ࡓෳട෸߮⃻࿾⺞ᩏ
࿖㓙ࠛࡦ࡝࠶࠴ࡔࡦ࠻ળ⼏ෳട࡮⊒⴫࡮⾗ᢱ෼㓸෸߮
േ‛㘺⢒ᣉ⸳⷗ቇ
㊁↢࠴ࡦࡄࡦࠫ࡯෸߮↢‛ߩ↢ᘒ⺞ᩏ
ᕁ⠨⸒⺆
ᰞᐸ㓁ሶ
2011/8/17㨪8/26
࠲ࡦࠩ࠾ࠕ
␠ળㅴൻ
㖧 ᤘ᥏
2011/9/19㨪
2012/3/16
࠼ࠗ࠷
♽⛔⊒↢
⷏ጟ૓৻㇢
2011/9/4㨪9/30
⧷࿖࡮࠼ࠗ࠷
ᦼ㑆
⋡⊛࿖
㧙㧙
⋡ ⊛
㊁↢࠴ࡦࡄࡦࠫ࡯෸߮↢‛ߩ↢ᘒ⺞ᩏ
㘺⢒ࡏࡁࡏߣ࠴ࡦࡄࡦࠫ࡯ߩᵴേߦ㑐ߔࠆ⺞ᩏ࡮࠺࡯
࠲෼㓸
ൻ⍹ᮡᧄⷰኤ
♽⛔⊒↢
⷏ጟ૓৻㇢
2011/11/1㨪12/11
ࡒࡖࡦࡑ࡯࡮࠲ࠗ ൻ⍹⊒ជ⺞ᩏ࡮࿖㓙ࠪࡦࡐࠫ࠙ࡓ಴Ꮸ࡮⊒⴫࡮⾗ᢱ෼㓸
ㆮવሶᖱႎ
I Jahan
ࡃࡦࠣ࡜࠺ࠪࡘ
࠹࠽ࠟࠩ࡞ߩㆮવ⹜ᢱ෼㓸
ᄢ⼱ᵗ੺
2011/11/2㨪
2012/2/8
2011/11/9㨪12/8
↢ᘒ଻ో
ࡑ࡟࡯ࠪࠕ
㊁ᄖ⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
㜞ᰴ⣖ᯏ⢻
⑧භ㓷ผ
2011/11/11㨪11/18
ࠕࡔ࡝ࠞ
㜞ᰴ⣖ᯏ⢻
㡨ᳰ⩿଻
2011/11/12㨪11/18
ࠕࡔ࡝ࠞ
╙ 41 ࿁ർ☨␹⚻⑼ቇળᄢળෳട࡮⊒⴫࡮⾗ᢱ෼㓸
╙ 41 ࿁ർ☨␹⚻⑼ቇળᄢળෳട࡮⊒⴫࡮⾗ᢱ෼㓸࡮
⎇ⓥᛂߜวࠊߖ
ᕁ⠨⸒⺆
ጊ᪸⵨⟤
ࡈ࡜ࡦࠬ࡮ࠡ࠾ࠕ ㊁↢࠴ࡦࡄࡦࠫ࡯ߩ⺞ᩏ࡮⾗ᢱ෼㓸
↢ᘒ଻ో
ᳯፉ ବ
2011/12/10㨪
2012/3/15
2012/1/4㨪2/29
࠙ࠟࡦ࠳
㔤㐳㘃⒳ߩⷰኤ࡮♮෼㓸࡮⎇ⓥㅪ⛊
↢ᘒ଻ో
ᄢ⼱ᵗ੺
2012/1/20㨪3/7
ࡑ࡟࡯ࠪࠕ
ᕁ⠨⸒⺆
Y Kim
2012/2/12㨪2/26
㖧࿖
ᕁ⠨⸒⺆
⁚㊁ᢥᶈ
2012/2/13㨪2/27
⧷࿖
ࡉ࠲ࠝࠩ࡞↢ᘒ⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
ᄢဳ㘃ੱ₎ߩᲧセ⹺⍮⑼ቇߦ㑐ߔࠆ࿖㓙౒ห⎇ⓥታ
ᣉߩᛂߜวࠊߖ
⎇ⓥᛂߜวࠊߖ࡮ᚑᨐႎ๔
♽⛔⊒↢
દ⮮ Პ
2012/3/12㨪3/15
ਛ࿖
ᮡᧄ⸘᷹࡮⎇ⓥᛂߜวࠊߖ
ㆮવሶᖱႎ
㋈ᧁධ⟤
2012/3/17㨪3/24
ਛ࿖
㊁↢ࠨ࡞ߩⷰኤ࡮⹜ᢱ෼㓸࡮⎇ⓥᛂߜวࠊߖ
(3) ᢎോ⵬૒ຬ࡮ᛛⴚ⵬૒ຬ࡮⎇ⓥᡰេផㅴຬ
ᚲዻ
᳁ฬ
ᦼ㑆
⋡⊛࿖
⋡ ⊛
ࠥࡁࡓᄙ᭽ᕈ
Ꮢ㊁ㅴ৻㇢
2011/3/1㨪8/10
࠼ࠗ࠷
ࡢࠝࠠ࠷ࡀࠩ࡞ߩ㐳ᦼ࠺࡯࠲ಽᨆ࡮⎇ⓥᛂߜวࠊߖ
↢ᘒ଻ో
દᏀᴦ⟤ᄹ
2011/4/26㨪10/28
࠙ࠟࡦ࠳
㊁↢࠴ࡦࡄࡦࠫ࡯ߩ↢ᘒቇ⊛⺞ᩏ
ㅴൻᒻᘒ
ᯅᧄ⵨ሶ
2011/6/20㨪6/24
㖧࿖
ᕁ⠨⸒⺆
ዊୖආବ
2011/8/12㨪8/27
ࠕࡔ࡝ࠞ
ࠥࡁࡓᄙ᭽ᕈ
Ꮢ㊁ㅴ৻㇢
2011/8/20㨪10/8
ࡑ࠳ࠟࠬࠞ࡞
ㅴൻᒻᘒ
ᯅᧄ⵨ሶ
2011/8/30㨪9/20
⧷࿖
ㅴൻᒻᘒ
ᯅᧄ⵨ሶ
2011/10/3㨪10/9
㖧࿖
ᕁ⠨⸒⺆
ዊୖආବ
ࠕࡔ࡝ࠞ
ࠥࡁࡓᄙ᭽ᕈ
Ꮢ㊁ㅴ৻㇢
2011/12/3㨪
2012/3/1
2011/12/5㨪12/12
࠼ࠗ࠷
ඳ‛㙚ᚲ⬿ߩ⾗ᢱ⺞ᩏ࡮෼㓸
╙ 10 ࿁࿖㓙ࠛࡦ࡝࠶࠴ࡔࡦ࠻ળ⼏ෳട࡮⊒⴫࡮⾗ᢱ
෼㓸෸߮ᣉ⸳⷗ቇ
ࠠ࠷ࡀࠩ࡞㘃ߩ㊁ᄖ⺞ᩏ࡮⎇ⓥㅪ⛊࡮⾗ᢱ෼㓸
ࠗࠡ࡝ࠬᒻ⾰ੱ㘃ቇળෳട࡮⊒⴫࡮⾗ᢱ෼㓸෸߮⾗ᢱ
ⷰኤ
ㆮ〔಴࿯ੱ㛽ߩⷰኤ
ࡑࠞࠢ㘃ߦኻߔࠆⅣႺࠛࡦ࡝࠶࠴ࡔࡦ࠻ߦ㑐ߔࠆ⾗
ᢱ෼㓸
࿖㓙ࠪࡦࡐࠫ࠙ࡓෳട࡮⊒⴫࡮⾗ᢱ෼㓸
↢ᘒ଻ో
દᏀᴦ⟤ᄹ
2012/1/4㨪3/28
࠙ࠟࡦ࠳
㊁↢࠴ࡦࡄࡦࠫ࡯ߩ↢ᘒቇ⊛
ㅴൻᒻᘒ
ᯅᧄ⵨ሶ
2012/2/24㨪3/1
㖧࿖
⎇ⓥᛂߜวࠊߖ࡮ᚲ⬿⾗ᢱߩⷰኤ࡮⾗ᢱ෼㓸
ㅴൻᒻᘒ
ട⾐⼱⟤ᐘ
2012/3/19㨪3/30
ࡒࡖࡦࡑ࡯
㔤㐳㘃ಽᏓߣ↢ᕷ⁁ᴫ⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
(4) 㕖Ᏹൕ⎇ⓥຬ
ᚲዻ
᳁ฬ
␠ળㅴൻ
ဈᏎື਽
ᦼ㑆
2011/4/30㨪8/20
⋡⊛࿖
ࠦࡦࠧ᳃ਥ౒๺࿖
⋡ ⊛
ࡏࡁࡏߩ㊁ᄖ⺞ᩏ࡮⎇ⓥㅪ⛊࡮⎇ⓥળ಴Ꮸ
࿖㓙࠮ࡦ࠲࡯
⷏Ỉ๺ሶ
2011/5/6㨪11/5
ࡉ࡯࠲ࡦ
ஜᐽ࡮ක≮࡮ᢥൻߦ㑐ߔࠆ⺞ᩏ⎇ⓥ
࿖㓙࠮ࡦ࠲࡯
AJJ MacIntosh
2011/5/7㨪5/29
ࡈ࡜ࡦࠬ
ࡍࡦࠡࡦߩⴕേߦ㑐ߔࠆ౒ห⎇ⓥ࡮⎇ⓥᛂߜวࠊߖ
␠ળㅴൻ
┻రඳᐘ
2011/7/2㨪10/3
ࠡ࠾ࠕ
ᕁ⠨⸒⺆
⪭ว⍮⟤
2011/8/13㨪8/25
ࠕࡔ࡝ࠞ
࿖㓙࠮ࡦ࠲࡯
M Seres
2011/8/17㨪8/26
࠲ࡦࠩ࠾ࠕ
㊁↢࠴ࡦࡄࡦࠫ࡯ߩ↢ᘒ⺞ᩏ
࿖㓙ࠛࡦ࡝࠶࠴ࡔࡦ࠻ળ⼏ෳട࡮⾗ᢱ෼㓸෸߮േ‛
࿦ᣉ⸳⷗ቇ
㊁↢࠴ࡦࡄࡦࠫ࡯ߩ⺞ᩏ࡮⾗ᢱ෼㓸
ࠬࠗࠬ㨯ࠝ࡯ࠬ࠻
࡝ࠕ㨯࠼ࠗ࠷㨯⧷࿖
3 ᰴరᢙℂ⸃ᨆࡊࡠࠣ࡜ࡓ㐿⊒ߩߚ߼ߩ౒ห⎇ⓥ
♽⛔⊒↢
⍫㊁ ⥶
2011/8/19㨪10/31
␠ળㅴൻ
ဈᏎື਽
ࠦࡦࠧ᳃ਥ౒๺࿖
㊁↢ࡏࡁࡏߩ⺞ᩏ࡮⎇ⓥᛂߜวࠊߖ
⛔ว⣖ࠪࠬ࠹ࡓ
ੑችᄥᐔ
2011/10/11㨪
2012/2/28
2011/11/12㨪11/18
ࠕࡔ࡝ࠞ
╙ 41 ࿁ർ☨␹⚻⑼ቇળᄢળෳട࡮⊒⴫࡮⾗ᢱ෼㓸
♽⛔⊒↢
⍫㊁ ⥶
2011/11/23㨪12/10
␠ળㅴൻ
┻రඳᐘ
ࡒࡖࡦࡑ࡯
࠙ࠟࡦ࠳࡮ࠦࡦ
ࠧ᳃ਥ౒๺࿖
ൻ⍹⊒ជ⺞ᩏ࡮ൻ⍹ᮡᧄ࠺࡯࠲෼㓸࡮ᮡᧄᢛℂ
࿖㓙࠮ࡦ࠲࡯
AJJ MacIntosh
2011/12/10㨪
2012/3/9
2012/1/28㨪2/9
ࡈ࡜ࡦࠬ㨯࠼ࠗ࠷
⎇ⓥᛂߜวࠊߖ࡮⾗ᢱ෼㓸
㧙㧙
㊁↢࠴ࡦࡄࡦࠫ࡯࡮ࡏࡁࡏߩ↢ᘒ⺞ᩏ࡮⎇ⓥㅪ⛊
࿖㓙࠮ࡦ࠲࡯
AJJ MacIntosh
2012/2/25㨪3/9
ࡌ࠻࠽ࡓ
㊁ᄖ⺞ᩏ࡮⎇ⓥㅪ⛊෸߮ቇⴚࠪࡦࡐࠫ࠙ࡓෳട࡮⊒⴫
(5) ቇᝄ․೎⎇ⓥຬ㧔PD㧕
ᚲዻ
᳁ฬ
ᦼ㑆
⋡⊛࿖
⋡ ⊛
↢ᘒ଻ో
᧻↰৻Ꮧ
2011/5/10㨪7/10
ࡑ࡟࡯ࠪࠕ࡮ࠪࡦ ࿖㓙࠻࡟࡯࠾ࡦࠣࡢ࡯࡚ࠢࠪ࠶ࡊෳട࡮࠹ࡦࠣࠩ࡞ㅊ
ࠟࡐ࡯࡞
〔⺞ᩏ࡮⎇ⓥㅪ⛊
ᕁ⠨⸒⺆
CD Dahl
2011/8/28㨪9/10
ࠬࠗࠬ࡮ࡈ࡜ࡦࠬ ⎇ⓥᛂߜวࠊߖ࡮⾗ᢱ෼㓸
↢ᘒ଻ో
᧻↰৻Ꮧ
2011/9/15㨪11/15
ࡑ࡟࡯ࠪࠕ
ᕁ⠨⸒⺆
CD Dahl
2012/1/3㨪1/8
ࠗࡦ࠼
⎇ⓥᛂߜวࠊߖ࡮࠹ࡦࠣࠩ࡞ߩㅊ〔⺞ᩏ
Looking Within:Interdisciplinary Approaches to
Consciousness ෳട࡮ᖱႎ෼㓸෸߮⎇ⓥᛂߜวࠊߖ
8. 㕖Ᏹൕ⻠Ꮷ
㧔ℂ㧕ᤊ⧷ม㧔⽷࿅ᴺੱ᧲੩ㇺකቇ⎇ⓥᚲ⹺⍮∝࡮㜞ᰴ⣖ᯏ⢻⎇ⓥಽ㊁೽ෳ੐⎇ⓥຬ㧕
‫ޟ‬㓐ᗧ⊛ⴕേ೙ᓮߩ␹⚻ၮ⋚‫ޠ‬2011.10.13㨪10.14
㧔㔤㧕ᄢ੗ᔀ㧔⁛┙ⴕ᡽ᴺੱ᫪ᨋ✚ว⎇ⓥᚲ㠽₞↢ᘒ⎇ⓥቶ㐳㧕
‫ޟ‬㊁↢േ‛ߩ㘩ᕈ⎇ⓥߣ଻ో߳ߩㆡ↪‫ޠ‬2011.11.29㨪11.30
㧔ℂ㧕㘖↰⪲ሶ㧔✚ว⎇ⓥᄢቇ㒮ᄢቇవዉ⑼ቇ⎇ⓥ⑼ᢎ᝼㧕
‫ޟ‬ㅴൻ㓸࿅ㆮવቇ‫ޠ‬2011.12.19㨪12.20
㧔ℂ㧕ᄢ⥪ᥓᔒ㧔ർᶏ㆏ᄢቇૐ᷷⑼ቇ⎇ⓥᚲഥᢎ㧕
‫ޟ‬ᣣᧄߩູ੃㘃ߦ߅ߌࠆ↢‛࿾ℂ‫ޔ‬ㅴൻ‫↢ޔ‬ᘒ㧦․ߦർߩⷞὐࠃࠅ‫ޠ‬2012.1.26㨪1.27
㧔ℂ㧕㧦ℂቇ⎇ⓥ⑼ᨒ 㧔㔤㧕㧦㔤㐳㘃⎇ⓥᚲᨒ
9. ࡝ࠨ࡯࠴࡮ࠕࠪࠬ࠲ࡦ࠻㧔R㨯A㧕
㧔᳁ฬ㧦ណ↪ᦼ㑆㧕
ᷡ㐳 ⼾ 㧦2011.4.1㨪2012.3.31
㋈ᧁ ධ⟤ 㧦2011.4.1㨪2012.3.31
⷏ጟ ૓৻㇢㧦2011.4.1㨪2011.9.30
ࠣࡠ࡯ࡃ࡞ COE
⷏ጟ ૓৻㇢㧦2011.10.1㨪2012.3.31
⑧භ 㓷ผ 㧦2011.4.1㨪2012.3.31
Ỉ↰ ⃍ሶ 㧦2011.4.1㨪2011.6.30
㘵↰ ᕺℂሶ㧦2011.4.1㨪2012.3.31
᧻Ꮉ ޽߅޿㧦2011.4.1㨪2012.3.31
቟੗ ᣧ♿ 㧦2011.4.1㨪2012.3.31
ศ↰ ᒎ↢ 㧦2011.4.1㨪2012.3.31
10. ࠹ࠖ࡯࠴ࡦࠣ࡮ࠕࠪࠬ࠲ࡦ࠻㧔T㨯A㧕
㧔᳁ฬ㧦ណ↪ᦼ㑆㧕
㔤㐳㘃ቇ࡮㊁↢േ‛♽⑼⋡ߦଥࠆ߽ߩ
ᣧᎹ ථᔒ 㧦2011.7.1㨪2011. 7.31
2011.10.1㨪2011.12.31
↰ਛ ⟤ㇺ 㧦2011.7.1㨪2011. 7.31
2011.10.1㨪2011.12.31
౗ሶ ፄ᣿ 㧦2011.10.1㨪2011.10.31
⏷᧛ ᦮ሶ 㧦2011.10.1㨪2011.12.31
ጊ↰ ᥓሶ 㧦2011.10.1㨪2011.12.31
૒⮮ ᧙ᄹ 㧦2011.10.1㨪2011.12.31
㋈ᧁ ධ⟤ 㧦2011.12.1㨪2011.12.31
ᓾጊ ᄹᏔሶ㧦2012.1.1㨪2012.1.31
㧙㧙
࿖㓙࠮ࡦ࠲࡯TA
᧛᧻ ᣿Ⓞ 㧦2011.12.1㨪2012.3.31
C Martin
㧦2011.12.1㨪2012.3.31
⏷᧛ ᦮ሶ 㧦2012.1.1㨪2012.3.31
ోቇ౒ㅢ⑼⋡ߦଥࠆ߽ߩ
ේ
ᥰ
㧦2011.5.1㨪2011.6.30
2011.9.1㨪2011.9.30
ᅏ↰ ᐽᐢ 㧦2011.6.1㨪2011.6.30
I Jahan
㧦2011.7.1㨪2011.7.31
ᓾጊ ᄹᏔሶ㧦2011.7.1㨪2011.7.31
ጊ↰ ᥓሶ 㧦2011.9.1㨪2011.9.30
ᬀ↰ ᗐ 㧦2011. 9.1㨪2011.9.30
్ ࡝࡜
㧦2011. 9.1㨪2011.9.30
ᳯፉ ବ
㧦2011. 9.1㨪2011.9.30
11. ᐕ㑆ࠬࠤࠫࡘ࡯࡞
2011 ᐕ
4᦬4ᣣ
4 ᦬ 22 ᣣ
5 ᦬ 7-8 ᣣ
6 ᦬ 11-12 ᣣ
6 ᦬ 18 ᣣ
7 ᦬ 30-31 ᣣ
8 ᦬ 2-3 ᣣ
9 ᦬ 25 ᣣ
9 ᦬ 26 ᣣ
9 ᦬ 28-30 ᣣ
10 ᦬ 12 ᣣ
10 ᦬ 30 ᣣ
ᣂ౉↢ࠝ࡝ࠛࡦ࠹࡯࡚ࠪࡦ
ᣂ౉ᚲຬ᱑ㄫળ
ᣣᧄ⿒ߜ߾ࠎቇળ ╙ 11 ࿁ቇⴚ㓸ળ㧔ਛㇱቇ㒮ᄢቇฦോේࠠࡖࡦࡄࠬ㧕
౒ห೑↪⎇ⓥળ‫ ╙ޟ‬12 ࿁࠾ࡎࡦࠩ࡞⎇ⓥ࠮ࡒ࠽࡯‫ޠ‬
ᧄቇഃ┙⸥ᔨᣣ
›ጊ౏㐿⻠ᐳ‫ޟ‬㔤㐳㘃ቇߩᗢߒߺ‫ޠ‬
ᄢቇ㒮ୃ჻⺖⒟౉ቇ⹜㛎
᧲੩౏㐿⻠ᐳ㧔ᣣᧄ⑼ቇᧂ᧪㙚㧕
ㆇ༡ᆔຬળ
ోቇ౒ㅢ⑼⋡‫ޟ‬㔤㐳㘃ቇߩ⃻࿷‫ޠ‬
ࠨ࡞ᘨ㔤⑂
Ꮢ᳃౏㐿
2012 ᐕ
1 ᦬ 10 ᣣ
1 ᦬ 23 ᣣ
2 ᦬ 2-3 ᣣ
2 ᦬ 21-22 ᣣ
2 ᦬ 23 ᣣ
3 ᦬ 2-3 ᣣ
3 ᦬ 10-11 ᣣ
3 ᦬ 11 ᣣ
3 ᦬ 12 ᣣ
ඳ჻⺰ᢥ⊒⴫ળ
ୃ჻⺰ᢥ⊒⴫ળ
౒ห೑↪⎇ⓥળ‫␠ޟ‬ળⴕേߣ⣖ 㧙ࠗࡔ࡯ࠫࡦࠣߣಽሶ㧙‫ޠ‬
ࠝ࡯ࡊࡦࠠࡖࡦࡄࠬ
ᄢቇ㒮ඳ჻⺖⒟✬౉⹜㛎
౒ห೑↪⎇ⓥળ‫ⴕޟ‬േ․ᕈࠍᡰ㈩ߔࠆࠥࡁࡓၮ⋚ߣ⣖ᯏ⢻ߩ⸃᣿‫ޠ‬
౒ห೑↪⎇ⓥળ‫ߩࠕࠫࠕޟ‬㔤㐳㘃ߩ଻ోߣ␠ળ↢ᘒ⎇ⓥߦ㑐ߔࠆㄭᐕߩᣂߚߥዷ㐿‫ޠ‬
ᷰㆻ㇌ᄦవ↢ᦨ⚳⻠⟵
ㆇ༡ᆔຬળ
3 ᦬ 15-16 ᣣ
3 ᦬ 17-18 ᣣ
3 ᦬ 24-25 ᣣ
ࡎࡒ࠾࠯࡯࡚ࠪࡦ⎇ⓥળ‫ߩࡦࡒࡄ࡯࠼ޟ‬ᓎഀ㧦ㆇേᯏ⢻߆ࠄ㜞ᰴᯏ⢻߳‫ޠ‬
౒ห೑↪⎇ⓥળ‫߇࡞ࠨޟ‬ੑ⿷ߢᱠߊߣ߈‫ޠ‬
౒ห೑↪⎇ⓥળ‫ ╙ޟ‬7 ࿁Ყセ␠ળ⹺⍮ࠪࡦࡐࠫ࠙ࡓ‫ޠ‬
㧙㧙
Υ. ⎇ⓥᵴേ
1. ⎇ⓥㇱ㐷෸߮㒝ዻᣉ⸳
ㅴൻ♽⛔⎇ⓥㇱ㐷
ㅴൻᒻᘒಽ㊁
㧨⎇ⓥ᭎ⷐ㧪
A) ࡑࠞࠢߩ♽⛔࿾ℂቇ⎇ⓥ
ự↰ⓛ㧘Ꮉᧄ⧐㧔ࠥࡁࡓᄙ᭽ᕈಽ㊁㧕㧘ᐔፒ㍈⍫㧘↰ਛᵗਯ㧔ࠥࡁࡓᄙ᭽ᕈಽ㊁㧕㧘ട⾐⼱⟤ᐘ
࡜ࠝࠬ࡮࠲ࠗ࡮ࡒࡖࡦࡑ࡯ߢฦ࿖⎇ⓥ⠪ߣߩ౒ห⎇ⓥ૕೙ߩ߽ߣߦ‫ޔ‬Ꮌ࿁⡞߈ขࠅ࡮ⷰኤ⺞ᩏߦࠃࠆࡑࠞࠢߩಽ
Ꮣ࡮ᒻᘒᄙ᭽ᕈ࡮↢ᕷታᘒ࠺࡯࠲ߣࠨࡦࡊ࡝ࡦࠣࠍⴕߞߚ‫⺞ߢ⋵ࠢ࠶ࠨ࡯ࡄࡦࡖ࠴ߣ⋵࡯ࡘࡇ࠲࠶ࠕߪߢࠬࠝ࡜ޕ‬
ᩏߒ‫ߩ࡯ࠠࡦࡕࡓࠨ࠶ࠕޔ‬ಽᏓධ㒢߅ࠃ߮ࠕࠞࠥࠩ࡞ߩࡔࠦࡦᴡ⷏ጯߦ߅ߌࠆಽᏓࠍ⷗޿ߛߒߚ‫ߪߢࠗ࠲ޕ‬ධㇱߢ
․ߦࠠ࠲࡮ࡒ࠽ࡒਔ⒳ࡉ࠲ࠝࠩ࡞ߩಽᏓߣᒻᘒቇ⊛⺞ᩏࠍⴕ޿‫ޔ‬ᓥ᧪޿ࠊࠇߡ޿ߚർ✲ 8-9 ᐲߩࠬ࡜࠲࠾㧙ࠢ࡜ࡆ
㑆㧔ࠢࡠ࡯ࡦࡑ࡯࡞ࠗᢿጀ㧕߇ਔ⒳ߩಽᏓႺ⇇ߢߪߥߊ‫ޔ‬Ⴚ⇇ၞߢ౉ࠅ⚵ߺ‫ޔ‬หᚲᕈߦಽᏓߒߡ޿ࠆ࿾ၞ߽⷗޿ߛ
ߐࠇߚ‫ޔߚ߹ޕ‬ർㇱߢࠕ࠶ࠨࡓࡕࡦࠠ࡯ߩ᝝₪⺞ᩏߦࠃࠅ‫ޔ‬ᒻᘒቇ⊛㨯ㆮવቇ⊛⾗ᢱࠍ෼㓸ߒߚ‫ߪߢ࡯ࡑࡦࡖࡒޕ‬
⷏ㇱ࡜ࠞࠗࡦጊ࿾࿾ၞ߅ࠃ߮ਛㇱߢಽᏓ⺞ᩏࠍⴕ޿‫ࡦࠗࠞ࡜ޔ‬ጊ࿾㜞✲ᐲ࿾ၞ㧔⷏ㇱ㧕ߢߪࠠ࠲ࡉ࠲ࠝࠩ࡞߿ࡌ࠾
ࠟࠝࠩ࡞ߪ⷗޿ߛߐࠇߕ‫✲ߣ࡯ࠠࡦࡕࡓࠨ࠶ࠕޔ‬ᐲ⊛ߦ᫮ߺಽߌࠍⴕߞߡ޿ࠆࠄߒ޿ߎߣࠍ⷗޿ߛߒߚ‫ޕ‬
B) ࠕࠞࠥࠩ࡞ߣ࠾ࡎࡦࠩ࡞ߩ੤㔀୘૕ߩᒻᘒቇ⊛ᬌ⸛
ự↰ⓛ㧘Ძ೑ବ㓶㧘Ꮉᧄ⧐㧔ࠥࡁࡓᄙ᭽ᕈಽ㊁㧕㧘ട⾐⼱⟤ᐘ
ජ⪲⋵ᚱ✚ඨፉ࿾ၞߦ⊒↢ߒߡ޿ࠆࠕࠞࠥࠩ࡞ߣ࠾ࡎࡦࠩ࡞ߩ੤㔀ߦ㑐ߒߡ‫ޔ‬ᒻᘒቇ⊛⺞ᩏ‫ߦ․ޔ‬የ㐳࡮૕Ძࡄ
࠲࡯ࡦ࡮૕⦡ࡄ࠲࡯ࡦߦ㑐ߔࠆᲧセಽᨆࠍⴕߞߚ‫ߺ⚵ߥ߹ߑ߹ߐߦ߁ࠂߓ߭ޕ‬วࠊߖߩ੤㔀߇⷗ࠄࠇ‫ࠅߥ߆ޔ‬㐳޿
㑆ߦࠊߚߞߡ੤㔀ߒߡ޿ࠆࠄߒ޿‫ޕ‬
C) ࡑࠞࠢߣ࠴ࡦࡄࡦࠫ࡯ߩᚑ㐳࡮ട㦂ᄌൻ⎇ⓥ
ự↰ⓛ㧘㋈ᧁ᮸ℂ㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕㧘ᐔፒ㍈⍫
ᕁᤐᦼ߆ࠄり૕ᚑᾫ߹ߢ‫߮ࠃ߅ޔ‬ട㦂ߦ઻߁り૕ᄌൻߦ㑐ߔࠆ⾗ᢱࠍ❑ᢿ⊛࡮ᮮᢿ⊛ߦ෼㓸ߒߚ‫ޕ‬⢵㐳ߪ 14 ᚽ
ߏࠈ߹ߢࠊߕ߆ߕߟߢߪ޽ࠆ߇Ⴧᄢߒ‫ࠄ߆ࠇߘޔ‬ᷫዋߒ‫ޔ‬20 ᚽએ㒠ᷫዋㅦᐲࠍჇߔ‫ߩઁޕ‬㐳ߐࠨࠗ࠭ߢߪ᦭ᗧߥ
ᷫዋߪ⷗ࠄࠇߥ޿‫ߪߢࠢࠞࡑޕ‬ᬁ㛽ߩ㛽┵≹ว߇ 15-20 ᚽߣ‫ߦ߁ࠂߓ߭ޔ‬ㆃ޿ߎߣ‫ߡߒߘޔ‬ᬁ૕㑆㑆㓒ߩ⍴❗߿ᄌ
ᒻᕈ㛽㑐▵∝ߩ⊒∝‫߮ࠃ߅ޔ‬㛽ኒᐲᷫዋߣᓸዋ㛽᛬ߦࠃߞߡ⣄ᩇߩ⍴❗߇߭߈߅ߎߐࠇߡ޿ࠆࠄߒ޿‫੹ޕ‬ᓟ‫ޔ‬⣄ᬁ
ߩ⹦⚦ߥᒻᘒቇ⊛ᐕ㦂ᄌൻߩ⎇ⓥ߇ᔅⷐߢ޽ࠆ‫ޕ‬
D) ⿷ౝ╭ߩ㈩⟎߆ࠄߺߚ⿷ߩᯏ⢻ゲߦ㑐ߔࠆ⸃೬ቇ⊛⎇ⓥ
ᐔፒ㍈⍫
㔤㐳㘃ߩ⿷ౝ╭ߩ⸃೬࡮ⷰኤࠍⴕ޿‫ޔ‬㛽㑆╭ߩ㈩⟎߆ࠄ⿷ߩᯏ⢻ゲߩ૏⟎ࠍផቯߔࠆ⹜ߺࠍ⛮⛯ਛߢ޽ࠆ‫ࠇߎޕ‬
߹ߢߦ‫ߪߡ޿߅ߦࡏࡁࡏޔ࡜࡝ࠧޔ‬⢛஥㛽㑆╭߇╙ 2 ⿾๟ࠅߦ㈩⟎ߐࠇߡ߅ࠅᯏ⢻ゲ߇ࡅ࠻ߣห᭽ߦ╙ 2 ⿾਄ߦ޽
ࠆߣផቯߐࠇࠆ৻ᣇߢ‫ࠢࠞࡑޔ‬㘃߿ࠢࡕࠩ࡞ߢߪᯏ⢻ゲ߇╙ 3 ⿾ߦሽ࿷ߔࠆߎߣ߇ࠊ߆ߞߡ޿ࠆ‫ޕ‬23 ᐕᐲߪ‫࠴ޔ‬
ࡦࡄࡦࠫ࡯ߦߟ޿ߡ⺞ᩏࠍⴕ޿‫ߪߢ࡯ࠫࡦࡄࡦ࠴ޔ‬⒳ౝᄌ⇣߇ᄢ߈ߊ‫ޔ‬2 㗡ߢߪࡅ࠻ߣห᭽ߦ‫ޔ‬⢛஥㛽㑆╭߇╙ 2
⿾๟ࠅߦ㈩⟎ߐࠇߡ޿ࠆ߇‫ ߩઁޔ‬2 㗡ߢߪࡑࠞࠢ㘃ߣห᭽ߦᯏ⢻ゲ߇╙ 3 ⿾ߦ޽ࠆߎߣ߇␜ໂߐࠇߚ‫੹ޕ‬ᓟ߽⛮⛯
ߒߡⷰኤ଀ࠍჇ߿ߔᔅⷐ߇޽ࠆ‫ޕ‬
E) ࠾ࡎࡦࠩ࡞ߩࡠࠦࡕ࡯࡚ࠪࡦߦ㑐ߔࠆታ㛎⊛⎇ⓥ
ᐔፒ㍈⍫㧘ự↰ⓛ㧘㋈ᧁ᮸ℂ㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕㧘ᣧᎹᷡᴦ㧔࿖㓙౒หవ┵⎇ⓥ࠮ࡦ࠲࡯㧕
㔤㐳㘃߇ੑ⿷߅ࠃ߮྾⿷ߢᱠߊ㓙ߦ‫ޔ‬り૕ฦㆇേಽ▵߇ߘࠇߙࠇߤߩࠃ߁ߦേ߈‫ޔ‬੕޿ߦද⺞ߔࠆߩ߆ࠍ᣿ࠄ߆
ߦߔࠆߎߣࠍ⋡⊛ߣߒ‫࡞ࠩࡦࡎ࠾ޔ‬ᱠⴕߩㆇേቇ⊛ಽᨆࠍ⛮⛯ਛߢ޽ࠆ‫ޕ‬23 ᐕᐲߦߪᣂߚߦ 2 ᱦߩ୘૕ࠍዉ౉ߒ
㚔ൻࠍⴕߞߚ‫ߪߕ߹ޕ‬ታ㛎⠪߅ࠃ߮ታ㛎ቶߦኻߔࠆⵍ㛎૕ߩ㚔ൻ߆ࠄᆎ߼‫ޔ‬ᐥ਄ߢߩੑ⿷߅ࠃ߮྾⿷ᱠⴕࡕ࠺࡞‫ޔ‬
࠻࡟࠶࠼ࡒ࡞਄ߢߩੑ⿷ᱠⴕࡕ࠺࡞ࠍ⏕┙ߒ‫ޔ‬ᰴߦ྾⿷ᱠⴕਛߩ྾⢇ߩേ߈ߦߟ޿ߡߩ੍஻⊛ߥ࠺࡯࠲ࠍ෼㓸ߒߚ‫ޕ‬
F) ᧲ධࠕࠫࠕߩࡑࠞࠢߩࡠࠦࡕ࡯࡚ࠪࡦߦ㑐ߔࠆㆇേቇ⊛⎇ⓥ
ᐔፒ㍈⍫㧘ự↰ⓛ
ߎࠇ߹ߢਥߣߒߡታ㛎ቶౝߢⴕࠊࠇߡ߈ߚ㔤㐳㘃ࡠࠦࡕ࡯࡚ࠪࡦߩㆇേቇ⊛ಽᨆࠍࡈࠖ࡯࡞࠼ߦ᜛ᒛߔࠆ⹜ߺ
ࠍ⛮⛯ਛߢ޽ࠆ‫ޕ‬23 ᐕᐲߪ‫ޔ‬22 ᐕᐲߦ෼㓸ߒߚ᧲ධࠕࠫࠕߩඨ㊁ᕈࡑࠞࠢ㧔ࠕ࠶ࠨࡓࡕࡦࠠ࡯‫ޔ࡞ࠩࠗࠢ࠾ࠞޔ‬
߅ࠃ߮ࡌ࠾ࠟࠝࠩ࡞㧕ߩࡠࠦࡕ࡯࡚ࠪࡦߩᤋ௝ࠍಽᨆߒߚ‫⚿ߩߘޕ‬ᨐ‫ߦ࡞ࠩࡦࡎ࠾ߪ࡯ࠠࡦࡕࡓࠨ࠶ࠕޔ‬Ყߴ‫ޔ‬ᱠ
ⴕᤨߦ⢋㑐▵ࠍᄢ߈ߊ protraction ߒ‫ޔ‬⢆㑐▵ࠍᄢ߈ߊિዷߐߖࠆߎߣ‫ߩ⿷ޔ‬ㆇ߮㗅ߦᄌ⇣߇ᄢ߈޿ߎߣ‫ޔ‬᝿ⴕᕈ࡮
ぞⴕᕈߩᱠⴕࠍࠃߊⴕ߁ߎߣߥߤ߇್᣿ߒߚ‫ޔߪߢ࡞ࠩࠝࠟ࠾ࡌߚ߹ޕ‬⢋↲㛽ߩᱠⴕ߳ߩነਈ߇ઁߩ 2 ⒳ࠃࠅᄢ߈
޿ߎߣ߇್᣿ߒߚ‫߇࡞ࠩࠝࠟ࠾ࡌޕ‬࿾਄ᱠⴕߦࠃࠅㆡᔕߒߡ޿ࠆߎߣࠍ␜ໂߔࠆ⚿ᨐߢ޽ࠆ‫ޕ‬
㧙㧙
G) 㔤㐳㘃ߩ㗡⬄ቇ
Ძ೑ବ㓶
߭߈⛯߈‫ޔ‬㔤㐳㘃ߩ㗡⬄ࠍ⸘᷹‫ޔ‬㕖⸘᷹ߩਔ㕙߆ࠄ⎇ⓥߒߡ޿ࠆ‫ޕ‬
H) 㔤㐳㘃ߩ⡻㛽ߩ⡻ᬁ㑐▵㕙ࠪࠚࠗࡊߣḧᦛߩಽᨆ
ട⾐⼱⟤ᐘ
㘃ੱ₎߿ࡅ࠻ߩᚳᐔ⢷ㇳߩ⿠Ḯ߿ㆡᔕ⊛ᗧ⟵ࠍ᣿ࠄ߆ߦߔࠆߎߣࠍ⋡⊛ߦ‫ޔ‬⢷ㇳߩᒻᘒಽᨆࠍⴕߞߡ޿ࠆ‫ᧄޕ‬ᐕ
ᐲߪ‫ޔ‬11 ⒳ߩ㔤㐳㘃ߩ⡻㛽ߩ㗡㗖ㇱߩ㐳᏷␜ᢙ߿⡻㛽㗖ㇱߩⷺᐲ‫ޔ‬ḧᦛ␜ᢙ‫ޔ‬⣺⡻╭☻㕙ㇱߩ⋧ኻ㐳ߥߤߩಽᨆࠍ
㐿ᆎߒߚ‫╙ߦ․ޕ‬ੑ⡻㛽ߪ‫ߩߘޔ‬ḧᦛ␜ᢙ߇ᚳᐔ⢷ㇳߩᜰᮡߦ߽೑↪ߐࠇࠆ߇‫࠹߿࡯ࠠࡦࡕ࡯࡝࡯࠙ޔ࡞ࠩࡕࠢޔ‬
ࡦࠣࠩ࡞ߣ޿ߞߚᄢဳߩ᮸਄ᕈࠨ࡞ߩ␜ᢙ୯ߪᄢဳ㘃ੱ₎ߩ߽ߩߣ㊀ⶄߒߡ޿ߚ‫ޕ‬
I) ࡑࠞࠢዻߩ೨⢇Ꮺ㛽ᩰ࡮਄⣨㛽㗡ᒻᘒߩ⒳㑆Ꮕߣ૏⟎⊛ⴕേ
ട⾐⼱⟤ᐘ㧘ự↰ⓛ
ࡑࠞࠢዻߪᄙ᭽ߥ⒳ࠍ฽ߺ‫ޔ‬⒳ߦࠃߞߡ೑↪ߔࠆᡰᜬၮ૕߿૏⟎⊛ⴕേߩ௑ะ߇⇣ߥࠆ‫ࠢࠞࡑޕ‬ዻ10⒳ߩ⢋↲
㛽‫ޔ‬㎮㛽‫ޔ‬਄⣨㛽ㄭ૏ㇱߩ⸘᷹୯ࠍಽᨆߒ‫ޔ‬⒳ߦࠃࠆ․ᓽ߇⷗޿ߛߖࠆ߆ߤ߁߆‫ޔ‬ᬌ⸛ߒߚ‫⚿ޕ‬ᨐ‫ޔ‬⒳㑆ᄌ⇣߇⒳
ౝᄌ⇣ߣᲧߴߡ㗼⪺ߢߥߊ‫ࠢࠞࡑޔ‬ዻߩ૕⢇㛽ᒻᘒߪၮᧄ⊛ߦߪ੕޿ߦࠃߊૃߡ޿ࠆߎߣ߇␜ߐࠇߚ‫ࡌޔߒߛߚޕ‬
࠾ࠟࠝࠩ࡞ߩ਄⣨㛽㗡ߪ⋧ኻ⊛ߦ⁜޿ߎߣ߇‫ޔ‬਄⣨㛽㗡ߩ⍫⁁ᓘ‫▵⚿ޔ‬ㇱߩ᏷‫ޔ‬⢋↲㛽㑐▵┃ߩ᏷ߣߩᲧセ߆ࠄ␜
ໂߐࠇߚ‫ߩߎޕ‬⒳߇߶ߣࠎߤ᮸਄ࠍ⒖േߦ↪޿ߥ޿ߎߣߦኻᔕߢ߈ࠆน⢻ᕈ߇޽ࠆ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
2)
3)
4)
5)
6)
ႎ๔
1)
2)
Fujita H, Hashimoto H, Shoda S, Suzuki T (2011) Dental Caries Prevalence as a Product of Agriculture and Subsistence
Pattern at the Yean-ri Site, South Korea. Caries Research 45:524-531.
Hamada Y, Sawada J, Cho F, Won MH, Hyun BHHyun (2011) Tubular anomalous bones found in both thighs of a
long-tailed macaque (Macaca fascicularis). Primates 53:25-30.
Kurita H, Suzumura T, Kanchi F, Hamada Y (2011) A photogrammetric method to evaluate nutritional status without
capture in habituated free-ranging Japanese macaques (Macaca fuscata): a pilot study. Primates 53:7-11.
Sakai T, Mikami A, Tomonaga M, Matsui M, Suzuki J, Hamada Y, Tanaka M, Miyabe-Nishiwaki T, Makishima H,
Nakatsukasa M, Matsuzawa T (2011) Differential Prefrontal White Matter Development in Chimpanzees and Humans.
Current Biology 21:1397-1402.
ጊፒஜ㧘ᯅᧄ⵨ሶ㧘⨃ේା↢ (2011) ੩ㇺᄢቇᄢቇ㒮ℂቇ⎇ⓥ⑼⥄ὼੱ㘃ቇ⎇ⓥቶᚲ⬿േ‛ᮡᧄ㧙ߣߊߦേ
‛ㆮሽ૕ߣേ‛ൻ⍹ߦߟ޿ߡ㧙. േ‛⠨ฎቇ 28:95-112.
Ogihara N, Makishima H, Hirasaki E, Nakatsukasa M (2012) Inefficient use of inverted pendulum mechanism during
quadrupedal walking in the Japanese macaque. Primates 53:41-48.
ự↰ⓛ (2011) ᧲ධࠕࠫࠕᄢ㒽ㇱߦ߅ߌࠆࡑࠞࠢߣߘߩㅴൻࡄ࠲࡯ࡦ. ᧲ධࠕࠫࠕᾲᏪᨋߩູ੃㘃 (1) 㔤㐳
㘃. ᶏᄖߩ᫪ᨋߣᨋᬺ No.81 (June, 2011) :57-62.
ᯅᧄ⵨ሶ (2012) ฬ⿧ಾㅢ಴࿯ੱ㛽. ผ〔ฬ⿧ಾㅢ ᢛ஻੐ᬺߦ઻߁⊒ជ⺞ᩏႎ๔ᦠ. ㅟሶᏒᢎ⢒ᆔຬળ㧦
42-47.
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
2)
3)
4)
5)
Hamada Y, Kurita H, Goto S, Morimitsu Y, Malaivijitnond S, Pathonton S, Pathontone B, Kingsada P, Vongsombath C,
Samouth F, Prazaysombath B (2011) Distribution and present status of long-tailed macaques (Macaca fascicularis) in
Laos and their ecological relationship with rhesus macaques (Macaca mulatta). (Monkeys on the Edge) (ed. Gumert MD,
Fuentes A, Jones-Engel L) p.72-98 Cambridge University Press.
Malaivijitnond S, Vazquez Y, Hamada Y (2011) Human impact on long-tailed macaques in Thailand. (Monkeys on the
Edge) (ed. Gumert MD, Fuentes A, Jones-Engel L) p.118-158 Cambridge University Press.
Ogihara N, Hirasaki E, Nakatsukasa M (2011) Experimental and computational studies of bipedal locomotion in the
bipedally-trained Japanese macaque. (Primate Locomotion: Linking in situ and ex situ Research) (ed. D'Aout K,
Vereecke EE) p.47-59 New York, Springer.
San AM, Hamada Y (2011) Distribution and current Status of Long-tailed macaques (Macaca fascicularis aurea) in
Myanmar. (Monkeys on the Edge) (ed. Gumert MD, Fuentes A, Jones-Engel L) p.45-71 Cambridge Univ. Press.
ᐔፒ㍈⍫ (2011) ࠨ࡞ߩᱠⴕ߆ࠄࡅ࠻ߩ⋥┙ੑ⿷ᱠⴕߩ⿠Ḯߣㅴൻࠍតࠆ㧚
‫ޟ‬ᣂ࡮㔤㐳㘃ቇߩߔߔ߼‫( ޠ‬੩ㇺᄢ
ቇ㔤㐳㘃⎇ⓥᚲ✬) p.19-35 ਣༀ㧚
ߘߩઁߩၫ╩
1)
2)
3)
ᐔፒ㍈⍫ (2011) ࠨ࡞ߩᱠ߈ᣇ. ੩ㇺᄢቇࠣࡠ࡯ࡃ࡞ COE‫ߩ‛↢ޟ‬ᄙ᭽ᕈߣㅴൻ⎇ⓥߩߚ߼ߩ᜚ὐᒻᚑ‫↢ޡ✬ޠ‬
߈‛ߚߜߩߟߠࠇ❱ࠅ ╙ 5 Ꮞ‫ ޢ‬p.121-127㧚
ᐔፒ㍈⍫ (2011) ੑ⿷ࠨ࡞ߩ㛽ᩰ㧘ᱠ㧘⣖. ᣣᧄੱ㘃ቇળㅴൻੱ㘃ಽ⑼ળ࠾ࡘ࡯ࠬ࡟࠲࡯. 25: p.4-5㧚
ᐔፒ㍈⍫㧘ᾢୖඳ㓶 (2011) ⿷ᐩㇱߩ╭㈩⟎ߣ⿷ᐩ࿶߆ࠄߺߚ㔤㐳㘃ߩ⿷ߩᯏ⢻ゲ. Anthropological Science
㧙㧙
(Japanese Series) 119: p.25-26㧚
ቇળ╬⊒⴫
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2)
3)
4)
5)
6)
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10)
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12)
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14)
15)
16)
17)
18)
19)
20)
21)
22)
23)
24)
Hamada Y (2011) Distribution and morphological characteristics of macaques in Thailand. International Symposium
“Biodiversity and Ecology of Wildlife in Thailand” (2011/09/06, Bangkok)㧚
Hamada Y (2011) Growth and Aging in Primates: Comparison of Life-History. First Thai National Symposium on
Animal Care and Use for Scientific Purposes & Laboratory Animal Trade Exhibition 2011 (2011/07/11-13, Bangkok).
Hamada Y (2011) Guideline for the care and use of laboratory primates. Workshop “Healthy Management for
Non-human Primates” (2011/05/18-20, Bangkok)㧚
Hashimoto H (2011) Mating Systems of the Prehistoric Jomon People from the Mainland Japan indicated by Dental
Traits. 13th Annual Conference of the Association for Biological Anthropology and Osteoarchaeology (2011/09/02-04,
Edinburgh, U.K).
Hirasaki E, Malaivijitnond S, Tojima S, Hamada Y (2011) A preliminary study on locomotor kinematics of the semi-wild
Assamese macaques (Macaca assamensis) in northern Thailand. The 81st annual meeting of the American Association
of Physical Anthropologists (2011/04, Oregon, USA).
Malaivijitnond S, Hamada Y, Arsaithamkul V (2011) How to catch free-ranging macaque monkeys? Workshop “Healthy
Management for Non-human Primates” (2011/05/18-20, Bangkok)㧚
Malaivjitnond S, Hamada Y (2011) What are nonhuman Primates? Workshop “Healthy Management for Non-human
Primates” (2011/05/18-20, Bangkok)㧚
⮮↰ዏ㧘ᯅᧄ⵨ሶ㧘Ꮉਭ଻ༀᥓ㧘ᄢ㊁ᙗ੖㧘ᐣ↰ᘕ⍫㧘㋈ᧁ㓉㓶 (2011) 㖧ඨፉ಴࿯ੱ㛽߆ࠄㄘ⠹ߩવ᠞࡮ฃ
ኈࠍ⠨ኤߔࠆ. ╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢળ (2011/11/04-06, ㇊ⷓᏒ)㧚
⮮↰ዏ㧘ᯅᧄ⵨ሶ㧘ᐣ↰ᘕ⍫㧘㋈ᧁ㓉㓶 (2011) ㄘ⠹ߪ㦏ᱤ₸ࠍჇടߐߖߚ߆㧫㧙㖧࿖൅ፉੱ㛽ߣ⑥቟㉿ੱ㛽
ߩૐ޿㦏ᱤ₸߆ࠄ↢ᬺࠍ⠨ኤߔࠆ㧙. ᣣᧄ⠨ฎቇදળ╙ 77 ࿁✚ળ (2011/05/28-29, ᧲੩ㇺ)㧚
ᮭ↰⛗㉿㧘 ጊ৻㆏㧘ự↰ⓛ (2011) ࠻ࡦࠟੱߩᚑ㐳ࡄ࠲࡯ࡦߣ⢈ḩߩᐕ㦂ᄌൻ. ╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢળ
(2011/11/04-06, ㇊ⷓᏒ)㧚
ự↰ⓛ㧘Pomchote P㧘↰ਛᵗਯ㧘Arsaithamakul V㧘Suryobroto B㧘Malaivijitnond S (2011) ࠠ࠲ࡉ࠲ࠝࠩ࡞㧔Macaca
leonina㧕ߣࡒ࠽ࡒࡉ࠲ࠝࠩ࡞㧔M. nemestrina㧕ߩᒻᘒቇ⊛Ყセ. ╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢળ (2011/11/04-06, ㇊
ⷓᏒ)㧚ᯅᧄ⵨ሶ (2011) ฎੱ㛽ࠍߤ߁⎇ⓥߔࠆ㧙ขࠅ਄ߍ߆ࠄ଻ሽ㧘޿ߑ⎇ⓥ㧙. ╙ 216 ࿁ ㄭᳯ⽴Ⴆ⎇ⓥળ
(2011/10/29, ṑ⾐⋵)㧚
ự↰ⓛ㧘᧲ፉ⚓ᒎ૫㧘Ძ೑ବ㓶㧘Ꮉᧄ⧐ (2011) ࠲ࠗࡢࡦࠩ࡞㧔Macaca cyclopis㧕ߣ࠾ࡎࡦࠩ࡞㧔Macaca fuscata㧕
ߩ੤㔀୘૕ߦ⷗ࠄࠇࠆየ㐳ࠍ᳿߼ࠆⷐ࿃. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/16-18, ›ጊᏒ)㧚
ᯅᧄ⵨ሶ (2011) ฎੱ㛽ࠍߤ߁⎇ⓥߔࠆ㧙ขࠅ਄ߍ߆ࠄ଻ሽ㧘޿ߑ⎇ⓥ㧙. ╙ 216 ࿁ ㄭᳯ⽴Ⴆ⎇ⓥળ
(2011/10/29, ṑ⾐⋵)㧚
ᯅᧄ⵨ሶ (2011) ੱ㛽ߣ⛗↹⾗ᢱ߆ࠄ⺒ߺขࠆೃ⟏‫ߒࠄߐޟ‬㚂‫ߩޠ‬ᣇᴺ. ⠨ฎቇ⎇ⓥળ╙ 57 ࿁✚ળ࡮⎇ⓥ㓸ળ
(2011/04/23-24, ጟጊᏒ)㧚
ᯅᧄ⵨ሶ㧘⮮↰ዏ㧘Ꮉਭ଻ༀᥓ㧘ᄢ㊁ᙗ੖㧘ᐣ↰ᘕ⍫㧘㋈ᧁ㓉㓶 (2011) ᱤߣਅ㗶㛽߆ࠄ⷗ߚ㖧࿖⑥቟㉿ฎზ
ੱ㛽. ╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢળ (2011/11/04-06, ㇊ⷓᏒ)㧚
ᐔፒ㍈⍫㧘ᾢୖඳ㓶 (2011) ⿷ᐩㇱߩ╭㈩⟎ߣ⿷ᐩ࿶߆ࠄߺߚ㔤㐳㘃ߩ⿷ߩᯏ⢻ゲ. ╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢળ
(2011/11/04-06, ㇊ⷓᏒ)㧚
ᐔ ፒ ㍈ ⍫ 㧘 ᄢ ⍹ ర ᴦ 㧘 ᷡ ᳓ ᄢ テ (2011) 㘃 ੱ ₎ ߩ ⿷ ߩ 㛽 㑆 ╭ ߦ ߟ ޿ ߡ . ╙ 27 ࿁ ᣣ ᧄ 㔤 㐳 㘃 ቇ ળ ᄢ ળ
(2011/07/16-18, ›ጊᏒ)㧚ട⾐⼱⟤ᐘ (2011) ⢷ㇳࡊࡠࡐ࡯࡚ࠪࡦߦߺࠆࠢࡕࠩ࡞ߣࡎࡒࡁࠗ࠼. ╙ 27 ࿁ᣣᧄ
㔤㐳㘃ቇળᄢળ⥄↱㓸ળ (2011/07/16-18, ›ጊᏒ)㧚
ᐔፒ㍈⍫㧘Malaivijitnond S㧘᧲ፉᴕᒎ૫㧘ự↰ⓛ (2011) ࠲ࠗ₺࿖ߦ᫮߻ඨ㊁↢ࡑࠞࠢ 2 ⒳ߩࡠࠦࡕ࡯࡚ࠪࡦ
ߩㆇേቇ⊛ಽᨆ. ╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢળ (2011/11/04-06, ㇊ⷓᏒ)㧚
ട⾐⼱⟤ᐘ (2011) ⢷ㇳࡊࡠࡐ࡯࡚ࠪࡦߦߺࠆࠢࡕࠩ࡞ߣࡎࡒࡁࠗ࠼. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ⥄↱㓸ળ
(2011/07/16-18, ›ጊᏒ)㧚
ട⾐⼱⟤ᐘ (2011) ⡻㛽ߩ⸘㊂⊛․ᓽߦࠃߞߡࡎࡒࡁࠗ࠼ߣ㕖ࡎࡒࡁࠗ࠼ߪ඙೎ߢ߈ࠆ߆㧫 ╙ 65 ࿁ᣣᧄੱ㘃
ቇળᄢળ (2011/11/04-06, ㇊ⷓᏒ)㧚
ਣᯅ⃨᮸㧘Nilpaung W㧘ự↰ⓛ㧘Malaivijitnond S (2011) ࠲ࠗ࡮ࠞࠝࠢ࡜ࡊ࠶ࠢ଻⼔඙ߦ↢ᕷߔࠆࡌ࠾ࠟࠝࠩ
࡞ߩ╙ 1 ૏ࠝࠬ੤ઍߣࠝࠬߩ⒖಴౉. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/16-18, ›ጊᏒ)㧚
ዊᎹ⑲ม㧘Malaivijitnond S㧘ự↰ⓛ (2011) ࡅࠟࠪࠕ࠶ࠨࡓࡕࡦࠠ࡯ߩ੤የⴕേ. Animal 2011㧦ᣣᧄേ‛ᔃℂ
ቇળ㧔╙ 71 ࿁㧕㧘ᣣᧄേ‛ⴕേቇળ㧔╙ 30 ࿁㧕
㧘ᔕ↪േ‛ⴕേቇળ/ᣣᧄኅ⇓▤ℂቇળ (2011 ᐕᐲ)วหᄢળ
(2011/09/09-11, ᧲੩ㇺ᷼඙)㧚
ᄢ㊁ᙗ੖㧘Ꮉਭ଻ༀᥓ㧘⮮↰ዏ㧘ᯅᧄ⵨ሶ㧘㋈ᧁ㓉㓶㧘ᐣ↰ᘕ⍫㧘ዊጊብ⟵㧘ୖጟᤩᄦ (2011) ᐞ૗ቇ⊛ᒻᘒ
᷹ቯቇࠍ↪޿ߚ㖧࿖␞቟㉿࡮൅ፉੱ㛽㗡⬄ߩ 3 ᰴరᒻᘒ⸃ᨆ. ╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢળ (2011/11/04-06, ㇊ⷓ
Ꮢ)㧚
㈬੗᦮ሶ㧘᧻੗ਃᨑ㧘Malkova L㧘ਃ਄┨మ㧘ਛോ⌀ੱ㧘෹᳗㓷Ꮖ㧘㋈ᧁ᮸ℂ㧘ự↰ⓛ㧘↰ਛᱜਯ㧘ችㇱ⾆ሶ㧘
Ꮞፉ⟤ᐘ㧘᧻ᴛື㇢ (2011) ࠴ࡦࡄࡦࠫ࡯ߩᄢ⣖⚵❱ߩ⊒㆐ㆊ⒟ߪࡅ࠻ߩ⣖ㅴൻࠍℂ⸃ߔࠆ਄ߢߩᣂߚߥᵢኤ
㧙㧙
ࠍਈ߃ࠆ. ╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢળ (2011/11/04-06, ㇊ⷓᏒ)㧚
25) ᮭ↰⛗㉿㧘 ጊ৻㆏㧘ự↰ⓛ (2012) ࠻ࡦࠟੱߩ㛽ᩰᚑ㐳ࡄ࠲࡯ࡦߣ⢈ḩߩᐕ㦂ᄌൻ. ╙ 29 ࿁ᣣᧄࠝ࠮ࠕ࠾ࠕ
ቇળᄢળ (2012/03/24-25, ୖᢝᏒ)㧚
26) ự↰ⓛ (2012) ᧲ධࠕࠫࠕߩ㔤㐳㘃㧘․ߦࡑࠞࠢߩᄙ᭽ᕈ⎇ⓥ. ੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲ౒ห೑↪⎇ⓥળ‫ࠫࠕޡ‬
ࠕߩ㔤㐳㘃ߩ଻ోߣ␠ળ↢ᘒ⎇ⓥߦ㑐ߔࠆㄭᐕߩᣂߚߥዷ㐿‫( ޢ‬2012/03/10, ›ጊᏒ)㧚
⻠Ṷ
1)
ᐔፒ㍈⍫ (2011/05) ੑ⿷ࠨ࡞ߩ㛽ᩰ㧘ᱠⴕ㧘⣖. ╙ 26 ࿁ᣣᧄੱ㘃ቇળㅴൻੱ㘃ಽ⑼ળࠪࡦࡐࠫ࠙ࡓ. ੩ㇺᏒ㧚
ࠥࡁࡓᄙ᭽ᕈಽ㊁
㧨⎇ⓥ᭎ⷐ㧪
A) ᨴ⦡૕┵ㇱࡋ࠹ࡠࠢࡠࡑ࠴ࡦߩታᘒ⸃᣿
ฎ⾐┨ᒾ㧘ේᥰ㧘ᐔ੗໪ਭ㧔ㆮવሶᖱႎಽ㊁㧕㧘ᐔ੗⊖วሶ㧔ㆮવሶᖱႎಽ㊁㧕
࠴ࡦࡄࡦࠫ࡯ߩᨴ⦡૕ߩ┵ㇱߦߪ‫ޔ‬ᄢⷙᮨߥࡋ࠹ࡠࠢࡠࡑ࠴ࡦ߇޽ࠆ‫⋧ߦࠇߎޕ‬ᒰߔࠆ᭴ㅧ‛ߪ‫ࠄߺߪߦ࠻ࡅޔ‬
ࠇߥ޿‫౒ޕ‬ㅢ␲వ߆ࠄಽጘߒߚᓟ‫♽ߩ߁߶ߩ࠻ࡅޔ‬೉ߢߎࠇ߇ᶖᄬߒߚߎߣࠍ‫ޔ‬೨ᐕߩ⎇ⓥߢ␜ߒߚ‫ߩߎޕ‬ᶖᄬ߇
ࡅ࠻ߩㅴൻߩේ࿃߹ߚߪ⚿ᨐߢ޽ࠆߎߣߪ‫ޔ‬น⢻ᕈߣߒߡ⠨߃ࠄࠇࠆߚ߼‫ޔ‬ᶖᄬߩᯏ᭴ࠍ⸃᣿ߔࠆߎߣࠍ㐳ᦼ⊛ߦ
⋡ᜰߒߡ޿ࠆ‫ޕ‬1 ߟߩᣇᴺߪ‫ߩߡߒߣ〔∥ߩࡦ࠴ࡑࡠࠢࡠ࠹ࡋߩߎޔ‬Ⴎၮ㈩೉ࠍࡅ࠻ߢ⷗ߟߌ಴ߒ‫ߩߘޔ‬᭴ㅧࠍ⸃
ᨆߔࠆߎߣߢ޽ࠆ‫ ߁߽ޕ‬1 ߟߩࠕࡊࡠ࡯࠴ߣߒߡ‫ޔ‬หߓࡅ࠻਄⑼ߩ࠹࠽ࠟࠩ࡞⑼ߦߺࠄࠇࠆห᭽ߩ⃻⽎ࠍ⺞ߴߡ޿
ࠆ‫ޕ‬
ࡈࠢࡠ࠹࠽ࠟࠩ࡞㧔Symphalangus syndactylus㧕ߦߪᨴ⦡૕┵ㇱߦᄢⷙᮨߥࡋ࠹ࡠࠢ࡞ࡑ࠴ࡦ߇޽ࠅ‫࠽࠹࠹ࡠࠪޔ‬
ࠟࠩ࡞㧔Hylobates lar㧕ߪߎࠇࠍ߽ߚߥ޿‫ߩߣ࠻ࡅߣ࡯ࠫࡦࡄࡦ࠴ޕ‬㑐ଥߦኻᔕߔࠆ‫ᦨޕ‬ೋߩᲑ㓏ߣߒߡ‫ࡠࠢࡈޔ‬
࠹࠽ࠟࠩ࡞ߩࡋ࠹ࡠࠢࡠࡑ࠴ࡦߩታᘒ⸃᣿ࠍ⋡ᜰߒߚ‫ ߩߎޕ‬2 ⒳ߢߩ㊂ߩᏅࠍ೑↪ߒߡࠢࡠ࡯ࡦࠍ․ቯߔࠆታ㛎ࠍ
ⴕ޿‫߇ࡦ࡯ࡠࠢޔ‬ᓧࠄࠇߚ‫ߩߘޕ‬Ⴎၮ㈩೉ߩ⸃ᨆ߆ࠄ‫ ࠻ࠗ࡜࠹ࠨࠔࡈ࡞ࠕޔ‬DNA ߢ޽ࠆߎߣ߇್᣿ߒߚ‫ࡈ࡞ࠕޕ‬
ࠔࠨ࠹࡜ࠗ࠻ DNA ߪ‫ޔ‬㔤㐳㘃ߩ࠹ࡠࡔࠕߩਥᚑಽߣߥߞߡ޿ࠆ࠹ࡋࡠࠢࡠࡑ࠴ࡦߢ޽ࠆ‫ߩࠕࡔࡠ࠹߇ࠇߎޕ‬㗔ၞ
ߢᄢⷙᮨߦჇ᏷ߒߡ޿ࠆߣ޿߁⚿⺰ߦ⥋ߞߚ‫ޕ‬ේ⪺⺰ᢥߣߒߡᛩⓂߒ‫ޔ‬ᡷ⸓ ߇⃻࿷ክᩏਛߢ޽ࠆ‫ޕ‬
B) ࡟࠻ࡠ࠻࡜ࡦࠬࡐ࠱ࡦߩࠥࡁࡓ߳ߩᓇ㗀
ฎ⾐┨ᒾ㧘ේᥰ㧘ᐔ੗໪ਭ㧔ㆮવሶᖱႎಽ㊁㧕㧘ᐔ੗⊖วሶ㧔ㆮવሶᖱႎಽ㊁㧕㧘I Jahan㧔ㆮવሶᖱႎಽ㊁㧕
SVA ࿃ሶߣࠃ߫ࠇࠆ࡟࠻ࡠ࠻࡜ࡦࠬࡐ࠱ࡦߪ‫࠻ࡅޔ‬਄⑼ߩ౒ㅢ␲వߢ↢ߓߚߣ⠨߃ࠄࠇߡ޿ࠆⶄวဳ࿃ሶߢ޽ࠆ‫ޕ‬
ࡅ࠻ߢㆮવሶߩ㗔ၞߦ౉ߞߡ∛᳇ߩේ࿃ߦߥࠆߥߤ‫⃻ޔ‬࿷ߢ߽ォ⒖ᵴᕈࠍ଻ᜬߒߡ޿ࠆ‫ߩߎޕ‬࿃ሶߩࠥࡁࡓ߳ߩᓇ
㗀ߦ㑐ߔࠆ⎇ⓥࠍⴕߞߚ‫ޕ‬
ߎߩ࿃ሶߪ 3 ߟߩ㗔ၞ߆ࠄߥࠅ‫ޔ‬ౝㇱߩ VNTR㧔variable number of tandem repeat㧕ߣࠃ߫ࠇࠆ㗔ၞߪ‫ޔ‬30-50 bp
ߩන૏߇❑೉ߦㅪߥߞߚ෻ᓳ㈩೉ߢ޽ࠆ‫ߩߎޕ‬㗔ၞߩ㐳ߐߪ‫ߪߢ࠻ࡅޕࠆߥ⇣ߦߣߏ࡯ࡇࠦޔ‬ᐔဋߪ⚂ 0.8 kb‫ᦨޔ‬
ᄢߪ⚂ 2.7 kb ߢ޽ࠆߎߣ߇ࠊ߆ߞߡ޿ࠆ‫࡞ࠩࠟ࠽࠹ࠢ࠶ࡠ࡯ࡈޕ‬㧔Hoolock hoolock㧕ߩࠥࡁࡓߢߩߎߩ࿃ሶߩ᭽ᘒ
ࠍ⺞ߴ‫ߣ࠻ࡅޔ‬ห᭽ߩࠦࡇ࡯ߦട߃ߡ㐳ߐ߇ 40 kb ࠍ⿥߃ࠆ VNTR 㗔ၞ߇ᄙᢙሽ࿷ߔࠆߎߣࠍ⷗಴ߒߚ‫ߩߎޕ‬㐳޿
VNTR 㗔ၞߪ࠮ࡦ࠻ࡠࡔࠕߦ޽ࠅ‫ࠍࡦ࠴ࡑࡠࠢࡠࡋ࠹ޔ‬ᒻᚑߒߡ޿ࠆߣ⠨߃ࠄࠇࠆ‫੹ߜࠊߥߔޕ‬࿁ߩ⊒⷗ߪ‫࡜࠻ޔ‬
ࡦࠬࡐ࠱ࡦ߇ᣂⷙߩ࠹ࡋࡠࠢࡠࡑ࠴ࡦࠍᄢ㊂ߦଏ⛎ߔࠆ⃻⽎ߢ޽ࠆ‫ ߽ߦࡓࡁࠥߩ࠻ࡅޕ‬SVA ࿃ሶߪᄙᢙሽ࿷ߔࠆ
ߎߣ߆ࠄ‫߽ߢ࠻ࡅޔ‬ห᭽ߩ⃻⽎߇⿠ߎࠆน⢻ᕈߪ޽ࠆߣ⠨߃ࠄࠇࠆ‫ޕ‬ේ⪺⺰ᢥߣߒߡᛩⓂߒ‫ޔ‬ᡷ⸓ ߇⃻࿷ክᩏਛ
ߢ޽ࠆ‫ޕ‬
C) ࠾ࡎࡦࠩ࡞ߩ㓸࿅ㆮવቇ⊛⎇ⓥ
Ꮉᧄ⧐㧘Ꮉᧄດᳯ㧘ᮘญ⠍ሶ㧘౐ᵄ⟜⡡㧔NPO ᴺੱࠨ࡞ߤߎࡀ࠶࠻㧕㧘㋈ᧁ⟵ਭ㧔NPO ᴺੱࠨ࡞ߤߎࡀ࠶࠻㧕
♮᛽಴ DNA ࠍ೑↪ߒߡಽᨆߒߚ᧲ർ࿾ᣇߩ࠾ࡎࡦࠩ࡞୘૕⟲ߩቅ┙ߦ㑐ߔࠆ⎇ⓥᚑᨐࠍ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળ
ᄢળ㧔›ጊᏒ㧕ߢ⊒⴫ߒߚ‫ߩ࡞ࠩࡦࡎ࠾ޕ‬㊁↢୘૕⟲ߩㆮવ⊛․ᓽࠍ⺞ᩏߔࠆᣂߒ޿⹜ߺߣߒߡ‫ޔ‬ਥⷐ⚵❱ㆡวㆮ
વሶⶄว૕㧔MHC㧕㗔ၞߩㆮવ⊛ᄙ᭽ᕈߦߟ߈‫࠻ࠗ࡜࠹ࠨࡠࠢࠗࡑޔ‬ᐳ૏ࠍಽᨆߔࠆᣇᴺߦߟ߈ᬌ⸛ߒߚ‫ޕ‬⠨᩺
ߒߚᣇᴺߩታ↪ᕈࠍ⹏ଔߔࠆߚ߼‫ޔ‬ችፒ⋵ᐘፉߩ࠾ࡎࡦࠩ࡞ߣߘߩ㑐ㅪ୘૕⟲ߦߟ޿ߡಽᨆߒߡ޿ࠆ‫ޕ‬ਃ㊀⋵ߩ࠾
ࡎࡦࠩ࡞ߦߟ޿ߡ‫੹ޔ‬ᐕᐲ߆ࠄᣂߚߦ౒ห೑↪⎇ⓥߢ࿾ၞ㑆੤ᵹߣᄖ᧪⒳᜛ᢔߩࡕ࠾࠲࡝ࡦࠣࠍ⋡⊛ߣߔࠆ⎇ⓥࠍ
㐿ᆎߒߚ‫ߩ࡞ࠩࡦࡎ࠾ޔߚ߹ޕ‬ᄖ᧪⒳໧㗴ߦߟ޿ߡ‫⎇ߩߢ߹ࠇߎޔ‬ⓥᚑᨐࠍ฽߻✚⺑ࠍනⴕᧄߩ 1 ┨ߣߒߡ಴ ౏
⴫ߒߚ‫ޕ‬
D) ࡑࠞࠞዻࠨ࡞ߩ♽⛔㑐ଥ
Ꮉᧄ⧐㧘Ꮉᧄດᳯ㧘ᮘญ⠍ሶ㧘ự↰ⓛ㧔ㅴൻᒻᘒಽ㊁㧕㧘↰ਛᵗਯ㧘ᄢ੗ᔀ㧔᫪ᨋ✚ว⎇ⓥᚲ㧕㧘ජ‫ޘ‬ጤື㧔㧔ᩣ㧕
࡜࡯ࠧ㧕
㧘P Wangda㧔ࡀࡄ࡯࡞ㄘᬺ⋭㧕㧘MA Haffman㧔␠ળㅴൻಽ㊁㧕㧘CAD Nahallage㧔Sri Jayawardenepura ᄢቇ㧕㧘
M Chalise㧔Tribhuvan ᄢቇ㧕㧘⯃⑲ᘢ㧔บḧ࿡┙ዳ᧲⑼ᛛᄢቑ㧕
6 ᦬ߦࠬ࡝࡜ࡦࠞߢ㐿௅ߐࠇߚ࿖㓙ࠪࡦࡐࠫ࠙ࡓߦෳടߒ‫౒ޔ‬ห⎇ⓥߢߔߔ߼ߡ޿ࠆ࠻ࠢࡕࡦࠠ࡯ߩ♽⛔࿾ℂ⎇
ⓥߩᚑᨐࠍ⊒⴫ߒߚ‫ޕ‬9 ᦬ߣ 12 ᦬ߦࡉ࡯࠲ࡦࠍ⸰໧ߒ‫ޔ‬ㄘᬺ⋭ߣㅴ߼ߡ޿ࠆ࠹࡟ࡔ࠻࡝࡯ߦࠃࠆࠕ࠶ࠨࡓࡕࡦࠠ
࡯ߩ↢ᘒ⺞ᩏ⚿ᨐࠍ⸃ᨆߔࠆߣߣ߽ߦ‫⎇ޔ‬ⓥᛂߜวࠊߖࠍⴕߞߚ‫ޕ‬10 ᦬ߦ⯃⑲ᘢ᳁ࠍ᜗߈‫ޔ‬ᣣᧄߢ㊁↢ൻߒߚ࠲
㧙㧙
ࠗࡢࡦࠩ࡞ߣบḧ࿷᧪⒳ߩᲧセࠍ⎇ⓥᚲ౒ห೑↪⎇ⓥߣߒߡ㐿ᆎߒߚ‫ޔߚ߹ޕ‬1 ᦬ߦ Chalise ᳁ߣ Nahallage ᳁ࠍ᜗
߈‫ޔ‬ධࠕࠫࠕߩ㔤㐳㘃ߩㅪ៤⎇ⓥߦะߌߚࡢ࡯࡚ࠢࠪ࠶ࡊࠍ㐿௅ߒߚ‫ޕ‬3 ᦬ߦ⎇ⓥᚲ౒ห೑↪⎇ⓥળߢධࠕࠫࠕߩ
㔤㐳㘃⎇ⓥ‫⛔♽ߦ․ޔ‬࿾ℂ⎇ⓥߩ⃻⁁ߣ⺖㗴ߦߟ߈⻠Ṷߒߚ‫ޕ‬
ࠠ࠲ࡉ࠲ࠝࠩ࡞ߩ♽⛔࿾ℂቇ⊛⎇ⓥࠍߔߔ߼ࠆߚ߼‫ޔ‬9 ᦬ߦ࠲ࠗධㇱߢࡍ࠶࠻ࠩ࡞ߩត⚝߅ࠃ߮ណ᧚‫↢ޔ‬ᕷ࿾ߩ
ⷰኤࠍⴕߞߚ‫ޔߚ߹ޕ‬2012 ᐕ 1 ᦬㨪2 ᦬ߦ࠲ࠗ࡮࠴ࡘ࡜ࡠࡦࠦ࡯ࡦᄢቇࠃࠅ⎇ⓥഥᚑࠍฃߌ‫ࠗ࠲ޔ‬ർㇱߢߩࠠ࠲ࡉ
࠲ࠝࠩ࡞ߩ⺞ᩏߣ࠴ࡘ࡜ࡠࡦࠦ࡯ࡦᄢቇℂቇㇱߦߡታ㛎ቶ૞ᬺࠍⴕߞߚ‫ޕ‬
E) ࡏࡁࡏߩ଻ోㆮવቇ⊛⎇ⓥ
Ꮉᧄ⧐㧘ᮘญ⠍ሶ㧘ฎᏒ೰ผ㧔␠ળㅴൻಽ㊁㧕㧘┻రඳᐘ㧔␠ળㅴൻಽ㊁㧕㧘ဈᏎື਽㧔␠ળㅴൻಽ㊁㧕㧘ᯅᧄජ⛗
㧔↢ᘒ଻ోಽ㊁㧕
⛘Ṍෂᯏߦ޽ࠆࡏࡁࡏࠍኻ⽎ߦ‫ࠧࡦࠦޔ‬᳃ਥ౒๺࿖ߢ㐳ᦼⷰኤ߇ⴕࠊࠇߡ޿ࠆ 6 ࠞᚲ㧔Wamba, Iyondji, Lac Tumba,
TL2, Salonga, Lomako㧕ߢ♮⹜ᢱࠍ෼㓸ߒ‫ޔ‬ㆮવሶߩᄙ᭽ᕈߣ࿾ၞ୘૕⟲㑆ߩㆮવಽൻࠍಽᨆߒߡ޿ࠆ‫ߩࠄ߆♮ޕ‬
DNA ಽᨆᴺࠍᡷༀߒߚߎߣߦࠃࠅ‫ߦࠄߐޔ‬ᄙ⒳㘃ߩㆮવሶಽᨆ߇น⢻ߦߥߞߚ‫⃻ޕ‬࿷‫ࠕ࡝࠼ࡦࠦ࠻ࡒޔ‬ㆮવሶ‫ޔ‬Y
ᨴ⦡૕ㆮવሶ‫ޔ‬Ᏹᨴ⦡૕ࡑࠗࠢࡠࠨ࠹࡜ࠗ࠻ߩᄌ⇣ࠍ୘૕⟲ߏߣߦಽᨆߒߡ޿ࠆ‫ޕ‬
F) ࡑࠞࠢࠩ࡞ࠦࡠ࠾࡯ߩ㓸࿅ㆮવቇ⊛⎇ⓥ
↰ਛᵗਯ㧘᫪ᧄ⌀ᑿ㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕㧘㊍ਛᘮᦶ㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕㧘Ꮉᧄດᳯ㧘Ꮉᧄ
⧐
㔤㐳㘃⎇ⓥᚲߢ⛽ᜬߐࠇߡ޿ࠆ࠾ࡎࡦࠩ࡞߅ࠃ߮ࠕࠞࠥࠩ࡞ߩ❥ᱺࠦࡠ࠾࡯ߦߟ޿ߡ‫ޔ‬ㆮવ⊛ᄙ᭽ᕈߩ⚻ᤨ⊛ᄌ
ൻࠍ᣿ࠄ߆ߦߔࠆߚ߼‫ޔ‬ಽᨆࠍߔߔ߼ߚ‫࡞ࠩࠥࠞࠕޕ‬ਛ࿖⟲߅ࠃ߮࠾ࡎࡦࠩ࡞㜞ᵿ⟲ߣ߽ߦ‫⺞ޔ‬ᩏኻ⽎ᦼ㑆ਛ‫ޔ‬ᓢ‫ޘ‬
ߦㆮવ⊛ᄙ᭽ᕈ߇ૐਅߒ‫✼ⴊޔ‬ᐲ߇਄᣹ߒߡ޿ࠆߎߣ߇᣿ࠄ߆ߦߥߞߚ‫ޔࠄ߇ߥߒ߆ߒޕ‬ଐὼߣߒߡ࠾ࡎࡦࠩ࡞㊁
↢⟲ߣห╬ߩㆮવ⊛ᄙ᭽ᕈࠍ଻ᜬߒߡ޿ࠆߎߣ߇ࠊ߆ߞߚ‫ޕ‬એ਄ߩߎߣࠍ‫⑼ޔ‬ቇ⎇ⓥ⾌⵬ഥ㊄⎇ⓥᚑᨐႎ๔ᦠ㧔ၮ
⋚⎇ⓥ(C)㧕ߣߒߡ߹ߣ߼‫ޔ‬ឭ಴ߒߚ‫ޕ‬
G) ࡢࠝࠠ࠷ࡀࠩ࡞ߥࠄ߮ߦᏗዋ࡟ࡓ࡯࡞ߩ㓸࿅ㆮવቇ⊛⎇ⓥ
Ꮉᧄ⧐㧘Ꮢ㊁ㅴ৻㇢㧘㜞⇌↱⿠ᄦ㧔㑐⷏ቇ㒮ᄢቇ㧕㧘⨥⼱⮍㧔ฬฎደ⧓ⴚᄢቇ㧕㧘૒⮮ብ᮸㧔੩ㇺᄢቇࠕࠫࠕ࡮ࠕࡈ
࡝ࠞ࿾ၞ⎇ⓥ⎇ⓥ⑼㧕㧘↰ਛᵗਯ㧘ቬㄭഞ㧔㧔⽷㧕ㅴൻ↢‛ቇ⎇ⓥᚲ㧕
8 ᦬ߦࡑ࠳ࠟࠬࠞ࡞ߩࡌ࡟ࡦ࠹ࠖ଻⼔඙ߢࡢࠝࠠ࠷ࡀࠩ࡞ߩ⺞ᩏࠍⴕߞߚ‫ޕ‬8 ⟲ࠍో㗡᝝₪ߒ‫⸘ޔ‬᷹⾗ᢱߣㆮવ
ሶ⹜ᢱࠍᓧߚ‫ޕ‬Ꮢ㊁ࠍਛᔃߦⴊ✼᭴ㅧߣ⟲㑆ಽൻߩᬌ⸛ࠍㅴ߼ߡ޿ࠆ‫ޕ‬
ේ↥࿖ߢ⛘Ṍ߇ෂ߱߹ࠇߡ޿ࠆࠢࡠࠠ࠷ࡀࠩ࡞‫ߦ࡞ࠩࡀ࠷ࠠࠠࡑ࡝ࠛࠞࠕ߮ࠃ߅࡞ࠩࡀ࠷ࠠࠠࡑ࡝ࠛࡠࠢࡠࠪޔ‬
ߟ޿ߡ‫ޔ‬ᣣᧄ࿖ౝߩേ‛࿦╬ߢ㘺⢒ߐࠇࠆ୘૕⟲ࠍኻ⽎ߦ‫ߩ࠻ࠗ࡜࠹ࠨࡠࠢࠗࡑޔ‬ಽሶࡑ࡯ࠞ࡯ߩ㐿⊒ࠍⴕߞߚ‫ޕ‬
ࠢࡠࠠ࠷ࡀࠩ࡞ߦߟ޿ߡߪ‫ޔ‬14 ㆮવሶᐳ߇ᄙဳ⊛ߢ޽ࠅ‫ⷫޔ‬ሶ್ቯߦ೑↪ߔࠆߎߣ߇ߢ߈ߚ‫ࠩࡀ࠷ࠠࠠࡑ࡝ࠛޕ‬
࡞ߦߟ޿ߡߪ‫ޔ‬ᣣᧄ࿖ౝߩ߶ߣࠎߤߩ୘૕ߦߟ޿ߡ‫ޔ‬7 ㆮવሶᐳߩ್ቯࠍ⚳߃ߚ‫ޕ‬
H) ኅ⇓ൻ⃻⽎ߣኅ⇓♽⛔ผߩ⎇ⓥ
Ꮉᧄ⧐㧘Ⓑ᧛ື਽㧔ᗲ⍮⋵┙ᄢቇ㧕㧘T Dorji㧔Bhutan ㄘᬺ⋭㧕
㧘ᄢጊୃ৻㧔੩ㇺᄢቇࠕࠫࠕ࡮ࠕࡈ࡝ࠞ࿾ၞ⎇ⓥ⎇ⓥ
⑼㧕
㧘ᧄᳯᤘᄦ㧔Ꮺᐢ⇓↥ᄢቇ㧕㧘⧛⼱ᗲᒾ㧔ኾୃᄢቇ㧕㧘㠽Ⴆ޽ࠁߜ㧔᧲ᶏᄢቇ㧕
ࡉ࡯࠲ࡦߣࠕ࡞࠽࡯࠴ࡖ࡞ࡊ࡜࠺࡯ࠪࡘ㧔ࠗࡦ࠼㧕ߦ߅ߌࠆ࠙ࠪ⑼ኅ⇓ࡒ࠲ࡦߩ೑↪ߣㆮવ⊛․ᕈߦߟ߈‫ࠇߎޔ‬
߹ߢߩ⎇ⓥᚑᨐࠍ 11 ᦬ߦ੩ㇺߢ㐿௅ߐࠇߚ࿖㓙ળ⼏ߢ⊒⴫ߒߚ‫ޕ‬ධ☨ࠕࡦ࠺ࠬ㜞࿾ߦ߅ߌࠆ࡜ࠢ࠳⑼ኅ⇓ߩ⎇ⓥ
ߢߪ‫ޔ‬
8 ᦬ߦࡍ࡞࡯ߢࠕࡗࠢ࡯࠴࡚⋵ߣࠢࠬࠦ⋵ߢⷰኤߣ⹜ᢱណขࠍⴕߞߚ‫ߩ࠻ࠗ࡜࠹ࠨࡠࠢࠗࡑޕ‬ᄙဳಽᨆ߆ࠄ‫ޔ‬
࡝ࡖࡑߣࠕ࡞ࡄࠞߩኅ⇓ൻߦ੤㔀߇㑐ਈߔࠆߣߩ␜ໂࠍᓧߚ‫⎇ߩߎޕ‬ⓥᚑᨐࠍ 3 ᦬ߦ㐿௅ߐࠇߚᣣᧄ⇓↥ቇળ╙
115 ࿁ᄢળ㧔ฬฎደ㧕ߢ⊒⴫ߒߚ‫ࡗ࡜ࡑࡅޕ‬㜞࿾ߩ⇓↥ߣ’⇓ߩ⎇ⓥߢߪ‫ޔ‬9 ᦬ߦࡉ࡯࠲ࡦ᧲ㇱߩࡔ࡜࠶ࠢ࿾ၞߢ
’᳃ߣኅ⇓ࠍ⺞ᩏߒ‫ޔ‬3 ⒳㘃ߩኅ⇓㧔࿷᧪‐‫ࡦ࠲ࡒޔࠢࡗޔ‬㧕߇ਃర੤㔀ߒߡ޿ࠆߎߣࠍೋ߼ߡ⏕⹺ߒߚ‫⃻ޕ‬࿷‫ޔ‬
ߎߩ࿾ၞߩኅ⇓ߩㆮવሶಽᨆࠍ⸘↹ߒߡ޿ࠆ‫ޕ‬
I) 㔤㐳㘃ߩ᳃ଶ↢‛ቇ⊛⎇ⓥ
Ꮉᧄ⧐㧘ਃᚭᐘਭ㧔NPO ᴺੱ࠾ࡎࡦࠩ࡞ࡈࠖ࡯࡞࠼ࠬ࠹࡯࡚ࠪࡦ㧕
⇓⥢ߦࠨ࡞㛽ࠍ␢ࠆାઔߩ⸥㍳ߣ‫⛘ޔ‬Ṍ࿾ߦ↢ᕷߒߚ࠾ࡎࡦࠩ࡞ߩ DNA ࠍಽᨆߔࠆ⋡⊛ߢ‫࡞ࠨߚࠇࠄ␢ޔ‬㛽㧔෡
₎ߣࠃ߫ࠇࠆ㧕ࠍฦ࿾ߢត⚝ߒߡ޿ࠆ‫ޕ‬ᣂߚߦ㐳㊁⋵ߩ⨆㊁Ꮢߢ⊒⷗ߒ‫ޔ‬㗡⬄㛽ߦᱷࠆ DNA ࠍಽᨆߒߚ‫ޕ‬㐳㊁⋵
ߢߪᧂ⊒⷗ߩࡒ࠻ࠦࡦ࠼࡝ࠕ DNA ࠲ࠗࡊ߇⏕⹺ߢ߈‫ޔ‬ਛㇱጊጪ࿾Ꮺߩ࠾ࡎࡦࠩ࡞୘૕⟲ߩᚑ┙ࠍ⠨߃ࠆᖱႎߣߥ
ߞߚ‫ޔߚ߹ޕ‬7 ᦬ߦ㚅Ꮉ⋵ߢ෡₎ߦ㑐ߔࠆ⡞߈ㄟߺ⺞ᩏࠍⴕߞߚ‫ޕ‬
J) ࡂ࠽ࡃ࠴ߩᱧผ↢‛࿾ℂቇ
↰ਛᵗਯ
7 ᦬ߦ㐳㊁⋵ߣർᶏ㆏ߢᬀ↢᥊ⷰߣࡑ࡞ࡂ࠽ࡃ࠴ߩಽᏓߩ㑐ଥߦ㑐ߔࠆ⺞ᩏࠍⴕ޿‫⨲ޔ‬ේᕈࡑ࡞ࡂ࠽ࡃ࠴ߩࡑࠗ
ࠢࡠࠨ࠹࡜ࠗ࠻ DNA ߩಽᨆࠍߔߔ߼ߚ‫ޕ‬
㧙㧙
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
2)
Koga A, Sasaki S, Naruse K, Shimada A, Sakaizumi M (2011) Occurrence of a short variant of the Tol2 transposable
element in natural populations of the medaka fish. Genetics Research 93(1):13-21.
Koga A, Notohara M, Hirai H (2011) Evolution of subterminal satellite (StSat) repeats in hominids. Genetica
139(2):167-175.
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
2)
3)
Koga A (2011) Contribution of DNA-based transposable elements to genome evolution: inferences drawn from behavior
of an element found in fish (Chapter 6). (Post-Genome Biology of Primates) (ed. Go, Imai, Hirai) Springer.
Koga A (2011) Transposable elements Tol1 and Tol2. (Medaka - Model for Organogenesis, Human Diseases and
Evolution) (ed. Naruse, Takeda, Tanaka) p.171-180 Springer.
⊕੗໪㧘Ꮉᧄ⧐ (2011) ࠲ࠗࡢࡦࠩ࡞ߣࠕࠞࠥࠩ࡞㧦੤㔀࿁ㆱߩߚ߼ߩᩮ⛘⸘↹. ‫ޟ‬ᣣᧄߩᄖ᧪ູ੃㘃‫( ޠ‬ጊ
↰ᢥ㓶㧘ᳰ↰ㅘ㧘ዊୖ೰✬) p.169-202 ᧲੩ᄢቇ಴ ળ㧚
ߘߩઁߩၫ╩
1)
Ꮉᧄ⧐ (2011) ࡉ࡯࠲ࡦߦ߅ߌࠆ㊁↢േ‛ߣߩኻ┙. ⑼ቇ 81: p.578㧚
ቇળ⊒⴫
ự↰ⓛ㧘᧲ፉᴕᒎ૫㧘Ძ೑ବ㓶㧘Ꮉᧄ⧐ (2011) ࠲ࠗࡢࡦࠩ࡞㧔Macaca cyclopis㧕ߣ࠾ࡎࡦࠩ࡞㧔Macaca fuscata㧕
ߩ੤㔀୘૕ߦ⷗ࠄࠇࠆየ㐳ࠍ᳿߼ࠆⷐ࿃. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/18, ›ጊᏒ)㧚
2) ᐔ੗໪ਭ㧘ᐔ੗⊖วሶ㧘ฎ⾐┨ᒾ㧘㡻Ლବผ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ޽ߞߡࡅ࠻ߦߥ޿ᨴ⦡૕┵ㇱࠥࡁࡓਇᲫ
࿾Ꮺ㧦(1) ሽ࿷᭽ᑼᄌ⇣߆ࠄផ᷹ߐࠇࠆ㕖⋧หᨴ⦡૕㑆ᧃ┵⚵឵߃. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/18,
›ጊᏒ)㧚
3) Ꮉᧄ⧐㧘ਃᚭᐘਭ㧘ᮘญ⠍ሶ㧘Ꮉᧄດᳯ (2011) ᵤシඨፉ୘૕⟲ߩㆮવ⊛․ᓽ߆ࠄߺߚർ㒢ߩ࠾ࡎࡦࠩ࡞ߩᚑ
┙. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/17, ›ጊᏒ)㧚
4) 㘑ᒛ༑ሶ㧘੗਄⧷ᴦ㧘Ꮉᧄ⧐㧘ਛᎹዏผ㧘੗਄㧙᧛ጊ⟤Ⓞ (2011) ㊄⪇ጊߩ࠾ࡎࡦࠩ࡞ߩㆮવ⊛ᄙ᭽ᕈ. ╙ 27
࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/7/17, ›ጊᏒ)㧚
5) ฎ⾐┨ᒾ (2011) ࡔ࠳ࠞߦߺࠄࠇࠆ 2 ߟߩኻᾖ⊛ߥォ⒖࿃ሶ㧦ଚ౉⋥ᓟߩర᳇ߥ࿃ሶߣ፣უߩෂᯏߦ┙ߟ࿃ሶ.
ᣣᧄㆮવቇળ╙ 83 ࿁ᄢળ (2011/09/20, ੩ㇺᏒ)㧚
6) ฎ⾐┨ᒾ㧘ᐔ੗⊖วሶ㧘ᐔ੗໪ਭ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ޽ߞߡࡅ࠻ߦߥ޿ᨴ⦡૕┵ㇱࠥࡁࡓਇᲫ࿾Ꮺ㧦(2) ၭ
㙃⚦⢩ࠍ↪޿ߚ⚵឵߃ߩᬌ಴. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/18, ›ጊᏒ)㧚
7) Michimoto K, Takami Y, Tanaka H, Ushimaru A (2012) Effects of urbanization on genetic structures in two
paddy-associated frog species with different dispersal habits. ᣣᧄ↢ᘒቇળ╙ 59 ࿁ᄢળ㧛╙ 5 ࿁᧲ࠕࠫࠕ↢ᘒቇળ
ㅪวᄢળ (2012/02/19, ᄢᵤᏒ)㧚
8) Suka T, Ushimaru A, Tanaka H, Yumoto T (2012) A post-gracial history of semi-natural grassland in central Japan and
its causation to distribution of grassland species of bumblebees. ᣣᧄ↢ᘒቇળ╙ 59 ࿁ᄢળ㧛╙ 5 ࿁᧲ࠕࠫࠕ↢ᘒቇ
ળㅪวᄢળ (2012/02/18, ᄢᵤᏒ)㧚
9) Tanaka H (2012) Genentic constitution and species status of the Bornean white-bearded gibbon (Hylobates albibarbis)
as inferred by amplified fragment length polymorphism (AFLP). ᣣᧄ↢ᘒቇળ╙ 59 ࿁ᄢળ㧛╙ 5 ࿁᧲ࠕࠫࠕ↢ᘒቇ
ળㅪวᄢળ (2012/02/20, ᄢᵤᏒ)㧚
10) Ꮉᧄ⧐㧘Ⓑ᧛ື਽㧘ᄢጊୃ৻㧘ᧄᳯᤘᄦ㧘⧛⼱ᗲᒾ㧘㠽Ⴆ޽ࠁߜ (2012) ධ☨࡜ࠢ࠳⑼േ‛ߩ⿠Ḯߦ㑐ߔࠆㆮ
વቇ⊛⎇ⓥ㧙ࡍ࡞࡯ߩࡆࠢ࡯࠾ࡖ㧘࡝ࡖࡑ㧘ࠕ࡞ࡄࠞߩㆮવሶᄙ᭽ᕈߣ੤㔀ߩ⺞ᩏ⚿ᨐ㧙. ᣣᧄ⇓↥ቇળ╙ 115
࿁ᄢળ (2012/03/29, ฬฎደᏒ)㧚
11) ᄢ੗ᔀ㧘ᷡ㊁⚔ౖ㧘ựፒિ৻㇢㧘Ꮉᧄ⧐ (2012) ቟ቯห૏૕Ყಽᨆߦࠃࠅ᣿ࠄ߆ߦߒߚ࠾ࡎࡦࠩ࡞ߩ㘩ᕈߩ⟲
ࠇౝᄌ⇣ߦߟ޿ߡ. ᣣᧄ↢ᘒቇળ╙ 59 ࿁ో࿖ᄢળ (2012/03/19, ᄢᵤᏒ)㧚
1)
⻠Ṷ
1)
2)
3)
4)
5)
Kawamoto Y (2011/06/21) Genetic diversity of macaques in Sri Lanka. International Symposium “Integrative Research
on Monkeys Man and Malaria in Asia. Colombo, Sri Lanka㧚
Kawamoto Y, Dorji T, Inamura T (2011/11/25) Utilization of mithun in Himalaya: A population genetic study on the
use of its hybrids in Bhutan. International Conference "Quality of life and optimal aging learning from wisdom of
highland civilizations". Kyoto.
Ꮉᧄ⧐ (2011/04/09) ࡉ࡯࠲ࡦߩࠨ࡞⺞ᩏ㧔⛯ႎ㧕. ੩ㇺᄢቇࡉ࡯࠲ࡦ෹ᅢࡊࡠࠣ࡜ࡓ ศ↰ᴰᲚ⥄ὼቇ࠮ࡒ࠽
࡯. ੩ㇺᏒ㧚
Ꮉᧄ⧐ (2012/03/10) ධࠕࠫࠕߩ㔤㐳㘃ߩ♽⛔࿾ℂ⎇ⓥ. ੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲ౒ห೑↪⎇ⓥળ‫ߩࠕࠫࠕޡ‬㔤㐳
㘃ߩ଻ోߣ␠ળ↢ᘒ⎇ⓥߦ㑐ߔࠆㄭᐕߩᣂߚߥዷ㐿‫ޢ‬. ›ጊᏒ㧚
ฎ⾐┨ᒾ (2011/09/25) ࠴ࡦࡄࡦࠫ࡯ߦ޽ߞߡࡅ࠻ߦߥ޿ᄢⷙᮨ෻ᓳ㈩೉. ੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲ᧲੩౏㐿⻠
ᐳ. ᧲੩ㇺ㧚
㧙㧙
♽⛔⊒↢ಽ㊁
㧨⎇ⓥ᭎ⷐ㧪
A) ᧲ㇱ࡙࡯࡜ࠪࠕ࿾ၞߦ߅ߌࠆᣂ╙ਃ♿ߩ㔤㐳㘃ㅴൻߦ㑐ߔࠆ⎇ⓥ
A-1) ࡒࡖࡦࡑ࡯↥ࠝ࠽ࠟࠩ࡞਄⑼ൻ⍹ߩ⎇ⓥ
㜞੗ᱜᚑ㧘⷏᧛೰㧘ᳯᧁ⋥ሶ㧘⍫㊁⥶㧘⷏ጟ૓৻㇢
ࡒࡖࡦࡑ࡯ߩ㞲ᣂ਎ޯᦝᣂ਎ߩ࿾ጀࠍኻ⽎ߦ㔤㐳㘃ࠍਛᔃߣߒߚູ੃㘃ൻ⍹ߩ⊒ជ⺞ᩏࠍ߅ߎߥߞߚ‫ࡦࠗࡖ࠴ޕ‬
ࠩ࠙࠶ࠢ࿾ၞߩਛᣂ਎ᧃޯ㞲ᣂ਎ೋ㗡ߩ࿾ጀ߆ࠄߪᣂ⒳ߩࠦࡠࡉࠬ੝⑼ߩൻ⍹߇⊒⷗ߐࠇߡ߅ࠅ‫⸥ޔ‬タ૞ᬺࠍⴕߞ
ߡ޿ࠆ‫ࡦࡆࠚ࠙ࠣޕ‬࿾ၞߩ㞲ᣂ਎ᓟඨߩ࿾ጀ߆ࠄߪ Semnopithecus㧔ࡂ࠿ࡑࡦ࡜ࡦࠣ࡯࡞ዻ㧕ߣᕁࠊࠇࠆൻ⍹߇ᄙ
ᢙ⷗ߟ߆ߞߡ߅ࠅ‫ߩߘޔ‬ಽ㘃࡮⸥タ૞ᬺࠍㅴ߼ߡ޿ࠆ‫࡯ࡌࠨߚ߹ޕ‬࿾ၞߩ೨ᦼ㧔?㧕ᦝᣂ਎ߣᕁࠊࠇࠆ࿾ጀ߆ࠄ⊒
⷗ߐࠇߡ޿ߚᄢဳߩࠝ࠽ࠟࠩ࡞੝⑼ߩൻ⍹ߩ♽⛔⊛૏⟎ߦ㑐ߔࠆᬌ⸛߽ㅴ߼ߡ޿ࠆ‫ޕ‬
A-2) ␹ᄹᎹ⋵↥ࠦࡠࡉࠬൻ⍹ߩ⎇ⓥ
⷏᧛೰㧘㜞੗ᱜᚑ
ᓟᦼᦝᣂ਎␹ᄹᎹ⋵↥ߩࠦࡠࡉࠬൻ⍹ߩಽ㘃ߩౣᬌ⸛ࠍⴕߞߚ‫࡞ࠞࠗࡃࠬࡦ࡜࠻ޔࠬࠢ࠹ࡇࠦ࡝࠼↥ࠬࡦ࡜ࡈޕ‬
↥ࡄ࡜ࡊ࡟ࠬࡃࠗ࠹ࠖࠬ‫ޔ‬ਛ࿖↥ࠠࡦࠪࠦ࠙ൻ⍹ߥߤߣᲧセᬌ⸛ߒ‫ࠬࠢ࠹ࡇࠦ࡝࠼ߢ߹ࠇߎޔ‬ዻߩ੝ዻߢ޽ߞߚࠞ
࠽ࠟࡢࡇ࠹ࠢࠬࠍዻฬߣߒߡᣂዻߣߒߡႎ๔ߒߚ‫↥ࠞ࡝ࡈࠕޔߚ߹ޕ‬ൻ⍹ࠦࡠࡉࠬ㘃߿⃻↢ࠕࠫࠕ↥ࠦࡠࡉࠬ㘃ߣ
ߩ♽⛔⊛㑐ㅪᕈߦߟ޿ߡ߽ᬌ⸛ߒߚ‫ޕ‬
A-3) ਛ࿖↥ᄢဳࡅࡅᣖൻ⍹ߩ⎇ⓥ
⷏᧛೰㧘㜞੗ᱜᚑ㧘⍫㊁⥶㧘દ⮮Პ
ᦝᣂ਎᧲࡮ධ࡙࡯࡜ࠪࠕ↥ࡊࡠࠨࠗࡁ࠮ࡈࠔ࡞ࠬߣ⷏࡙࡯࡜ࠪࠕ↥ࡄ࡜࠼࡝ࠦࡇ࠹ࠢࠬߩಽ㘃ߩౣᬌ⸛ࠍⴕߞߡ
޿ࠆ‫ࡅࡅ↢⃻ޕ‬ᣖ߿ࡑࠞࠢߩ㗡⬄㛽ߩ CT ↹௝࠺࡯࠲ࠍ♖ᩏߒ‫ޔ‬ਛ࿖↥ࡊࡠࠨࠗࡁ࠮ࡈࠔ࡞ࠬൻ⍹ߩ㜞⸃௝ᐲ CT
↹௝߆ࠄ‫⛔♽ߩߘޔ‬ቇ⊛૏⟎ࠍផቯߢ߈ߥ޿߆ᬌ⸛ߒߚ‫ޕ‬
A-4) บḧ↥ࠝ࠽ࠟࠩ࡞⑼ൻ⍹ߩ⎇ⓥ
㜞੗ᱜᚑ
บḧ࿖┙⥄ὼ⑼ቇඳ‛㙚ߩᒛ㊒⠍ඳ჻ߣ౒หߢ‫ޔ‬บḧධㇱߩਛᦼᦝᣂ਎ߩ࿾ጀ߆ࠄ⷗ߟ߆ߞߚࠝ࠽ࠟࠩ࡞⑼ߩ߽
ߩߣ⠨߃ࠄࠇࠆㆆ㔌ᱤൻ⍹ߩ⸥タ‫ߦ߮ࠄߥޔ‬ฎ↢‛࿾ℂቇ⊛⎇ⓥࠍⴕߞߚ‫ޕ‬
A-5) ਛ࿖↥ࡑࠞࠢൻ⍹ߩ㗡㛽ౝㇱᒻᘒߦ㑐ߔࠆ⎇ⓥ
દ⮮Პ㧘⷏᧛೰㧘㜞੗ᱜᚑ
ਛ࿖↥ Macaca anderssoni ߩൻ⍹ᮡᧄࠍ CT ᠟௝ߒ‫ߩߘޔ‬㗡㛽ౝㇱ᭴ㅧߩ⸃ᨆߣ⃻↢⒳ߣߩᲧセࠍⴕ޿‫⛔♽ߩߘޔ‬
⊛૏⟎ߦߟ޿ߡᬌ⸛ߒߚ‫ޕ‬
A-6) ᣣᧄ೉ፉߦ߅ߌࠆ࠾ࡎࡦࠩ࡞ߩㅴൻߦ㑐ߔࠆฎ↢‛ቇ⊛⎇ⓥ
⷏ጟ૓৻㇢㧘㜞੗ᱜᚑ㧘⷏᧛೰㧘દ⮮Პ
ᣣᧄ೉ፉߩ╙྾♿ߩᵢ─࡮ⵚߞ߆ၸⓍ‛‫✽߮ࠃ߅ޔ‬ᢥㆮ〔߆ࠄ⷗ߟ߆ߞߡ޿ࠆ࠾ࡎࡦࠩ࡞ൻ⍹ࠍ⸃ᨆߒ‫ߩߘޔ‬ᒻ
ᘒ⊛ᄌൻߣ࿾ℂ⊛ಽᏓߩᄌㆫߦ㑐ߒߡ‫ޔ‬ฎ↢‛ቇ⊛ߦᬌ⸛ߒߚ‫࡞ࠩࡦࡎ࠾ޔߚ߹ޕ‬ൻ⍹ߩ↥࿾ߣᮡᧄߩ࠺࡯࠲ࡌ࡯
ࠬࠍ૞ᚑߒ‫ ⸘ޔ‬27 ࿾ὐߩᮡᧄࠍᢛℂߒߚ‫ޕ‬
A-7) ᦺ㞲ඨፉߩࡑࠞࠢൻ⍹ߩᬌ⸛
㜞੗ᱜᚑ
㖧࿖వผᢥൻ⎇ⓥ㒮ߩ᧘㓉ഥᢎ᝼ߣ౒หߢ‫ᦺޔ‬㞲ඨፉߩᦝᣂ਎ߩㆮ〔߆ࠄ⊒⷗ߐࠇߡ޿ࠆࡑࠞࠢൻ⍹ߩᬌ⸛ࠍⴕ
ߞߚ‫ޕ‬
A-8) ࠗࡦ࠼ࡀࠪࠕߩࡑࠞࠢൻ⍹ߩᬌ⸛
㜞੗ᱜᚑ
ࠗࡦ࠼ࡀࠪࠕߩࠛࡀ࡞ࠡ࡯㋶‛⾗Ḯ⋭ᐡߩࠕࠫࠬඳ჻ߣ౒หߢ‫ࡢࡖࠫޔ‬ፉ߆ࠄ⷗ߟ߆ߞߡ޿ࠆᦝᣂ਎ߩ㔤㐳㘃ൻ
⍹ߩⷰኤߣ⸘᷹ࠍⴕ޿‫᧲ߦ․ޔ‬ධࠕࠫࠕߩࠝ࠽ࠟࠩ࡞⑼ߩࠨ࡞ߩㅴൻผߦߟ޿ߡᬌ⸛ߒߡ޿ࠆ‫ޕ‬
B) ᧲ㇱ࡙࡯࡜ࠪࠕ࿾ၞߦ߅ߌࠆฎ╙ਃ♿ߩ㔤㐳㘃ㅴൻߦ㑐ߔࠆ⎇ⓥ
㜞੗ᱜᚑ㧘⷏᧛೰㧘ᳯᧁ⋥ሶ㧘⷏ጟ૓৻㇢
ࡒࡖࡦࡑ࡯ߩࡐࡦ࠳࠙ࡦ࿾ၞߦᐢ߇ࠆਛᦼᆎᣂ਎ᧃߩ࿾ጀ߆ࠄ↥಴ߔࠆ㔤㐳㘃ൻ⍹ߪ‫ޔ‬ේᆎ⊛ߥᦛ㥦₎㘃ߣ⌀₎
㘃ߩਛ㑆⊛ߥᒻᘒࠍ␜ߒ‫₎⌀ޔ‬㘃ߩ⿠Ḯ࿾ߣ⿠Ḯᤨᦼߦ㑐ߔࠆ⺰੎ࠍ⿠ߎߒߡ޿ࠆ‫ߩࠄࠇߘޕ‬ൻ⍹ߩᒻᘒቇ⊛߅ࠃ
߮♽⛔⊛ߥ⸃ᨆࠍ߅ߎߥߞߚ‫ޕ‬
C) ⃻↢㔤㐳㘃ߩᯏ⢻ᒻᘒቇ⊛⎇ⓥ
C-1) ࠹࠽ࠟࠨ࡞ߩ㖸ჿ↢ℂߦ㑐ߔࠆታ㛎ⴕേቇ⊛⎇ⓥ
⷏᧛೰㧘㚅↰໪⾆㧔⹺⍮ቇ⠌ಽ㊁㧕
࠹࠽ࠟࠩ࡞㘃ߩ༄㗡ㇱߩᲧセ⸃೬ߣᨴ⦡⚵❱ಾ ૞ᚑߦࠃࠅ‫ޔ‬༄㗡ߩ╭㛽ᩰ♽ᒻᘒߩዻ㑆ᄌ⇣ࠍಽᨆߔࠆߣߣ߽
㧙㧙
ߦ‫ޔ‬ᄢဳ㘃ੱ₎ߩ⸃೬⚿ᨐߣᲧセᬌ⸛ߒ࠹࠽ࠟࠩ࡞㘃ߩ․ᓽࠍಽᨆߒߚ‫ࡓ࠙࡝ࡋߩ࡞ࠨࠟ࠽࠹࠹ࡠࠪޔߚ߹ޕ‬㖸ჿ
ታ㛎࠺࡯࠲ߩᚑᨐࠍߣࠅ߹ߣ߼ߚ‫ޕ‬
C-2) ࠴ࡦࡄࡦࠫ࡯ߩ㥦⣧ߩ↢ℂቇ⊛ᯏ⢻ߦ㑐ߔࠆᵹ૕Ꮏቇ⊛ಽᨆ
⷏᧛೰㧘㋈ᧁ᮸ℂ㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕㧘ችㇱ⾆ሶ㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕㧘᧻ᴛື㇢㧔ᕁ⠨⸒
⺆ಽ㊁㧕
㧘෹᳗㓷Ꮖ㧔ᕁ⠨⸒⺆ಽ㊁㧕㧘ᨋ⟤㉿㧔ᕁ⠨⸒⺆ಽ㊁㧕
ࡅ࠻ߩ㥦⣧ߩ↢ℂቇ⊛ᯏ⢻ߩ․㐳ࠍ᣿ࠄ߆ߦߔࠆߚ߼ߦ‫↢ߩ࠽࠻ࠝޔ‬૕࠴ࡦࡄࡦࠫ࡯୘૕ߩਛ㗻㕙ㇱߩ CT ᠟௝
ߣ‫ޔ‬࿖㓙㔚᳇ㅢାၮ␆ᛛⴚ⎇ⓥᚲߦ߅޿ߡࡅ࠻ߩ㥦⣧ㇱߩ MRI ᠟௝ࠍⴕ޿‫ޔ‬ᱜ⏕ߥ㥦⣧ਃᰴరᒻᘒߩ࠺ࠫ࠲࡞࠺
࡯࠲ࠍ૞ᚑߒߚ‫ޔߚߒߦߣ߽ࠍࠇߘޕ‬㥦⣧ౝߩๆ᳇ߩᵹࠇ‫ޔ‬᷷ᐲ࡮Ḩᐲᄌൻߦ㑐ߔࠆᵹ૕Ꮏቇ⊛ࠪࡒࡘ࡟࡯࡚ࠪࡦ
ࠍታᣉߒߚ‫ޕ‬
C-3) ᦛ㥦₎㘃ߩ೽㥦⣧ᒻᘒߩᄌ⇣ߦ㑐ߔࠆ⎇ⓥ
⷏᧛೰
㔤㐳㘃ߦ߅ߌࠆ೽㥦⣧ߩㅴൻࡊࡠ࠮ࠬࠍ᣿ࠄ߆ߦߔࠆߚ߼‫ޔ‬㜞⸃௝ᐲ CT ࠍ↪޿ߡ࿖ౝᄖᯏ㑐ߦᚲ⬿ߐࠇߡ޿ࠆ
ᦛ㥦₎㘃㗡㛽ᮡᧄࠍㅊട᠟௝ߒ‫ߩ⑼ోޔ‬೽㥦⣧ᒻᘒߩᄌ⇣ࠍ⏕⹺ߒߚ‫ޕ‬
C-4) 㔤㐳㘃ߩ྾⢇ߦߟ޿ߡߩᯏ⢻ᒻᘒቇ⊛⎇ⓥ
ᳯᧁ⋥ሶ
micro CT ߦࠃࠆ᠟௝࠺࡯࠲ࠍ↪޿ߡ‫ޔ‬྾⢇㛽ߩౝㇱ᭴ㅧߩ⸃ᨆࠍⴕߞߡ޿ࠆ‫ᧄޕ‬ᐕᐲߪ‫ࠬ࡝ࡠޔ‬㘃߿ࡏࡁࡏߦߟ
޿ߡߩ᠟௝ࠍⴕߞߚ‫ޔߚ߹ޕ‬㔤㐳㘃ߦ߅ߌࠆ྾⢇㛽ᒻᘒ߿ᆫ൓ߩ㆑޿ߣ㛽ߦ߆߆ࠆ⩄㊀ߣߩ㑐ଥࠍജቇ⊛ߦᬌ⸛ߔ
ࠆߚ߼ߦ‫╭ޔ‬㛽ᩰ♽ߩᢙℂࡕ࠺࡞ߩ᭴▽ࠍⴕߞߡ޿ࠆ‫ߥ⊛⥸৻ޕ‬㔤㐳㘃ߣߒߡࠝࡑࠠࠩ࡞ࠍ૶޿‫੹ޔ‬ᐕᐲߪࠦࡦࡇ
ࡘ࡯࠲࡯࡮ࡕ࠺࡞਄ߢᱠⴕߩធ࿾ᦼ㑆ਛߢߩᆫ൓ᄌൻߩㄭૃᓳరࠍⴕߞߚ‫ޕ‬
C-5) ᧲ࠕࠫࠕ↥ࡑࠞࠢߩ㗡㛽ᒻ⁁ߩᲧセ⎇ⓥ
દ⮮Პ㧘⷏᧛೰㧘㜞੗ᱜᚑ
ࡑࠞࠢዻߩ⃻↢⒳ࠍኻ⽎ߦ‫ޔ‬CT ࠍ↪޿ߚ㗡㛽ౝㇱ᭴ㅧߩ⸃ᨆߣᐞ૗ቇ⊛ᒻᘒ᷹ቯࠍ↪޿ߚ㗡㛽߅ࠃ߮ᱤ‎ߩ⸃
ᨆࠍⴕ޿‫ޔ‬ᒻ⁁ᄌ⇣ߩ᳇୥ⅣႺㆡᔕߦߟ޿ߡᬌ⸛ߒߚ‫ޕ‬
C-6) ࡗࠢࠪࡑࠩ࡞㗡⬄ᒻᘒߩ⍵ዊൻߣᒻ⁁ᄌൻߩ⎇ⓥ
⍫㊁⥶
ደਭፉߦ޿ࠆ࠾ࡎࡦࠩ࡞ߪ‫ޔ‬ፉᎩലᨐߦࠃࠅ‫ᧄޔ‬࿯ߦ޿ࠆ߽ߩߦᲧߒߡࠨࠗ࠭߇ዊߐߊ‫ޔ‬㗻㕙ᒻ⁁߇⇣ߥࠆ‫ޕ‬ਔ
⠪ߩᚑ㐳ࠕࡠࡔ࠻࡝࡯゠〔߇⇣ߥߞߡ޿ࠆߎߣࠍ␜ߒ‫ߪߡ޿߅ߦ࡞ࠩࡑࠪࠢࡗޔ‬නߦፉᎩൻߦࠃࠆዊဳൻߦઃ㓐ߒ
ߡ㗻㕙ᒻ⁁߇ᄌൻߒߚߩߢߪߥߊ‫⿠ߚߒ┙⁛ߦࠇߙࠇߘޔ‬࿃߇޽ࠆߎߣࠍ␜ໂߒߚ‫ޕ‬
D) 㔤㐳㘃એᄖߩ߶੃㘃ࠍਥߥኻ⽎ߣߒߚฎ↢‛ቇ⊛⎇ⓥ
D-1) ฎ╙ਃ♿ູ੃㘃⋧ߩ⸃ᨆ
ᳯᧁ⋥ሶ㧘㜞੗ᱜᚑ
ฎ╙ਃ♿㧔6500 ਁᐕ೨㨪2400 ਁᐕ೨㧕ߩ㒽᫮⣄ᬁേ‛⋧ࠍ⸃ᨆߔࠆߎߣߦࠃߞߡ‫੃ູޔ‬㘃ߩㅴൻߩታᘒࠍ᣿ࠄ
߆ߦߔࠆߎߣࠍ⋡ᜰߒߡ޿ࠆ‫ᧄޕ‬ᐕᐲߪ‫ޔ‬Ԙࡒࡖࡦࡑ࡯ߩࡐࡦ࠳࠙ࡦጀ߿࠲ࠗߩࠢ࡜ࡆ⋧‫ࡦ࡝ࠡ࡞ࠛߩ࡞ࠧࡦࡕޔ‬
࠱࡯ጀ߆ࠄ↥಴ߒߚ㘩⡺㘃ൻ⍹ߩ♽⛔ಽ㘃ቇ⊛ᬌ⸛ߣ⸥タ‫ޔ‬ԙࠕࠫࠕ᧲ㇱߩฎ╙ਃ♿⡺㘩ູ੃㘃⋧ߩᲧセ⸃ᨆ‫ޔ‬Ԛ
⡺ᱤ⋡ߩ♽⛔⊛૏⟎ߩᬌ⸛ߩߚ߼ߩᒻᘒ࠺࡯࠲෼㓸ࠍⴕߞߚ‫ޕ‬
D-2) ࡒࡖࡦࡑ࡯ਛㇱߦ߅ߌࠆਛᣂ਎߆ࠄᦝᣂ਎ߩᣂ╙ਃ♿⣄ᬁേ‛ൻ⍹⋧ߩ⸃ᨆ
⷏ጟ૓৻㇢㧘㜞੗ᱜᚑ㧘ᳯᧁ⋥ሶ㧘⷏᧛೰
ࡒࡖࡦࡑ࡯ߩᣂ╙ਃ♿ູ੃㘃↢ጀᐨߩ⸃᣿ࠍ⋡ᜰߒ‫ޔ‬ਛᣂ਎߆ࠄᦝᣂ਎ߦ↢ᕷߒߡ޿ߚູ੃㘃⋧ߩᒻᘒ‫ߣ⛔♽ޔ‬
ㅴൻߦ㑐ߔࠆ⎇ⓥࠍⴕߞߡ޿ࠆ‫ࠢ࠙ࠩࡦࠗࡖ࠴ޕ‬࿾ၞ߿ࠣࠚࡆࡦ࿾ၞߩࠗ࡜ࡢࠫጀ߆ࠄ↥಴ߒߚൻ⍹߿ࠨࡌ࡯࿾ၞ
ߩ╙྾♿ၸⓍ‛߆ࠄ↥಴ߒߚൻ⍹ࠍหቯߒ‫ઁޔ‬࿾ၞ↥಴േ‛ߣߩ♽⛔⊛߅ࠃ߮ฎ↢‛ಽ㘃࿾ℂ⊛㑐ଥࠍᬌ⸛ߒߡ޿
ࠆ‫੃ູޕ‬㘃ߩ㘩⡺㘃ߣߍߞᱤ㘃ߩൻ⍹⒳ߩ⸥タࠍⴕ޿‫ޔ‬ਗⴕߒߡ࠙ࠪ⑼ߩൻ⍹ߩಽ㘃࡮⸥タ૞ᬺࠍㅴ߼ߡ޿ࠆ‫߹ޕ‬
ߚῬ⯻㘃ߩࠞࡔ㘃ߩൻ⍹ߩ੍஻⊛ߥಽ㘃ࠍⴕߞߚ‫↥ޔߚ߹ޕ‬಴ູ੃㘃ߩᱤߩࠛ࠽ࡔ࡞⾰ߦ฽߹ࠇࠆ㉄⚛ߣ὇⚛቟ቯ
ห૏૕ࠍ↪޿ߡฦേ‛⋧ߩฎⅣႺ߿ฎ↢ᘒߩ⸃ᨆࠍㅴ߼ߡ޿ࠆ‫ޕ‬
D-3) ፉᩮ⋵↥ߩ೨ᦼਛᣂ਎ࡆ࡯ࡃ࡯ൻ⍹ߩ⎇ⓥ
⷏ጟ૓৻㇢㧘㜞੗ᱜᚑ
ፉᩮඨፉߩฎᶆጀ㧔⚂ 2000 ਁᐕ೨ߩ࿾ጀ㧕߆ࠄ↥಴ߒߚᄢဳߩൻ⍹ࡆ࡯ࡃ࡯Youngofiber㧔ࡗࡦࠧࡈࠔࠗࡃ࡯ዻ㧕
ߩᱤߩౝㇱࠛ࠽ࡔ࡞ࡄ࠲࡯ࡦࠍ pQCT ࠬࠠࡖࡦࠍ↪޿ߡⷰኤߒߡ⸥タߒߚ‫ߢ߹ࠇߎޕ‬ᣣᧄߩࡆ࡯ࡃ࡯ൻ⍹ߪℰᶉᏒ‫ޔ‬
นఽᏒ‫ޔ‬૒਎଻Ꮢ߆ࠄ⍮ࠄࠇߡ޿ߚ߇‫੹ޔ‬࿁ߩႎ๔ߦࠃߞߡᣂߒ޿↥࿾ࠍㅊടߔࠆߎߣ߇ߢ߈ߚ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
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㧙㧙
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Takai M (2012) Origins and evolution of early primates. (Post-Genome Biology of Primates) (ed. Hirai H, et al.)
p.269-280 Primatology Monographs, Springer.
ߘߩઁߩၫ╩
1)
2)
3)
ᳯᧁ⋥ሶ (2011) ૗ࠍ⷗ߡࠆ߆ࠊ߆ࠄߥ޿. ‫ ╙ ࠅ❱ࠇߠߟߩߜߚ‛߈↢ޟ‬5 Ꮞ‫ ޠ‬੩ㇺᄢቇࠣࡠ࡯ࡃ࡞ COE
ࡊࡠࠣ࡜ࡓ ↢‛ߩᄙ᭽ᕈߣㅴൻ⎇ⓥߩߚ߼ߩ᜚ὐᒻᚑ 㧙ࠥࡁࡓ߆ࠄ↢ᘒ♽߹ߢ㧙 p.41-42 ੩ㇺᄢቇᄢቇ㒮
ℂቇ⎇ⓥ⑼↢‛⑼ቇኾ᡹࡮↢ᘒቇ⎇ⓥ࠮ࡦ࠲࡯࡮㔤㐳㘃ቇ⎇ⓥᚲ࡮㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯㧦੩ㇺ㧚
⷏᧛೰ (2011) ⹤ߒߎߣ߫ߩ㔤㐳㘃⊛⿠Ḯ㨪⪲ጊࠦ࡟࡚ࠢࠪࡦࠍ߽ߣߦ. ᣣᧄੱ㘃ቇળㅴൻੱ㘃ಽ⑼ળ࠾ࡘ࡯
ࠬ࡟࠲࡯ 25: p.6-9㧚
⷏᧛೰㧘ਛോ⌀ੱ㧘ᐔፒ㍈⍫㧘┻ᧄᶈౖ㧘ᷡ᳓ᄢテ㧘࿡᧻⼾㧘⍹↰⧷ኪ㧘ጟ↰቞ᒾ (2011) ᣣᧄੱ㘃ቇળㅴൻ
ੱ㘃ቇಽ⑼ળ╙ 25 ࿁ࠪࡦࡐࠫ࠙ࡓ ⪲ጊ᧖ᄦవ↢ㅊᖬ⸥ᔨࠪࡦࡐࠫ࠙ࡓ‫⎇ߩ࠻ࡅߣ࡞ࠨޟ‬ⓥ߆ࠄ⷗߃ࠆ߽ߩ‫ޠ‬㧚
Anthropological Science (Japanese Series) 119(2): p.137-142㧚
ቇળ⊒⴫
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9)
Egi N, Tsubamoto T, Takai M (2011) Carnivorous mammal faunas in the Paleogene of East Asia: timing of faunal
turnovers and geographical differences. 22nd International Senckenberg Conference “The world at the time of Messel:
puzzles in paleobiology, paleoenvironment and the history of early primates” (2011/11, Frankfurt, Germany)㧚
Nishimura T, Lebrun R, Ponce de León M, Zollikofer CPE (2011) Morphological variation of the paranasal sinuses in
strepsirrhines. The 80th Annual Meeting of the American Association of Physical Anthropologists (2011/4/14-16, Hilton
Minneapolis, Minneapolis, USA).
Nishioka Y, Thaung-Htike, Zin-Maung-Maung-Thein, Egi N, Tsubamoto T, Takai M (2011) Evolutionary history of Old
World porcupines in Eurasia with an emphasis on the new Hystrix materials from Myanmar. World Conference on
Paleontology and Sedimentology (2011/11/28-12/2, Nakhon Rachashima, Thailand).
Sonoda T, Hirayama R, Takai M, Thaung-Htike, Zin-Maung-Maung-Thein, Ando H (2011) The morphological variation
in gigantic tortoises (Testudinidae; Testudines) from the Irrawaddy Sediments in Central Myanmar. World Conference
on Paleontology and Sedimentology. (2011/11/28-12/02, Nakhon Rachashima, Thailand).
Takai M, Thaung-Htike, Zin-Maung-Maung-Thein, Maung-Maung, Aung-Naing-Soe, Egi N, Tsubamoto T (2011) First
discovery of cercopithecid monkeys from the late Miocene to Pliocene Irrawaddy sediments in central Myanmar. World
Conference on Paleontology and Sedimentology (2011/11/28-12/02, Nakhon Rachashima, Thailand).
Tsubamoto T, Thaung-Htike, Zin-Maung-Maung-Thein, Egi N, Takai M (2011) The anthracotheres (Mammalia,
Artiodactyla) from the Neogene of central Myanmar. 71st Annual Meeting Society of Vertebrate Paleontology (2011/11,
Las Vegas, USA).
Yano W, Egi N, Takano T, Ogihara N. (2011) Craniofacial ontogenetic trajectories of two subspecies of Japanese
macaque (Macaca fuscata).From fetus to adult. The 80th Annual Meeting of the American Association of Physical
Anthropologists (2011/04, Minneapolis, USA).
Yano W, Egi N, Takano T, Ogihara N (2011) Ontogenetic trajectories of two subspecies of Macaca fuscata. The 5th
International Symposium of the Global COE project “from Genome to Ecosystem.” (2011/07, Kyoto)㧚
ᳯᧁ⋥ሶ㧘⩆ේ⋥㆏ (2011) ᮸਄ᕈ྾⿷ᱠⴕဳ㔤㐳㘃ߩ೨⢇╭㛽ᩰࡕ࠺࡞ߩ᭴▽. ᣣᧄੱ㘃ቇળᄢળ (2011/11,
㇊ⷓᏒ)㧚
㧙㧙
10) ᳯᧁ⋥ሶ㧘㎄ᧄᱞਭ㧘㜞੗ᱜᚑ (2011) ฎ╙ਃ♿࡙࡯࡜ࠪࠕ᧲ㇱߦ߅ߌࠆ⡺㘩ູ੃േ‛⋧㧦ᤨ㑆⊛ᄌㆫߣ࿾ℂ
⊛ᄌ⇣. ᣣᧄ࿾⃿ᖺᤊ⑼ቇㅪว 2011 ᐕᄢળ (2011/05, ජ⪲)㧚
11) ᳯᧁ⋥ሶ㧘㎄ᧄᱞਭ㧘㜞੗ᱜᚑ (2011) ฎ╙ਃ♿᧲࡙࡯࡜ࠪࠕߩ⡺㘩ູ੃േ‛⋧㧦᭴ᚑⷐ⚛ߩᤨ㑆⊛ᄌൻߣ࿾
ℂ⊛ᄌ⇣. ᣣᧄฎ↢‛ቇળᐕળ (2011/07, ㊄ᴛᏒ)㧚
12) ᳯᧁ⋥ሶ㧘㎄ᧄᱞਭ㧘㜞੗ᱜᚑ㧘ࠫࡦ࡮ࡑ࠙ࡦ࡮ࡑ࠙ࡦ࡮࠹ࠗࡦ㧘࠲࠙ࡦ࡮࠲ࠗ (2011) ೋᦼ㔤㐳㘃ࠍ↥ߔࠆ
ࡐࡦ࠳࠙ࡦູ੃േ‛⋧㧔ਛᦼᆎᣂ਎㧧ࡒࡖࡦࡑ࡯㧕ߩฎ↢‛࿾ℂ⊛․ᓽ. ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07, ›ጊ)㧚
13) દ⮮Პ㧘⷏᧛೰㧘㜞੗ᱜᚑ (2011) ࠕࠫࠕߩ㔤㐳㘃ࡑࠞࠢ㘃ߦ߅ߌࠆ㗡㛽㗻㕙ㇱߩࠕࡠࡔ࠻࡝࡯ߣ⒳㑆Ꮕ㆑. ᣣ
ᧄ࿾⃿ᖺᤊ⑼ቇㅪว 2011 ᐕᄢળ, ᐀ᒛࡔ࠶࠮࿖㓙ળ⼏႐ (2011/05/24, ජ⪲)㧚
14) ᫪↰⥶㧘⍫㊁⥶㧘ਛോ⌀ੱ (2011) ਄㗶╙৻ᄢ⥓ᱤᱤ㗖✢ࠍ↪޿ߚᐞ૗ቇ⊛ᒻᘒ⸃ᨆ. ╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢ
ળ (2011/11, ᴒ✽)㧚
15) ⷏᧛೰ (2011) ⹤ߒߎߣ߫ߩ㔤㐳㘃⊛⿠Ḯ㨪⪲ጊࠦ࡟࡚ࠢࠪࡦࠍ߽ߣߦ. ᣣᧄੱ㘃ቇળㅴൻੱ㘃ቇಽ⑼ળ╙
25 ࿁ࠪࡦࡐࠫ࠙ࡓ ⪲ጊ᧖ᄦవ↢ㅊᖬ⸥ᔨࠪࡦࡐࠫ࠙ࡓ‫⎇ߩ࠻ࡅߣ࡞ࠨޟ‬ⓥ߆ࠄ⷗߃ࠆ߽ߩ‫( ޠ‬2011/05/14, ੩
ㇺ)㧚
16) ⷏᧛೰㧘᫪ᄥᔒ㧘ᾢ⇌ᷡᔒ㧘᧻Ỉᾖ↵ (2011) ࡑࠞࠢࠩ࡞ߦ߅ߌࠆ਄㗶ᵢߩ↢ℂቇ⊛ᯏ⢻ߦ㑐ߔࠆᢙ୯ᵹ૕ജ
ቇ⊛⎇ⓥ. ╙ 26 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/18, ›ጊ)㧚
17) ⷏᧛೰㧘㜞੗ᱜᚑ (2011) ࡙࡯࡜ࠪࠕߦ߅ߌࠆᄢဳࠝ࠽ࠟࠩ࡞੝⑼ൻ⍹ߩㅴൻผ㧦㗡㛽ౝㇱ᭴ㅧߩᒻ⾰ߣ᜛ᢔ
⚻〝. ᣣᧄ࿾⃿ᖺᤊ⑼ቇㅪว 2011 ᐕᄢળ, ᐀ᒛࡔ࠶࠮࿖㓙ળ⼏႐ (2011/05/24, ජ⪲)㧚
18) ⷏ጟ૓৻㇢ (2011) ࡒࡖࡦࡑ࡯ਛᄩㇱߩᣂ╙ਃ♿ᓟඨߩⅣႺᄌേߦ઻ߞߚࡗࡑࠕ࡜ࠪߩㅴൻ. ᣣᧄ࿾⃿ᖺᤊ⑼
ቇㅪวᄢળ (2011/05/23-28, ᐀ᒛ)㧚
19) 㜞੗ᱜᚑ (2011) ᧲ㇱࠕࠫࠕߦ߅ߌࠆᣂ╙ਃ♿ᓟඨߩ⁜㥦₎㘃ߩㅴൻผߣߘߩ᜛ᢔ⚻. ᣣᧄ࿾⃿ᖺᤊ⑼ቇㅪว
ᄢળ (2011/05/23-28, ᐀ᒛ)㧚
20) ㎄ᧄᱞਭ㧘ᳯᧁ⋥ሶ㧘㜞੗ᱜᚑ㧘࠲࠙ࡦ࡮࠲ࠗ㧘ࠫࡦ࡮ࡑ࠙ࡦ࡮ࡑ࠙ࡦ࡮࠹ࠗࡦ (2011) ⃻↢㔤㐳㘃ߩ〒㛽ࠨ
ࠗ࠭ߩ⸘᷹ߣࡐࡦ࠳࠙ࡦߩൻ⍹㔤㐳㘃ߩ૕㊀ផቯ߳ߩᔕ↪. ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07, ›ጊ)㧚
21) ㎄ᧄᱞਭ㧘Thaung-Htike㧘Zin-Maung-Maung-Thein㧘ᳯᧁ⋥ሶ㧘㜞੗ᱜᚑ (2011) ࡒࡖࡦࡑ࡯ߦ߅ߌࠆᣂ╙ਃ♿
ߩࠕࡦ࠻࡜ࠦ࠹࡝࠙ࡓ㘃㧔ູ੃✁㧘஧か⋡㧕ߩㅴൻ. ᣣᧄ࿾⃿ᖺᤊ⑼ቇㅪว 2011 ᐕᄢળ (2011/05, ජ⪲)㧚
22) ㎄ᧄᱞਭ㧘࠲࠙ࡦ࡮࠲ࠗ㧘ࠫࡦ࡮ࡑ࠙ࡦ࡮ࡑ࠙ࡦ࡮࠹ࠗࡦ㧘ᳯᧁ⋥ሶ㧘⷏ጟ૓৻㇢㧘ࡑ࠙ࡦ࡮ࡑ࠙ࡦ㧘㜞੗
ᱜᚑ (2011) ࡒࡖࡦࡑ࡯ਛᄩㇱߩᣂ╙ਃ♿ߦ߅ߌࠆࠕࡦ࠻࡜ࠦ࠹࡝࠙ࡓ㘃㧔ູ੃✁஧か⋡㧕ߩㅴൻ. ᣣᧄฎ↢
‛ቇળᐕળ (2011/07, ㊄ᴛᏒ)㧚
23) ⍫㊁⥶㧘⩆ේ⋥㆏ (2011) ፉᎩൻߦ઻߁ࡗࠢࠪࡑࠩ࡞㧔Macaca fuscata yakui㧕㗡⬄㛽ߩዊဳൻߣᒻ⁁ߩ․ᱶൻ.
╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢળ (2011/11, ᴒ✽)㧚
24) ⍫㊁⥶㧘㜞㊁ᥓ㧘ᳯᧁ⋥ሶ㧘⩆ේ⋥㆏ (2011) ಴↢೨ᓟߦ߅ߌࠆ㗡⬄㛽ᚑ㐳᭽ᑼߩᄌൻ. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇ
ળ (2011/07, ›ጊ)㧚
25) ⷏ጟ૓৻㇢㧘Rössner GE㧘㜞੗ᱜᚑ (2012) ࡒࡖࡦࡑ࡯ਛㇱߩ਄ㇱਛᣂ⛔㧙ਅㇱᦝᣂ⛔߆ࠄ↥಴ߒߚ࠙ࠪ⑼ൻ
⍹ߩ੍ኤ⊛ᬌ⸛. ᣣᧄฎ↢‛ቇળ╙ 161 ࿁଀ળ (2012/01/20-22, ንጟ)㧚
26) ⭽↰ືᐔ㧘ᐔጊᑇ㧘㜞੗ᱜᚑ㧘࠲࠙ࡦ࡮࠲ࠗ㧘ࠫࡦ࡮ࡑ࠙ࡦ࡮ࡑ࠙ࡦ࡮࠹ࠗࡦ㧘቟⮮ኼ↵ (2012) ࡒࡖࡦࡑ࡯
ਛᄩㇱࠗ࡜ࡢࠫၸⓍ‛ࠃࠅ↥಴ߒߚᏂᄢ࡝ࠢࠟࡔ㘃ߩᄖ⣻↲ߦ⷗ࠄࠇࠆᒻᘒ⊛ᄌ⇣. ᣣᧄฎ↢‛ቇળ╙ 161 ࿁
଀ળ (2012/01/20-22, ንጟ)㧚
27) 㜞੗ᱜᚑ㧘⷏ጟ૓৻㇢㧘࠲࠙ࡦ࡮࠲ࠗ㧘ࠫࡦ࡮ࡑ࠙ࡦ࡮ࡑ࠙ࡦ࡮࠹ࠗࡦ㧘ࡑ࠙ࡦ࡮ࡑ࠙ࡦ (2012) ࡒࡖࡦࡑ࡯
ਛᄩㇱߩᓟᦼ㞲ᣂ਎ዊဳູ੃㘃ൻ⍹฽᦭ጀߦ㑐ߔࠆ੍ኤ⊛ႎ๔. ᣣᧄฎ↢‛ቇળ╙ 161 ࿁଀ળ (2012/01/20-22,
ንጟ)㧚
28) ㎄ᧄᱞਭ㧘Tsogtbaatar Kh㧘ታศ₵⾆㧘Mainbayar B㧘ᷰㇱ⌀ੱ㧘Chinzorig Ts㧘㋈ᧁ⨃㧘Khatanbaatar P㧘ᳯᧁ⋥
ሶ (2012) ࡕࡦࠧ࡞ߩ਄ㇱᆎᣂ⛔ Ergilin Dzo ጀ↥ߩູ੃㘃ൻ⍹⟲㧦ᦨᣂߩ⎇ⓥᚑᨐ. ᣣᧄฎ↢‛ቇળ╙ 161 ࿁
଀ળ (2012/01, ንጟ)㧚
⻠Ṷ
1)
2)
3)
⷏᧛೰ (2011/08/28) ๭ๆ‫ྚޔ‬ਅ‫ߩ⹤⊒ߡߒߘޔ‬ㅴൻ. ╙ 31 ࿁ᱤߩᒻᘒቇࠍ߼ߋࠆᙣ⹤ળ, ฬฎደ࡙ࡑ࠾࠹ࠢ
ᱤ⑼⵾⩻ኾ㐷ቇᩞ㧚
⷏᧛೰ (2011/10/15) ࠨ࡞ߩჿߣࡅ࠻ߩ⹤ߒߎߣ߫. ੩ㇺᄢቇ✚วඳ‛㙚࡟ࠢ࠴ࡖ࡯ࠪ࡝࡯࠭ No.92, ੩ㇺᄢቇ
✚วඳ‛㙚㧚
⷏᧛೰ (2011/12/05) ๭ๆ‫ྚޔ‬ਅ‫ߩ⹤⊒ߡߒߘޔ‬ㅴൻ. ╙ 7 ࿁㕍ⓨႶ, ฬ㋕›ጊࡎ࠹࡞㧚
ߘߩઁ
1)
㜞੗ᱜᚑ (2011) 2011/07/16 ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળ ⥄↱㓸ળ‫ߩ࡞ࠩࡦࡎ࠾ޟ‬ൻ⍹‫ޠ‬㐿௅⽿છ⠪㧔㔤㐳㘃⎇ⓥ
Vol.27 No.2 P.161㧕
㧚
㧙㧙
␠ળ↢ᘒ⎇ⓥㇱ㐷
↢ᘒ଻ోಽ㊁
㧨⎇ⓥ᭎ⷐ㧪
A) ࠾ࡎࡦࠩ࡞ߩ↢ᘒቇ࡮ⴕേቇ
ᷰㆻ㇌ᄦ㧘ඨ⼱๋㇢㧘ᒛ㢈㧘ㇹ߽߃㧘Ỉ↰᥏ሶ㧘ᄢ⼱ᵗ੺
ੱὑ⊛ᓇ㗀ߩዋߥ޿ⅣႺߦߔ߻‫ޔ‬㊁↢ߩ࠾ࡎࡦࠩ࡞߇‫⥄ޔ‬ὼⅣႺ߆ࠄฃߌࠆᓇ㗀ߦ⌕⋡ߒߥ߇ࠄ‫ޔ‬୘૕⟲↢ᘒቇ‫ޔ‬
ណ㘩↢ᘒቇ‫ⴕޔ‬േ↢ᘒቇߥߤߩⷰὐ߆ࠄ‫⎇ޔ‬ⓥࠍㅴ߼ߡ޿ࠆ‫ޕ‬
ደਭፉߩἑಾᎹ਄ᵹၞߢߪ‫ޔ‬᫪ᨋબណߣᨐታߩ⼾ರߩᐕᄌേ߇࠾ࡎࡦࠩ࡞୘૕⟲ߦਈ߃ࠆᓇ㗀ࠍ᣿ࠄ߆ߦߔࠆ⋡
⊛ߢ‫ޔ‬
‫⺞࡞ࠩࠢࡗޟ‬ᩏ㓌‫߁޿ߣޠ‬ቇ↢ߥߤߩࡏ࡜ࡦ࠹ࠖࠕ߆ࠄߥࠆ⺞ᩏࠣ࡞࡯ࡊࠍ⚵❱ߒ‫ޔ‬1998 ᐕએ᧪⺞ᩏࠍ⛮⛯
ߒߡ޿ࠆ‫੹ޕ‬ᐕ߽ᄐቄߦ৻ᢧ⺞ᩏࠍⴕߞߡ‫ੱޔ‬ญቇ⊛⾗ᢱࠍ㓸߼ߚ‫⾗ߩߎޕ‬ᢱࠍၮߦ‫ߩ࡞ࠩ࡝࠻ࡅޔ‬ኒᐲߣ‫ߩߘޔ‬
࿾ၞᄌ⇣࡮࿾ℂᄌ⇣ߦߟ޿ߡಽᨆߒߚ‫ߩ࡞ࡅޔߣࠞࠪ࡮࡞ࠨޔߚ߹ޕ‬ಽᏓߣߩ㑐ଥߦߟ޿ߡಽᨆߒߚ‫ޕ‬
ደਭፉᶏጯㇱߢߪ‫ޔ‬㊁↢࠾ࡎࡦࠩ࡞ߩࠠࡁࠦ㘩ߩᄙ᭽ᕈߣⴕേࡄ࠲࡯ࡦߦߟ޿ߡ⎇ⓥࠍⴕߞߚ‫ߦࠇ⟲ޔߚ߹ޕ‬ᚲ
ዻߔࠆࠝࠬ߇৻ᤨ⊛ߦ⟲ࠇࠍ㔌ࠇࠆⴕേߦߟ޿ߡ⎇ⓥߒߚ‫ޕ‬
ዊ⼺ፉߩ࠾ࡎࡦࠩ࡞ߩ␠ળ੤ᷤߦߟ޿ߡߩ⎇ⓥࠍⴕߞߚ‫ޕ‬
B) ࠾ࡎࡦࠩ࡞ߣหᚲ⊛ߦ↢ᕷߔࠆ↢‛ߣߩ㑐ଥ
ඨ⼱๋㇢㧘Ỉ↰᥏ሶ㧘ỳ↰㘧㠽
ደਭፉߢ‫ߣ࡞ࠩࡦࡎ࠾ޔ‬หᚲ⊛ߦ↢ᕷߔࠆ↢‛ߣߩ㑐ଥߦߟ޿ߡ⎇ⓥࠍⴕߞߚ‫ߩࠢࠫ࠴ࠗޕ‬ખ㑆ߢ޽ࠆࠕࠦ࠙ߩ
ᨐታࠍណ㘩ߔࠆ‫ߣ࡞ࠩࡦࡎ࠾ޔ‬㠽ߥߤߩᨐታ㘩⠪ߦߟ޿ߡ⺞ᩏߒߚ‫ࠦࡁࠠߩ࡞ࠩࡦࡎ࠾ޔߚ߹ޕ‬㘩ߦࠃࠆ⩶ߩ⢩ሶ
ᢔᏓߦߟ޿ߡ⎇ⓥࠍⴕߞߚ‫ޕ‬
C) ࠾ࡎࡦࠩ࡞ߩᶖൻ⢻ജߩ⎇ⓥ
Ỉ↰᥏ሶ㧘ඨ⼱๋㇢
㘺⢒୘૕ࠍኻ⽎ߦ‫ޔ‬ᶖൻ₸ߣ㘩‛ߩᶖൻ▤ㅢㆊᤨ㑆ߦߟ޿ߡߩታ㛎⊛⎇ⓥࠍⴕߞߚ‫ޕ‬
D) ࠾ࡎࡦࠩ࡞ߩ୘૕⟲▤ℂ
ᷰㆻ㇌ᄦ
ᄙ᭽ߥⷰὐ߆ࠄ࠾ࡎࡦࠩ࡞ߦࠃࠆㄘ૞‛ⵍኂߩ໧㗴⸃᳿ࠍ࿑ࠆߚ߼‫ޔ‬ㄘ૞‛ⵍኂࠍ⿠ߎߒߡ޿ࠆ࠾ࡎࡦࠩ࡞ߩ㘩
ᕈ߿࿯࿾೑↪ߦᓇ㗀ࠍਈ߃ࠆⷐ࿃ߩಽᨆ‫᦭ޔ‬ലߥⵍኂ▤ℂᚻᴺߩ㐿⊒‫₎ޔ‬ኂߦߟ޿ߡߩ␠ળቇ⊛⎇ⓥߥߤࠍⴕߞߚ‫ޕ‬
E) 㘺⢒࠾ࡎࡦࠩ࡞ߩㆆേߦߟ޿ߡߩ⎇ⓥ
ㇹ߽߃㧘C Sueur
㔤㐳㘃⎇ⓥᚲߩደᄖ᡼㘺႐ߦ㘺⢒ߐࠇߡ޿ࠆ࠾ࡎࡦࠩ࡞ࠍኻ⽎ߦ‫ޔ‬ㆆേߩࡔࠞ࠾࠭ࡓߦߟ޿ߡߩ⎇ⓥࠍⴕߞߚ‫ޕ‬
F) ㊁↢࠴ࡦࡄࡦࠫ࡯ߣࡏࡁࡏߩ⎇ⓥ
ᯅᧄජ⛗㧘દᏀᴦ⟤ᄹ
࠙ࠟࡦ࠳౒๺࿖ࠞ࡝ࡦ࠭᫪ᨋ‫ࠧࡦࠦޔ‬᳃ਥ౒๺࿖ࡢࡦࡃ࿾඙ߢ‫␠ߩࡏࡁࡏߣ࡯ࠫࡦࡄࡦ࠴ࠇߙࠇߘޔ‬ળቇ⊛࡮↢
ᘒቇ⊛⎇ⓥࠍⴕߞߚ‫ߩ࡯ࠫࡦࡄࡦ࠴ޕ‬ㆆേ߿ⴕേߩ࠺࡯࠲ࠍߣࠆߣߣ߽ߦ‫ޔ‬ቯ㊂⊛ߥᬀ↢⺞ᩏ߿ᨐታ㊂⺞ᩏࠍᐔⴕ
ߒߡⴕ޿‫ⴕߩ࡯ࠫࡦࡄࡦ࠴ޔ‬േ߿␠ળ㑐ଥ߇ⅣႺ߆ࠄߤߩࠃ߁ߥᓇ㗀ࠍฃߌߡ޿ࠆ߆ߣ޿߁ὐߦᵈ⋡ߒߡ‫⎇ޔ‬ⓥࠍ
ⴕߞߡ޿ࠆ‫ޕ‬
G) ᧲ධࠕࠫࠕᾲᏪᨋߩ㔤㐳㘃ߩ␠ળ↢ᘒቇ⊛⎇ⓥ
ᷰㆻ㇌ᄦ㧘ඨ⼱๋㇢㧘᧻↰৻Ꮧ㧘ᄢ⼱ᵗ੺
ࠗࡦ࠼ࡀࠪࠕ࡮ਛㇱࠬ࡜࠙ࠚࠪߦ߅޿ߡ‫࡯ࠠࡦࡕࠢ࠶ࡋߣࠢࠞࡑࡦࠕࠤࡦ࠻ޔ‬㑆ߩ⒳㑆㔀⒳ߩ❥ᱺߦߟ޿ߡߩ⛮
⛯ⷰኤࠍ߅ߎߥߞߡ޿ࠆ‫ࠕࠪ࡯࡟ࡑޕ‬㗔ࡏ࡞ࡀࠝፉ㨯ࠨࡃᎺߩ࠳࠽ࡓࡃ࡟࡯᫪ᨋ଻⼔඙ߢߪ‫ޔ‬ᤤⴕᕈ㔤㐳㘃 5 ⒳ߩ
౒ሽߩ↢ᘒቇ⊛ࡔࠞ࠾࠭ࡓࠍ᣿ࠄ߆ߦߔࠆߚ߼‫ޔ‬ኒᐲ࠮ࡦࠨࠬ‫ⴕޔ‬േⷰኤߦࠃࠆ㘩ᕈ߿ㆆേߩ⺞ᩏࠍⴕߞߚ‫੹ޕ‬ᐕ
ᐲߪ‫ߩߘޔߒ⋡⌕ߦ࡯ࠠࡦࡕࡈ࡯࡝࠼࠶࡟ޔ‬㘩ᕈߦߟ޿ߡߩಽᨆࠍⴕߞߚ‫ࡃࠨࠕࠪ࡯࡟ࡑޕ‬Ꮊߩࠬࠞ࠙ߢ‫ⴕޔ‬േⷰ
ኤߣ࠮ࡦࠨࠬࠍ߽ߣߦ‫↢ߩ࡞ࠩࠣࡦ࠹ޔ‬ᘒ߿␠ળ᭴ㅧߦߟ޿ߡߩ⎇ⓥࠍⴕߞߚ‫␠ߩ࡞ࠩࠝ࠲ࡉޔߪߢ࠙ࠞࠬޕ‬ળ↢
ᘒߦߟ޿ߡߩ੍஻⊛⺞ᩏࠍⴕߞߚ‫ޕ‬
H) ࠞ࡝ࡦ࠭᫪ᨋ଻⼔඙ߦ᫮ᕷߔࠆ㊁↢㔤㐳㘃ߩነ↢⯻ߩ⎇ⓥ
ᳯፉବ㧘ᯅᧄජ⛗㧘ฎᏒ೰ผ㧔␠ળㅴൻಽ㊁㧕㧘ጟᧄቬ⵨㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕
㊁↢㔤㐳㘃߇หᚲ⊛ߦ᫮ᕷߔࠆ࠙ࠟࡦ࠳౒๺࿖ࠞ࡝ࡦ࠭᫪ᨋ଻⼔඙ߢ‫ޔ‬ኋਥߣነ↢⯻ߩ㑐ଥࠍℂ⸃ߔߴߊነ↢⯻
ቇ⊛⺞ᩏࠍⴕߞߚ‫ޕ‬6 ⒳ߩᤤⴕᕈ㊁↢㔤㐳㘃ࠍኻ⽎ߣߒ‫ࠍ࡞ࡊࡦࠨ♮ޔ‬ណ㓸ߒߚ‫⃻ߪ࡞ࡊࡦࠨޕ‬࿾ߢ⸃ᨆࠍⴕ޿‫ޔ‬
⯻ෆⷰኤߣ⯻ෆᢙ᷹ቯࠍታᣉߒߚ‫ ߚ߹ޕ‬DNA ⸃ᨆ↪ߣߒߡ 70%ࠛ࠲ࡁ࡯࡞ߦࠨࡦࡊ࡞ࠍ଻ሽߒᣣᧄ߳ᜬߜᏫߞߚ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
Bernard H, Matsuda I, Hanya G, Ahmad AH (2011) Characteristics of Night Sleeping Trees of Proboscis Monkeys
㧙㧙
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3)
4)
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✚⺑
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ႎ๔
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3)
(Nsalis larvatus) in Sabah, Malaysia. International Journal of Primatology 32:259-267.
Bernard H, Matsuda I, Hanya G, Ahmad AH (2011) Effects of river width on the selection of sleeping-site by proboscis
monkeys (Nasalis larvatus) in Sabah, Malaysia. Journal of Tropical Biology and Conservation 8:9-12.
Dufour V, Sueur C, Whiten A, Buchanan-Smith H (2011) The impact of moving to a novel environment on social
networks, activity and wellbeing in two new world primates. American Journal of Primatology 73:802-811.
Hanya G, Stevenson P, van Noordwijk M, Wong ST, Kanamori T, Kuze N, Aiba S, Chapman CA, van Schaik C (2011)
Seasonality in fruit availability affects frugivorous primate biomass and species richness. Ecography 34:1009-1017.
Hanya G, Aiba S (2011) Annual periodicity in fruiting in temperate forests in Yakushima, Japan. Forestry Studies in
China 13:112-122.
Hanya G, Menard N, Qarro M, Ibn Tattou M, Fuse M, Vallet D, Yamada A, Go M, Takafumi H, Tsujino R, Agetsuma N,
Wada K (2011) Dietary adaptations of temperate primates: comparisons of Japanese and Barbary macaques. Primates
52:187-198.
Matsuda I, Murai T, Clauss M, Yamada T, Tuuga A, Bernard H, Higashi S (2011) Regurgitation and remastication in the
foregut-fermenting proboscis monkey (Nasalis larvatus). Biology Letters 7:786-789.
Matsuda I, Tuuga A, Bernard H (2011) Riverine refuging by proboscis monkeys (Nasalis larvatus) and sympatric
primates: implications for adaptive benefits of the riverine habitat. Mammalian Biology 76:165-171.
Pyritz L, King AJ, Sueur C, Fitchel C (2011) Reaching a consensus: terminology used in coordination and
decision-making research. International Journal of Primatology 32:1268-1278.
Sueur C, Petit O, Marco AD, Jacobs A, Watanabe K, Thierry B (2011) A comparative network analysis of social style in
macaques. Animal Behaviour 82:845-852.
Sueur C, King AJ, Conradt L, Kerth G, Lusseau D, Mettke-Hoffmann C, Schaffner CM, Williams L, Zinner D, Aureli F
(2011) Collective decision-making and fission-fusion dynamics: a conceptual framework. Oikos 120:1608-1617.
Sueur C, Jacobs A, Petit O, Amblard F, King AJ (2011) How can social network analysis improve the study of primate
behaviour? American Journal of Primatology 73:703-709.
Zhang P, Li BG, Watanabe K, Qi XG (2011) Sleeping cluster patterns and retiring behaviors during winter in a
free-ranging band of the Sichuan snub-nosed monkey. Primates 52:221-228.
Matsuda I, Tuuga A, Bernard H, Furuichi T (2012) Inter-individual relationships in proboscis monkeys: a preliminary
comparison with other non-human primates. Primates 53:13-23.
Yasuoka H, Kimura D, Hashimoto C, Furuichi T (2012) Quantitative Assessment of Livelihoods around Great Apes
Reserves: Cases in LuoScienctific Reserve, DR Congo, and Kalinzu Forest Reserve, Uganda. African Study Monographs
43:137-157.
᧻↰৻Ꮧ (2011) ࠹ࡦࠣࠩ࡞߆ࠄ⚌⸃ߊࠦࡠࡉࠬ੝⑼ߩᄙ᭽ߥ↢ᘒߣ␠ળ. 㔤㐳㘃⎇ⓥ 27:75-93.
ㄞᄢ๺㧘๺↰৻㓶㧘ᷰㆻ㇌ᄦ (2011) ㊁↢࠾ࡎࡦࠩ࡞ߩណ㘩ߔࠆᧁᧄᬀ‛. 㔤㐳㘃⎇ⓥ 27:27-49.
᧻↰৻Ꮧ (2011) 㔤㐳㘃ߩ㊀ጀ␠ળ㧔ડ↹ࠪࡦࡐࠫ࠙ࡓߩ⊒⴫ౝኈႎ๔㧕. 㔤㐳㘃⎇ⓥ 26:258-259.
ᷰㆻ㇌ᄦ (2011) 㔤㐳㘃ߩ೑߈ᚻ. Clinical Neuroscince 29(6):634-635.
ᷰㆻ㇌ᄦ㧘Ᏹ↰㇌ᒾ㧘ᳯᚑᐢ᢯ (2011) ࠾ࡎࡦࠩ࡞୘૕⟲▤ℂߩ⃻႐ߣ੹ᓟߩ⺖㗴. ູ੃㘃⑼ቇ㧔2010 ᐕᐲ⥄
↱㓸ળ⸥㍳㧕51:212-214.
⪺ᦠ㧔න⪺㧕
1)
2)
Sha J, Matsuda I, Bernard H (2011) The Natural History of Proboscis Monkey. Natural History Publications. p.126 Kota
Kinabalu.
᧻↰৻Ꮧ (2012) ࠹ࡦࠣࠩ࡞㧙ᴡߣ↢߈ࠆࠨ࡞. p.146 ᧲ᶏᄢቇ಴ ળ㧚
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
2)
Sueur C (2012) Social network, information flow and decision making efficiency: comparing humans to animals. (Social
Networking and Community Behavior Modeling: Qualitative and Quantitative Measures.) (ed. Safar, Maytham, and
Khaled Mahdi) p.164-177 IGI Global.
ᯅᧄජ⛗ (2012) ࠞ࡝ࡦ࠭ߩ᫪ߣ࠴ࡦࡄࡦࠫ࡯ߩ଻⼔㧙ࠛࠦ࠷࡯࡝࠭ࡓߩ⃻⁁‫ࠍ࠳ࡦࠟ࠙ޟ‬⍮ࠆߚ߼ߩ 53 ┨‫ޠ‬
(ศ↰᣽ᄦ㧘⊕⍹ჽ৻㇢✬) p.322-327 ᣿⍹ᦠᐫ㧚
⪺ᦠ㧔⠡⸶㧕
1)
ඨ⼱๋㇢ ⸶ (2011) ╙ 5 ┨ 㔤㐳㘃ߩᄙ᭽ᕈߣ↢ᘒ. ╙ 6 ┨ 㔤㐳㘃ߩ㈩஧ࠪࠬ࠹ࡓ. (ࡠࡃ࡯࠻㨯ࡏࠗ࠼㧘࡚ࠫ
࡯ࡦ㨯B㨯ࠪ࡞ࠢ⪺㧘᧻ᧄ᥏ሶ㧘ዊ↰੫⋙⸶⪺,‫ߦ߁ࠃߩߤߪ࠻ࡅޟ‬ㅴൻߒߡ߈ߚ߆‫ )ޠ‬p.189-232,233-274 ࡒࡀ࡞
ࡧࠔᦠᚱ㧘੩ㇺ㧔ಽᜂ⸶㧕.
ߘߩઁߩၫ╩
1)
2)
ᯅᧄජ⛗ (2011) ࠪ࠾ࠕߩሶ⢒ߡᡰេ㨪㔤㐳㘃ߩሶ⢒ߡߣࡅ࠻ߩㅴൻߩⷞὐ߆ࠄ p.7-9 ߎߤ߽ᧂ᧪ 3 ภ㧚
ᯅᧄජ⛗ (2011) ࠴ࡦࡄࡦࠫ࡯ߣ᫪ࠍᱠߊ㧚↢߈‛ߚߜߩߟߠࠇ❱ࠅ╙ 5 Ꮞ p.119-120㧚
㧙㧙
ቇળ⊒⴫
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4)
5)
6)
7)
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13)
14)
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⻠Ṷ
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2)
Go M, Hashimoto C (2011) Vocal synchronization in mixed-species associations of blue monkeys and red-tailed
monkeys (Cercopithecus mitis and C. ascanius) in the Kalinzu Forest, Uganda. Biodiversity and Evolution gCOE
International Symposium. (2011/07/09-10, Kyoto, Japan)㧚
Sawada A (2012) Fungivory in wild monkeys on Yakushima Island. ᣣᧄ↢ᘒቇળ╙ 59 ࿁ో࿖ᄢળ (2012/03/18, ᄢ
ᵤ)㧚
ᒛ㢈㧘ᷰㆻ㇌ᄦ (2011) A comparative study on the dominance style in threeprovisioned free-ranging groups of
Japanese macaque. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/16-18, ›ጊ)㧚
ㇹ߽߃㧘ᯅᧄජ⛗ (2011) ࠨ࡞ߩᷙ⟲ᒻᚑ㧙⇣ߥࠆ⒳ߤ߁ߒ߇ߤߩࠃ߁ߦ৻✜ߦ޿ࠆߩ߆㧫 ᣣᧄູ੃㘃ቇળ
2011 ᐕᐲᄢળ (2011/09/08-11, ችፒ)㧚
ㇹ߽߃㧘ᯅᧄජ⛗ (2011) ࠞ࡝ࡦ࠭᫪ᨋߦ↢ᕷߔࠆࡉ࡞࡯ࡕࡦࠠ࡯ߣ࡟࠶࠼࠹ࠗ࡞ࡕࡦࠠ࡯ߩᷙ⟲ߦ߅ߌࠆ⊒
ჿߩหᦼᕈ. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળ (2011/07/16-18, ›ጊ)㧚
ඨ⼱๋㇢㧘Stevenson P㧘Noordwijk M㧘Wong TS㧘㊄᫪ᦺሶ㧘ਭ਎Ớሶ㧘⋧႐ᘕ৻㇢㧘Chapman CA㧘Schaik C (2011)
ᨐታ↢↥ߩቄ▵ᕈ߇ᨐታ㘩㔤㐳㘃ߩࡃࠗࠝࡑࠬߣ⒳ᢙߦᓇ㗀ߔࠆ. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/17,
›ጊ)㧚
ᯅᧄජ⛗㧘቟ጟብ๺㧘ᚻႦ⾫⥋㧘ฎᏒ೰ผ (2011) ࠙ࠟࡦ࠳౒๺࿖ߦ߅ߌࠆ᫪ᨋ଻⼔඙๟ㄝߩ࿾ၞ૑᳃ߦࠃࠆ
᫪ᨋ⾗Ḯߩ೑↪ߩታᘒ. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/18, ᗲ⍮⋵›ጊᏒ)㧚
᧻↰৻Ꮧ㧘᧛੗഼⵨㧘Clauss M㧘ጊ↰᦮⟤㧘Augustine T㧘Henry B㧘᧲ᱜ೰ (2011) 㔤㐳㘃ߩ෻⧖ⴕേߩ⊒⷗㧦
࠹ࡦࠣࠩ࡞ߩ੐଀. 2011 ᐕᐲᣣᧄູ੃㘃ቇળᄢળ (2011/09, ችፒ)㧚
᧻↰৻Ꮧ㧘Tuuga A㧘Bernard H㧘ฎᏒ೰ผ (2011) ࠹ࡦࠣࠩ࡞␠ળߩ․ᓽ㧦ࡔࠬߩ⒖☋᭽ᑼߣ୘૕㑆㑐ଥ. ╙
27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07, ›ጊ)㧚
ਃ⼱㓷⚐㧘ᷰㆻ㇌ᄦ (2011) ࠪ࡞ࡃ࡯࡞࠻ࡦߩ୘૕ᢙኒᐲߪߤߩࠃ߁ߦ᳿߹ࠆߩ߆㧫 ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળ
ᄢળ (2011/07/16-18, ›ጊ)㧚
ᄢ⼱ᵗ੺㧘Ỉ↰᥏ሶ㧘⑔᳗ᕶ໪㧘ඨ⼱๋㇢ (2011) ࠾ࡎࡦࠩ࡞㓶ߩ㓸࿅߆ࠄߩ৻ᤨቅ┙ⴕേ㧦ណ㘩ᚢ⇛ߩⷰὐ
߆ࠄ. ᣣᧄ㔤㐳㘃ቇળ ╙ 26 ࿁ቇⴚᄢળ (2011/07, ᗲ⍮⋵)㧚
ᄢ⼱ᵗ੺㧘Ỉ↰᥏ሶ㧘ඨ⼱๋㇢ (2011) ࠾ࡎࡦࠩ࡞㓶ߩ㓸࿅߆ࠄߩ৻ᤨቅ┙ⴕേ. ᣣᧄູ੃㘃ቇળ 2011 ᐕᐲ
ቇⴚᄢળ (2011/09, ችፒ)㧚
Ỉ↰᥏ሶ㧘ඨ⼱๋㇢ (2011) ደਭፉߦ߅ߌࠆ࠾ࡎࡦࠩ࡞ߩࠠࡁࠦ㘩ⴕേ. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ
(2011/07/17, ›ጊ)㧚
ᷰㆻ㇌ᄦ㧘ࠗࠬ࡜ࡓ࡞࡮ࡂ࠺ࠖ㧘↰ਛବ᣿㧘ᧁ႐␞ሶ㧘㚅↰໪⾆ (2011) ࠗࡦ࠼ࡀࠪࠕ‫࠾ࡖࠫࡦ࡝ޔ‬ጊ㜞࿾ߦ
᫮߻ࠞ࠾ࠢࠗࠩ࡞ߩ᷷ᴰᶎⴕേ. ᣣᧄູ੃㘃ቇળ 2011 ᐕᐲᄢળ (2011/09/08-11, ችፒ)㧚
Hanya G (2012) Mammal abundance affects the distribution of terrestrial blood-feeding leeches in Yakushima. ᣣᧄ↢
ᘒቇળ╙ 50 ࿁ᄢળ (2012/03/21, ᄢᵤ)㧚
Otani Y, Sawada A, Hanya G (2012) Ranging behavior of male Japanese macaques. ╙ 59 ࿁ᣣᧄ↢ᘒቇળ࡮╙ 5 ࿁
EAFES㧔᧲ࠕࠫࠕ↢ᘒቇળㅪว㧕ᄢળ (2012/03, ṑ⾐⋵)㧚
ඨ⼱๋㇢ (2011) ࠨ࡞ߩᢙࠍ᳿߼ࠆ߽ߩ㧙ደਭፉߩ࠾ࡎࡦࠩ࡞ߩု⋥ಽᏓߣ␠ળᄌേ. ᣣᧄ↢ᘒቇળ╙ 59 ࿁
ᄢળ. ╙ 5 ࿁ᣣᧄ↢ᘒቇળᄢፉ⾨ฃ⾨⻠Ṷ㧚
ᷰㆻ㇌ᄦ (2011/11/03) ᧲ධࠕࠫࠕፉᎩㇱߩࠨ࡞㧦ߘߩ․ᓽߣ⒳ಽൻ‫ࠅࠊ߆߆ߩߣੱޔ‬. ╙ 6 ࿁ੱ㘃ቇ㑐ㅪቇળ
ද⼏ળวหࠪࡦࡐࠫ࠙ࡓ. ᴒ✽⋵┙ඳ‛㙚࡮⟤ⴚ㙚㧚
␠ળㅴൻಽ㊁
㧨⎇ⓥ᭎ⷐ㧪
A) ࠦࡦࠧ᳃ਥ౒๺࿖ᚢᤨਅߩ᫪ᨋߩ⎕უߣࡏࡁࡏ୘૕⟲ߦ߅ࠃ߷ߔᓇ㗀ߦߟ޿ߡߩ⎇ⓥ
ฎᏒ೰ผ㧘J Nackoney
1990 ᐕઍೋ㗡߆ࠄߩ᡽ᴦ⊛ᷙੂߣ‫ ߊ⛯ߦࠇߘޔ‬2 ᐲߩᚢ੎ߦࠃࠅ‫ޔ‬1973 ᐕએ᧪㊁↢ࡏࡁࡏ㧔Pan paniscus㧕ߩ⎇
ⓥࠍ⛯ߌߡ߈ߚ⿒㆏Ꮊ࡞ࠝ࡯ቇⴚ଻⼔඙ߩ᫪ᨋߣߘߎߦ૑߻㊁↢േ‛ߪ‫ޔ‬ᄢ߈ߥ࠳ࡔ࡯ࠫࠍฃߌߚ‫ߩߎޕ‬㑆ߩ᫪ᨋ
⎕უߩㅴⴕߣࡏࡁࡏ୘૕⟲ߦ߅ࠃ߷ߔᓇ㗀ࠍ⏕߆߼ࠆߚ߼‫ޔ‬NASA ߩឭଏߔࠆⴡᤊ↹௝ࠍ↪޿ߡ‫ޔ‬1990 ᐕ‫ޔ‬2000
ᐕ‫ޔ‬2005 ᐕߩ 3 ߟߩᤨᦼߦ߅ߌࠆ᫪ᨋߩ⁁ᴫࠍᲧセಽᨆߒߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬1990-2000 ᐕߩᤨᦼߦબណ࿾ߩ㕙Ⓧ߇⚂
3 ୚ߦߥߞߡ޿ࠆߎߣ‫ߩߘߚ߹ޔ‬ᄙߊ߇‫ޔ‬㆏〝߆ࠄ㔌ࠇߚ᫪ᨋߩᅏᷓߊߦὐ࿷ߔࠆዊߐߥબណ࿾ߢ޽ࠆߎߣ߇ࠊ߆
ߞߚ‫ᦼߩߎߜࠊߥߔޕ‬㑆‫ޔ‬᫪ߦㅏߍㄟࠎߛ૑᳃߇ኅᣖන૏ߩዊߐߥኅߣࠠࡖ࠶ࠨࡃ⇌ࠍ૞ߞߡ↢ᵴߒ‫ޔ‬᫪ᨋ߇ⱎߩ
Ꮍ⁁ߦߥߞߡ޿ߚߎߣ߇⏕߆߼ࠄࠇߚ‫ޕ‬᫪ߩᅏߦᢔ࿷ߔࠆዊߐߥዬ૑࿾ߦߪᓎੱ߿ઁߩ૑᳃ߩ⋡߽ዯ߆ߕ‫ࡏࡁࡏޔ‬
ߥߤߩ଻⼔₞ࠍ฽߼ߚ⁚⁸ߪ⥄↱ߦⴕ߁ߎߣ߇ߢ߈ࠆ‫⷗৻ޕ‬ᄢ߈ߥ⎕უࠍฃߌߡ޿ߥ޿ࠃ߁ߦ⷗߃ߥ߇ࠄ߽ᄢဳ߶
੃㘃߇߶ߣࠎߤ޿ߥ޿ ”empty forest” ᒻᚑߩࡔࠞ࠾࠭ࡓߩ߭ߣߟ߇‫⎇ᧄޔ‬ⓥߦࠃߞߡ⏕߆߼ࠄࠇߚ‫ޕ‬
㧙㧙
B) ࡏࡁࡏߩ␠ળ㑐ଥߣㆆേߦ㑐ߔࠆ⎇ⓥ
ฎᏒ೰ผ㧘R Heungjin㧘ᓼጊᄹᏔሶ
ࠦࡦࠧ᳃ਥ౒๺࿖࡞ࠝ࡯଻⼔඙ߦ↢ᕷߔࠆࡏࡁࡏࠍኻ⽎ߦ‫␠ޔ‬ળ㑐ଥߣㆆേߦ㑐ߔࠆ⎇ⓥࠍⴕߞߚ‫ޕ‬୘૕ㅊ〔ߦ
߽ߣߠߊⷰኤߦࠃߞߡ‫ޔ‬Უⷫߩ␠ળ⊛ⴕേ߇⥄ࠄߩ↢ᵴผࠬ࠹࡯ࠫߦࠃߞߡߤ߁ᄌൻߔࠆ߆ࠍಽᨆߔࠆߚ߼ߩ࠺࡯
࠲ࠍ෼㓸ߒߚ‫ߣࠆߔ࠼࡯࡝߇ࠬࡔޔߚ߹ޕ‬⠨߃ࠄࠇߡ޿ࠆࡏࡁࡏߩㆆേߩ࠲ࠗࡒࡦࠣ߿ᣇะ߇‫ޔ‬ታ㓙ߦߤߩࠃ߁ߥ
ࡔࠞ࠾࠭ࡓߢ᳿߹ࠆߩ߆ࠍ⺞ߴࠆߚ߼ߩ੍஻⊛⺞ᩏࠍⴕߞߚ‫ߡߞ߆߆ߦ⟂ޔߦࠄߐޕ‬㓸࿅ߦߟ޿ߡߎࠄࠇߥߊߥߞ
ߚ୘૕ࠍ⠉ᣣߦߥߞߡ㓸࿅ߩ߶߷ో୘૕ߢᝡ⚝ߦᚯࠆߣ޿߁‫ޔ‬ᐢߊ㔤㐳㘃ߢ߽ೋ߼ߡߣߥࠆ੐଀ࠍⷰኤߒ‫⺰ޔ‬ᢥߣ
ߒߡႎ๔ߒߚ‫ޕ‬
C) ࠬ࡝࡜ࡦࠞߦ↢ᕷߔࠆ㔤㐳㘃ߩⴕേ↢ᘒቇ⊛⎇ⓥ
MA Huffman, CAD Nahallage (University of Sri Jayewardenepura)
2004 ᐕᧃߦ㐿ᆎߒߚ‫↢ߦࠞࡦ࡜࡝ࠬޔ‬ᕷߔࠆ㊁↢㔤㐳㘃ߩಽᏓ⺞ᩏࠍ⛮⛯ߒߚ‫ޔߚ߈ߡߞⴕߢ߹ࠇߎޕ‬ධ⷏࡮
ධ࡮ධ᧲࡮ਛᄩ࡮ർ᧲࿾ၞߦ߅ߌࠆ‫࡞ࠩࠢ࠻ޔ‬㧔Macaca sinica㧕‫࡞࡯ࠣࡦ࡜࡟ࠣޔ‬㧔Semnopithecus priam thersites㧕‫ޔ‬
ࠞࠝࡓ࡜ࠨࠠ࡜ࡦࠣ࡯࡞㧔Trachypithecus vetulus㧕ߩಽᏓ⺞ᩏࠍ᜛ᄢߒ‫ోࠞࡦ࡜࡝ࠬޔ‬࿯ߦ߅ߌࠆಽᏓࠍ⏕߆߼ࠆ
ߚ߼ߦฦ⋵‫ޔ‬࿾඙࡟ࡌ࡞ߦࠕࡦࠤ࡯࠻⺞ᩏࠍ߅ߎߥߞߚ‫ޕ‬
D) ࡑࠞࠢߩᢥൻ⊛ⴕേ⎇ⓥ
MA Huffman, CAD Nahallage (University of Sri Jayawardenepura), JB Leca (University of Lethbridge)
⍹ㆆ߮ߥߤߩᢥൻ⊛ⴕേߩ␠ળ⊛ⷰኤቇ⠌࡮વ᠞ᯏ᭴ߩᲧセ⎇ⓥࠍⴕߞߚ‫ޕ‬
E) ࡌ࠻࠽ࡓߦ߅ߌࠆ‫ࠕ࡝࡜ࡑޔ‬વ᠞ⅣႺߩᄌኈߣੱ₞౒ㅢᗵᨴᕈࡑ࡜࡝ࠕߩ಴⃻ߩℂ⸃ߦะߌߚቇ㓙⊛⎇ⓥ
MA Huffman㧘ਛỈ⑲੺㧘R Culleton㧔㐳ፒᄢቇ㧕㧘೨㊁⧐ᱜ㧔⮮↰଻ஜⴡ↢ᄢቇ㧕㧘Ꮉวⷡ㧔₡දක⑼ᄢቇ㧕㧘QN
Yuyen㧘R Marchand (Khanh Phu Malaria Research Center,Medical CommitteeNetherlands-Vietnam)
2010 ᐕ߆ࠄࡌ࠻࠽ࡓ࡮ਛධㇱߦ޽ࠆࠞࡦࡈ࡯᧛ߩਐ㒺ㇱߦ᫮ᕷߔࠆ㊁↢ࠩ࡞ߩ⺞ᩏࠍ㐿ᆎߒߚ‫࡞ࠨߣ࠻ࡅޕ‬㑆
ߢવ᠞ߒ‫౒⇓ੱޔ‬ㅢᗵᨴࠍᒁ߈⿠ߎߔࡑ࡜࡝ࠕේ⯻ࠍ㊁↢ࠞ࠾ࠢࠗࠩ࡞߿ࡉ࠲ࠝࠩ࡞ߩ♮߆ࠄᬌ಴ߔࠆᣇᴺࠍ㐿⊒
ߒ‫ޔ‬㊁↢ࡑࠞࠢ㘃࡮ࡅ࠻㑆ᗵᨴߩ⎇ⓥࠍߔߔ߼ߚ‫ޕ‬
F) ࠾ࡎࡦࠩ࡞ߩነ↢⯻↢ᘒቇߣஜᐽ⛽ᜬߦኻߔࠆ㘩‛ㆬᛯߩᓎഀߦߟ޿ߡߩ⎇ⓥ
AJJ MacIntosh, MA Huffman
࠾ࡎࡦࠩ࡞ߩነ↢⯻ᗵᨴ∝‫ⴕޔ‬േᵴേࡄ࠲࡯ࡦ߿ᬀ‛ᕈ㘩‛ߦ㑐ߔࠆ⎇ⓥࠍ⛮⛯ߒߚ‫ޕ‬
G) ධࠕࡈ࡝ࠞ⷏ㇱጧߦ↢ᕷߔࠆ࠴ࡖࠢࡑࡅࡅߩណ㘩ⴕേ‫ޔ‬ነ↢⯻↢ᘒቇߣⓨ㑆೑↪ߩ↢ᘒቇ⊛⎇ⓥ
PA Pebsworth, MA Huffman
2009 ᐕ߆ࠄ㐿ᆎߒߚ‫ޔ‬ධࠕࡈ࡝ࠞ⷏ධㇱߦ޽ࠆࡢࠗ࡞࠼ࠢ࡝ࡈ⥄ὼ଻⼔࿾඙ߦ↢ᕷߔࠆ㊁↢࠴ࡖࠢࡑࡅࡅ⟲ߩ
⺞ᩏࠍ⛮⛯ߒߚ‫ޕ‬ᬀ‛ᕈ㘩‛ߩណ㘩ⴕേ‫ޔ‬࿯㘩޿ⴕേߣነ↢⯻ᗵᨴߣߩ࿃ᨐ㑐ଥࠍតߞߡ࠺࡯࠲⸃ᨆࠍⴕߞߚ‫ޕ‬
H) ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆᕈ⊛ࠪࠣ࠽࡞ߣ᳞ᗲⴕേ
C Garcia (CNRS, Paris)㧘L Rigaill (CNRS, Paris)㧘MA Huffman㧘FB Bercovitch
ࠝࠬߩ࠾ࡎࡦࠩ࡞㧔Macaca fuscata㧕ߩ੤የⴕേࠍ⺃⊒ߔࠆࡔࠬߩᕈ⊛ࠪࠣ࠽࡞ߣߘߩା㗬ᕈߦ㑐ߔࠆ⎇ⓥࠍⴕ
޿‫ߩࠬࡔޔ‬ᕈ⊹ߩ⦡ߣ൬޿߇ࠪࠣ࠽࡞ߣߥߞߡ޿ࠆ߆ߤ߁߆ࠍ⺞ߴߚ‫ޕ‬㔤㐳㘃⎇ⓥᚲߢ㘺⢒ߐࠇߡ޿ࠆ୘૕ࠍኻ⽎
ߦ‫ⴕޔ‬േⷰኤ㧔ᕈⴕേ㧕‫ޔ‬ᕈ⊹ߩ࠺ࠫ࠲࡞౮⌀᠟ᓇ㧔ᕈ⊹ߩ⦡ߩቯ㊂⊛⹏ଔ㧕‫⸃ࡦࡕ࡞ࡎޔ‬ᨆ㧔♮ਛߩෆࠬ࠹ࡠࠗ
࠼㧕ࠍⴕߞߚ‫⎇ᧄޕ‬ⓥߪࡅ࠻ߩᕈࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߦᓇ㗀ߔࠆ↢ᘒቇ⊛࡮ㅴൻ⊛ߥⷐ࿃ߩℂ⸃ߦነਈߔࠆߣ⠨߃
ࠄࠇࠆ‫ޕ‬
I) 㘩‛ଏ⛎ߩᐕᰴᄌേߣ࠾ࡎࡦࠩ࡞ߩ୘૕⟲ࡄ࡜ࡔ࡯࠲ߩ㑐ଥߦߟ޿ߡߩ⎇ⓥ
ㄞᄢ๺
⟲ࠇ↢ᵴߔࠆേ‛ߩ႐ว‫ޔ‬㘩‛߇㒢ࠄࠇࠆ႐วߦߘࠇࠍᎼࠆ┹ว߇↢ߓ‫ޔ‬㗼࿷ൻߒߚ㗅૏㑐ଥߩᓇ㗀߇ណ㘩ᚑഞ
߿❥ᱺᚑഞߦ෸߱႐ว߇޽ࠆ‫┹ޕ‬วߩᒝߐߪ㘩‛ߩଏ⛎⁁ᘒ߆ࠄᓇ㗀ࠍฃߌ‫▎৻ޔ‬ᚲߦ㓸ਛߒߡ޿ࠇ߫ఝ૏ߥ୘૕
߇⁛භߢ߈‫ޔ‬ಽᢔߒߡ޿ࠇ߫⟲ࠇߩోຬ߇೑↪ߢ߈ࠆߛࠈ߁‫ߩ⑺ޕ‬ਥⷐ㘩‛ߢ޽ࠆၷᨐ㘃ߩଏ⛎⁁ᘒߪ᮸⒳ߏߣߦ
⇣ߥࠅ‫↥↢ߩߘ߽߆ߒޔ‬㊂߿⚿ታ᮸⒳ߩ⚵ߺวࠊߖ߇ᐕߦࠃࠅᄢ߈ߊᄌൻߔࠆ‫ޔߡߞ߇ߚߒޕ‬㘩‛ࠍᎼࠆ┹วߩ⒟
ᐲ߽ᐕߏߣߦᄢ߈ߊᄌൻߒ‫ޔ‬ណ㘩ᚑഞߩ㗅ߦᏅߪᦨ⚳⊛ߦ❥ᱺᚑഞߦᓇ㗀ߔࠆߣ੍ᗐߐࠇࠆ‫⎇ᧄޕ‬ⓥߢߪߎࠇ߹ߢ
ߩ㐳ᦼ⎇ⓥߢᓧߚ࠺࡯࠲ࠍ⸃ᨆߔࠆߎߣߦࠃࠅ‫ޔ‬㘩‛⾗Ḯߩଏ⛎⁁ᘒߩᄌേ߇㗅૏㑐ଥࠍㅢߓߡ୘૕⟲ࡄ࡜ࡔ࡯࠲
ߦ෸߷ߔᓇ㗀ࠍ᣿ࠄ߆ߦߒߚ‫ޔߚ߹ޕ‬⒳ሶ࠻࡜࠶ࡊߦࠃࠆ⚿ታ㊂ߩ⺞ᩏࠍᧄᐕᐲ߽⛮⛯ߒߡⴕߞߚ‫ޕ‬
J) ࠾ࡎࡦࠩ࡞ߩᴱ߹ࠅ႐ㆬᛯߦ㑐ߔࠆ⎇ⓥ
ㄞᄢ๺
࠾ࡎࡦࠩ࡞ߩᴱ߹ࠅ႐ㆬᛯߦਈ߃ࠆᬀ↢࠲ࠗࡊ࡮࿾ᒻ࡮ᮡ㜞ߩᓇ㗀ࠍ⺞ߴߚ‫ޕ‬᝝㘩⠪߇޿ߥ޿㊄⪇ጊߢߪ‫ޔ‬㘩‛
ߩ⼾ንߥ႐ᚲ߿‫ޔ‬㘑ࠍߒߩߍࠆ⼱╭‫ߪ޿ࠆ޽ޔ‬ᮡ㜞ߩૐ޿႐ᚲ߇ᴱ߹ࠅ႐ߣߒߡㆬᛯߐࠇࠆ௑ะ߇ᒝ޿ߎߣࠍ᣿ࠄ
߆ߦߒߚ‫ޕ‬
K) ࡑࠞࠢ㘃࡮ࠦࡠࡉࠬ㘃ߩ㘩ᕈߩ࿾ၞᄌ⇣ߦ㑐ߔࠆ⎇ⓥ
㧙㧙
ㄞᄢ๺㧘๺↰৻㓶㧘ᷰㆻ㇌ᄦ㧘ඨ⼱๋㇢㧘CC Grueter (Max Plank Institute of Evolutionally Anthropology)
ㆊ෰ߦ౏⴫ߐࠇߚ㔤㐳㘃㧔ࡑࠞࠢ㘃‫ࠬࡉࡠࠦޔ‬㘃㧕ߩ㘩ᕈ⾗ᢱࠍ෼㓸࡮ಽᨆߒ‫ޔ‬࿾ၞᄌ⇣ߣߘࠇࠍ߽ߚࠄߔⅣႺ
ⷐ࿃ߦߟ޿ߡ⸃ᨆࠍⴕߞߚ‫ޕ‬
L) ᣣᧄ↥ູ੃㘃ߩ⒳ሶᢔᏓߦ㑐ߔࠆ⎇ⓥ
ㄞᄢ๺㧘⷗ᶆᴕ⡍ሶ㧘ዊጼථ਽㧘᧛੗ੳᔒ㧔ንጊᏒࡈࠔࡒ࡝࡯ࡄ࡯ࠢ㧕
࠾ࡎࡦࠩ࡞ߦࠃࠆ⒳ሶᢔᏓߩ⎇ⓥߩ৻Ⅳߣߒߡ‫᧲ޔ‬੩㇠ᄖߩ࠾ࡎࡦࠩ࡞ߩ♮ߦ฽߹ࠇࠆ⒳ሶߩ․ᕈ㧔ᬀ‛ߩ⒳ᢙ‫ޔ‬
♮৻୘޽ߚࠅߦ฽߹ࠇࠆ⒳ሶߩᢙ‫ޔ‬ஜో₸ߥߤ㧕ߣߘߩቄ▵ᄌൻࠍ⹏ଔߔࠆߣߣ߽ߦ‫ߩઁޔ‬࿾ၞߣᲧセߒߚ‫ߞ޿ޕ‬
߸߁‫ޔ‬ᤓᐕᐲߦᒁ߈⛯߈㘺⢒᧦ઙਅߩࡎࡦ࠼࠹ࡦ㧔Martes melampus㧕ࠍኻ⽎ߦ⛎㙄ታ㛎ࠍⴕ޿‫ޔ‬⒳ሶߩឃᴭᤨ㑆ࠍ
⹏ଔߒߚ‫ᧄޕ‬ᐕᐲߪ㘩‛ߩ៨ข㊂߇૕ౝㅢㆊᤨ㑆ߦਈ߃ࠆᓇ㗀ߦߟ޿ߡ⺞ߴ‫ޔ‬៨ข㊂ߩᓇ㗀ߪߣߊߦ⷗ࠄࠇߥ޿ߎ
ߣࠍ᣿ࠄ߆ߦߒߚ‫ޕ‬
M) ᧲ධࠕࠫࠕ↥㔤㐳㘃ߩណ㘩↢ᘒߦ㑐ߔࠆ⎇ⓥ
ㄞᄢ๺㧘B Suryobroto㧘I Hadi㧘KA Udayati㧔ࡏࠧ࡯࡞ㄘ⑼ᄢቇ㧕㧘MV Nguyen㧘VH Nguyen㧔ࡈࠛᄢቇ㧕
ࠗࡦ࠼ࡀࠪࠕ࡮ࡄࡦࠟࡦ࠳࡜ࡦ⥄ὼ଻⼔඙߅ࠃ߮ࡌ࠻࠽ࡓ࡮࠰ࡦ࠴ࡖ⥄ὼ଻⼔඙ߢࠫࡖࡢ࡞࠻ࡦ㧔Trachypithecus
auratus㧕‫࡞࡯ࠣࡦ࡜ࠢ࠘࠼ࡀࠬࠞࠕޔ‬㧔Pygathrix nemaeus㧕‫࡞ࠩࠗࠢ࠾ࠞޔ‬㧔Macaca fascicularis㧕‫ࠥࠞࠕ߮ࠃ߅ޔ‬
ࠩ࡞㧔Macaca mulatta㧕ߩၮ␆↢ᘒߦ㑐ߔࠆ⺞ᩏࠍⴕ޿‫ޔ‬ᓐࠄߩ㘩ᕈ‫ޔ‬ᵴേᤨ㑆㈩ಽ‫ߩઁޔ‬േ‛ߣߩ⒳㑆㑐ଥߥߤ
ߩ࠺࡯࠲ࠍ෼㓸ߒߚ‫⺞ߪߢࠕࠪࡀ࠼ࡦࠗޕ‬ᩏ࿾ౝߦᬀ↢ࡊࡠ࠶࠻ࠍ⸳⟎ߒ‫ߩ࠻ࡦ࠲ࠬࠪࠕޔ‬දജࠍᓧߡࡈࠚࡁࡠࠫ
࡯⺞ᩏࠍᲤ᦬ⴕߞߚ‫ޕ‬
N) ㊁↢ࡏࡁࡏߩ㓸࿅᭴ㅧߩ↢ᚑߦߟ޿ߡߩ⎇ⓥ
ဈᏎື਽
㊁↢ࡏࡁࡏߩᲫߠߊࠈ޿੤ᷤ‫ޔ‬㓸࿅ࡔࡦࡃ࡯ߩ಴ળ޿ߩ੤ᷤ‫ޔ‬⒖౉ࡔࠬߩ␠ળ㑐ଥߩᄌㆫߦ㑐ߔࠆ࠺࡯࠲෼㓸ߣ
ಽᨆ‫⚿ޔ‬ᨐߩ౏⴫ࠍㅴ߼ߚ‫ޕ‬㓸࿅㑆㑐ଥࠍ࠹࡯ࡑߦ‫ޔ‬ᣂߚߥ⺞ᩏ㓸࿅ߩㅊ〔ⷰኤ‫ޔߌߠੱޔ‬୘૕⼂೎ࠍ⛮⛯ߒߚ‫ޕ‬
ⴕേߩᄙ᭽ᕈࠍ࠹࡯ࡑߦ‫ޔ‬㓞ធ୘૕⟲ߩ⺞ᩏ߽⛮⛯ߒߚ‫ޕ‬
O) ㊁↢ࡏࡁࡏ࡮࠴ࡦࡄࡦࠫ࡯ߩᐢၞ⺞ᩏ
┻రඳᐘ
ⅣႺ⋭ⅣႺ⎇ⓥផㅴ⾌‫ޟ‬㜞ੱญኒᐲ࿾ၞߦ߅ߌࠆቅ┙ߒߚ㔤㐳㘃୘૕⟲ߩᜬ⛯⊛଻⼔▤ℂ‫৻ߩ࠻ࠢࠚࠫࡠࡊߩޠ‬
Ⅳߣߒߡ‫ࠧࡦࠦޔ‬᳃ਥ౒๺࿖࿾ၞߦ↢ᕷߔࠆࡏࡁࡏ㧔Pan paniscus㧕߅ࠃ߮࠙ࠟࡦ࠳౒๺࿖‫౒ࠕ࠾ࠡޔ‬๺࿖ߦ↢ᕷ
ߔࠆ࠴ࡦࡄࡦࠫ࡯㧔P.troglodytes㧕ߩ㊁↢୘૕⟲ߩ⺞ᩏࠍ߅ߎߥߞߚ‫ߩࠇߙࠇߘޕ‬࿾ၞߢ↢ᘒቇ⊛⺞ᩏࠍ߅ߎߥ޿‫ޔ‬
หᤨߦㆮવቇ⊛߅ࠃ߮ੱ₞౒ㅢᗵᨴ∝⸃ᨆߩߚ߼ߩ⹜ᢱࠍ෼㓸ߒߚ‫ޕ‬
P) ࠾ࠪࡠ࡯࡜ࡦ࠼ࠧ࡝࡜ߩ␠ળ⊛⊒㆐ߣⅣႺߩ㑐ㅪߦߟ޿ߡߩ⎇ⓥ
᧻ේᐙ
2010 ᐕߦࠗࠡ࡝ࠬධㇱߩࡂ࠙࡟࠶࠷࡮ࡐ࡯࠻࡝ࡓ㊁↢േ‛౏࿦ߢⴕߞߚ࠾ࠪࡠ࡯࡜ࡦ࠼ࠧ࡝࡜ߩ␠ળ⊛⊒㆐⺞
ᩏߢ෼㓸ߒߚⴕേ࠺࡯࠲ࠍ⸃ᨆߒߚ‫ޕ‬10 ᐕએ਄߆ߌߡࠦ࠼ࡕ߇ᄙߊ⢒ߞߚ⟲ࠇߣ‫ߩࠇ⟲ޔ‬ഃ⸳߆ࠄ 5 ᐕ⋡ߩ⟲ࠇ
ߩࠦ࠼ࡕߦ߅ߌࠆㆆ߮ⴕേࠍᲧセߔࠆߎߣߢ‫ߩࡕ࠼ࠦޔ‬ᚑ㐳⊒㆐ᤨߦ߅ߌࠆઁ୘૕ߩᓇ㗀ࠍ⺞ߴߚ‫࠽࠻ࠝޔߚ߹ޕ‬
ࡔࠬห჻߿ࠝ࠻࠽ࡔࠬߣࠦ࠼ࡕߩㆆ߮ⴕേߩ⸃ᨆࠍⴕ޿‫ޔ‬1 ᱦએ਄ߩࠧ࡝࡜ߦ߅ߌࠆ㕖ⴊ✼ࡔࠬߣߩ␠ળ੤ᷤ߇‫ޔ‬
⧯޿⟲ࠇߦ߅޿ߡ㗫❥ߦ⷗ࠄࠇࠆߎߣ߆ࠄ‫ߩࡕ࠼ࠦޔ‬ዋߥ޿ഃ⸳㑆߽ߥ޿⟲ࠇߢߪ‫ޔ‬㕖ⴊ✼ࡔࠬ߇ࠦ࠼ࡕߩ␠ળ⊛
ࠬࠠ࡞ߩ⊒㆐ߦᓇ㗀ࠍਈ߃ߡ޿ࠆน⢻ᕈ߇ផ᷹ߐࠇߚ‫ޕ‬
Q) ࠾ࡎࡦࠩ࡞߇ォᇾ᡹᠄ࠍⴕ߁ᗧ⟵ߩ⸃᣿
ᓼጊᄹᏔሶ
᡹᠄ࠍฃߌߚ࠾ࡎࡦࠩ࡞ߪ‫ߦߋߔޔ‬೨ߩ᡹᠄ߦߪ㑐ࠊࠅߥ޿୘૕ߦ᡹᠄ߔࠆߎߣ߇޽ࠆ㧔ォᇾ᡹᠄㧕‫ⵍޕ‬᡹᠄୘
૕ߦߣߞߡߩォᇾ᡹᠄ࠍⴕ߁ᗧ⟵ࠍ᣿ࠄ߆ߦߔࠆߚ߼‫ޔ‬፲ጊࡕࡦࠠ࡯ࡄ࡯ࠢߩ㙄ઃߌ⟲ࠍኻ⽎ߦ⺞ᩏࠍⴕߞߚ‫ޕ‬
R) 㘺⢒ਅࡏࡁࡏ࡮࠴ࡦࡄࡦࠫ࡯ߩࠕࠢ࠹ࠖࡆ࠹ࠖߦ㑐ߔࠆ⎇ⓥ
H Sojung
㊁↢ߩࡏࡁࡏߩࡔࠬߪ࠴ࡦࡄࡦࠫ࡯ߩࡔࠬߣᲧߴߡ᡹᠄ᕈ߇ૐ޿ߣߐࠇߡ߅ࠅ‫ߩߘޔ‬㆑޿ߪਔ⠪ߩ↢ᕷⅣႺߩ㆑
޿‫ߒ޿ߥޔ‬ਔ⠪ߩᕈฃኈᕈߩ㆑޿ߢ⺑᣿ߢ߈ࠆߣ⠨߃ࠄࠇࠆ‫ߩߎޕ‬໧㗴ࠍ⠨߃ࠆ♻ญߣߒߡ‫ࠫࡦࡄࡦ࠴ߣࡏࡁࡏޔ‬
࡯߇߶߷ห৻ߩⅣႺߢ㘺⢒ߐࠇߡ޿ࠆ࠼ࠗ࠷ߩ࡜ࠗࡊ࠴ࡅേ‛࿦ߦ߅޿ߡ 6 ࡩ᦬㑆ߦࠊߚࠅਔ⒳ߩⴕേ࠺࡯࠲ߩ
෼㓸ࠍⴕߞߚ‫ޕ‬
S) ࡔࠬ࠾ࡎࡦࠩ࡞ߩട㦂ߦ઻߁ࡃࠗࠝࡑ࡯ࠞ࡯DHES ߩᄌൻߩ⎇ⓥ
RS Cicalise Takeshita㧘MA Huffman㧘FB Bercovitch㧘ᷡ᳓ᘮሶ㧔ጟጊℂ⑼ᄢ㧕
ࠬ࠹ࡠࠗ࠼ࡎ࡞ࡕࡦߩ߁ߜ‫ࡦࡠ࠹ࠬࡠ࠼ࡦࠕࡇࠛࡠ࠼ࡅ࠺ޔ‬㧔DHEA㧕ߣߘߩ⎫㉄ࠛࠬ࠹࡞㧔DHEA-S㧕ߪ‫ࡅޔ‬
࠻߅ࠃ߮ࡅ࠻એᄖߩ㔤㐳㘃ߩ⧯㦂୘૕ߦᐢߊߺࠄࠇࠆࡎ࡞ࡕࡦߢ޽ࠆ‫ޕ‬DHEA-S ߩỚᐲߪ࠴ࡦࡄࡦࠫ࡯‫ࠩࠥࠞࠕޔ‬
࡞‫࡞ࠩࡀ࠷ࠠޔ‬㘃‫ߪߢ࠻ࡅޔ‬ᐕ㦂ߣ౒ߦૐਅߔࠆߚ߼‫ޔ‬⠧ൻߩ↢‛ቇ⊛ࡑ࡯ࠞ࡯ߣߒߡᓎ┙ߟߣ⠨߃ࠄࠇߡ޿ࠆ‫ޕ‬
ߒ߆ߒ‫ޔ‬ᐕ㦂ߣ఺∉ቇ⊛ߥᯏ⢻ߣߩ㑐ଥߪߎࠇ߹ߢ߶ߣࠎߤ᣿ࠄ߆ߦߐࠇߡ޿ߥ޿‫⎇ᧄޕ‬ⓥߢߪ㘺⢒ਅߩ࠾ࡎࡦࠩ
㧙㧙
࡞ࠍኻ⽎ߦ‫♮ޔ‬ਛߩ DHEA-S Ớᐲߩᐕ㦂ᄌൻ㧔1-26 ᱦ㧕߅ࠃ߮ቄ▵ᄌൻࠍ⺞ߴߚ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
Garcia C, Huffman MA, Shimizu K, Redman P, Speakman JR (2011) Energetic consequences of seasonal breeding in
female Japanese macaques (Macaca fuscata). American Journal of Physical Anthropology 146:161-170.
2) Kaur T, Singh J, Huffman MA, Petrzelkova KJ, Taylor NS, Xu S, Dewhirst FE, Paster BJ, Debrunyne L, Vandamme P,
Fox JG (2011) Campylobacter troglodytes – sp. nov., isolated from feces of human-habituated wild chimpanzees (Pan
troglodytes schweinfurthii) in Tanzania. Applied and Environmental Microbiology 77:2366-2373.
3) Kim TW, Ryu HJ, Choi JB, Choe JC (2011) Tower construction by the manicure crab Cleistostoma dilatatum during dry
periods on an intertidal mudflat. Journal of Ethology 29:459-465.
4) Leca J-B, Gunst N, Huffman MA (2011) Complexity in object manipulation by Japanese macaques (Macaca fuscata): A
cross-sectional analysis of manual coordination in stone handling patterns. Journal of Comparative Psychology
125:61-71.
5) Petrášová J, Uzliková M, Kostka M, Petrželková KJ, Huffman MA, Modrý D (2011) Diversity and host specificity of
Blastocystis in syntopic primates on Rubondo Island, Tanzania. International Journal of Primatology 11:1113-1120.
6) Sakamaki T (2011) Submissive pant-grunt greeting of female chimpanzees in Mahale Moutains National Park, Tanzania.
African Study Monographs 32:25-41.
7) Sugiura H, Shimooka Y, Tsuji Y (2011) Variation in spatial cohesiveness in a group of Japanese macaques (Macaca
fuscata). International Journal of Primatology 32:1348-1366.
8) Tsuji Y (2011) Sleeping site preferences of wild Japanese macaques (Macaca fuscata): the importance of non-predatory
factors. Journal of Mammalogy 92:1261-1269.
9) Tsuji Y (2011) Seed dispersal by Japanese macaques (Macaca fuscata) in western Tokyo, Japan: a preliminary report.
Mammal Study 36:165-168.
10) Nahallage CAD, Huffman MA (2012) Stone handling behavior in rhesus macaques (Macaca mulatta), a behavioral
propensity for solitary object play shared with Japanese macaques. Primates 53:71-78.
11) Pebsworth PA, Bardi MA, Huffman MA (2012) Geophagy in chacma baboons: Patterns of soil consumption by age
class, sex, and reproductive state. American Journal of Primatology 74:48-57.
12) Tsuji Y, Takatsuki S (2012) Inter-annual variation in nut abundance is related to agonistic interactions of foraging
female Japanese macaques (Macaca fuscata). International Journal of Primatology 33:489-512.
✚⺑
1)
Furuichi T (2011) Female contributions to the peaceful nature of bonobo society. Evolutionary Anthropology
20:131-142.
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
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8)
Furuichi T, Idani G, Ihobe H, Hashimoto C, Tashiro Y, Sakamaki T, Mulavwa BN, Yangozene K, Kuroda S (2011)
Long-term studies on wild bonobos at Wamba, Luo Scientific Reserve, D.R. Congo: towards the understanding of
female life history in a male-philopatric species. (Long-term field studies of primates) (ed. Kappeler P, Watts D)
p.413-433 Springer-Verlag, Berlin, Heidelberg.
Yamagiwa J, Basabose AK, Kahekwa J, Bikaba D, Ando C, Matsubara M, Iwasaki N, Sprague DS (2011) Long-term
research on Grauer's gorillas in Kahuzi-Biega National Park, DRC: life history, foraging strategies, and ecological
differentiation from sympatric chimpanzees. (Long-term field studies of primates) (ed. Kappeler PM, Watts DP)
p.385-412 Springer, New York.
Chalmers A, Huffman MA, Koyama N, Takahata Y (2012) Fifty years of female macaque demography at Arashiyama,
with special reference to long-lived female (> 25 years). (The Monkeys of Stormy Mountain: 60 Years of Primatological
Research on the Japanese Macaques of Arashiyama.) (ed. Leca JB, Huffman MA, Vasey PL) p.51-67 Cambridge
University Press, Cambridge.
Huffman MA, Vasey P, Leca JB (2012) Introduction. (The Monkeys of Stormy Mountain: 60 Years of Primatological
Research on the Japanese Macaques of Arashiyama.) (ed. Leca JB, Huffman MA, Vasey PL) p.1-10 Cambridge
University Press, Cambridge.
Huffman MA, Fedigan LM, Leca J-B, Vasey P (2012) A brief historical time-line of research on the Arashiyama
macaques. (The Monkeys of Stormy Mountain: 60 Years of Primatological Research on the Japanese Macaques of
Arashiyama) (ed. Leca JB, Huffman MA, Vasey PL) p.13-27 Cambridge University Press, Cambridge.
Huffman MA, MacIntosh AJJ (2012) Plant-food diet of the Arashiyama Japanese macaques and its potential medicinal
value. (The Monkeys of Stormy Mountain: 60 Years of Primatological Research on the Japanese Macaques of
Arashiyama.) (ed. Leca JB, Huffman MA, Vasey PL) p.356-431 Cambridge University Press, Cambridge.
Huffman MA, Takahata Y (2012) Long-term Trends in Mating Relations of Japanese Macaques at Arashiyama, Japan.
(The Monkeys of Stormy Mountain: 60 Years of Primatological Research on the Japanese Macaques of Arashiyama) (ed.
Leca JB, Huffman MA, Vasey PL) p.71-86 Cambridge University Press, Cambridge.
Leca JB, Gunst N, Huffman MA (2012) Thirty years of stone handling tradition in Arashiyama-Kyoto macaques:
implications for cumulative culture and tool use in non-human primates. (The Monkeys of Stormy Mountain: 60 Years
㧙㧙
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of Primatological Research on the Japanese Macaques of Arashiyama) (ed. Leca JB, Huffman MA, Vasey PL)
p.223-257 Cambridge University Press, Cambridge.
ㄞᄢ๺ (2012) ࠪࠞߩ⪭ߜⓄᜪ޿㧙ࡈࠖ࡯࡞࠼ࡁ࡯࠻ߩ⸥㍳߆ࠄ. ‫ޟ‬ᐔᚑ 24 ᐕᐲ ਛቇᩞ‫ޡ‬࿖⺆‫ ޠޢ‬p.118-127
శ᧛࿑ᦠ಴ 㧚
✬㓸
1)
Leca JB, Huffman MA, Vasey PL (2012) The Monkeys of Stormy Mountain: 60 Years of Primatological Research on
the Japanese Macaques of Arashiyama. p.498 Cambridge University Press, Cambridge.
ߘߩઁߩၫ╩
1)
2)
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4)
ฎᏒ೰ผ (2011) ⃻ઍߩߎߣ߫㧦ⓨ⊕ߩ 5 ᤨ㑆. 2011 ᐕ 5 ᦬ 6 ᣣᄕೀ ੩ㇺᣂ⡞㧚
ฎᏒ೰ผ (2011) ⃻ઍߩߎߣ߫㧦ടㅦߔࠆᄌൻ. 2011 ᐕ 6 ᦬ 24 ᣣᄕೀ ੩ㇺᣂ⡞㧚
Koyama N (2012) Touches of Humanity in monkey society. In: The Monkeys of Stormy Mountain: 60 Years of
Primatological Research on the Japanese Macaques of Arashiyama. Leca JB, Huffman MA, Vasey PL p.42-50
Cambridge University Press, Cambridge, (translated by Huffman MA).
Ohta E (2012) In search of the phantom monkeys. In: The Monkeys of Stormy Mountain: 60 Years of Primatological
Research on the Japanese Macaques of Arashiyama. Leca JB, Huffman MA, Vasey PL p.28-33 Cambridge University
Press, Cambridge, (translated by Huffman MA).
ቇળ⊒⴫
1)
2)
3)
4)
5)
6)
7)
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13)
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16)
Huffman MA (2011) ࠨ࡞ࡑ࡜࡝ࠕߩኋਥߣߥࠆ᧲ධࠕࠫࠕߩࡑࠞࠢዻࠍㅊߞߡ. ᣣᧄቇⴚᝄ⥝ળࠕࠫࠕ࡮ࠕࡈ
࡝ࠞቇⴚၮ⋚ᒻᚑ੐ᬺ㧦᫪ᨋ⊒↢ࡅ࠻ࡑ࡜࡝ࠕߣੱ₞౒ㅢᗵᨴ∝ࠨ࡞ࡑ࡜࡝ࠕߩ↢ᘒ⎇ⓥળ (2011/07, ₡දක
⑼ᄢቇ)㧚
Huffman MA (2012) How primates deal with disease and zoonoses potentially affecting humans in the tropics.
International Symposium on Forest Malaria: Man-Mosquitoes-Monkeys (2012/03, Nha Trang, Vietnam).
Huffman MA (2012) 㔤㐳㘃ߩ⥄Ꮖᴦ≮ⴕേ㧙੍㒐ߣᴦ≮. One World, One Health Symposium, ␹ᚭࠕ࠾ࡑ࡞ࠤ
ࠕ࿖㓙ળ⼏ (2012/02, ␹ᚭ)㧚
Huffman MA, Nahallage CAD (2011) Non-invasive methods for the study of the diseases of monkeys and man in nature.
International Symposium: Integrative Research on Monkeys, Malaria and Man in Asia (2011/06, Colombo, Sri Lanka).
Nahallage CAD, Huffman MA (2011) Primates of Sri Lanka. International Symposium: Integrative Research on
Monkeys, Malaria and Man in Asia (2011/06, Colombo, Sri Lanka).
Tokuyama N (2011) Why do Japanese macaques perform redirected aggression? International Symposium for
Conservation of Wild Bonobos and Chimpanzees, Research Center of Ecology and Forest (2011/08, Kinshasa, DRC).
Tsuji Y, Fujita S, Sugiura H, Nakagawa N (2012) Rome was not built in a day: time to grasp information on plant
feeding of wild Japanese macaques (Macaca fuscata). ESJ59/EAFES5 (2012/03, Otsu).
Tsuji Y, Miura S, Shiraishi T (2011) Gastrointestinal passage time of seeds ingested by captive Japanese martens Martes
melampus. ᣣᧄູ੃㘃ቇળ (2011/09, ችፒᏒ)㧚
ฎᏒ೰ผ (2011) ࡏࡁࡏ㧦᳓߇ࡏࡁࡏࠍ↢ࠎߛ㨪ࠦࡦࠧ⋆࿾ߩࡏࡁࡏߩㅴൻߣ↢ᵴ. SAGA ࠪࡦࡐࠫ࠙ࡓ
(2011/11, ᾢᧄᏒേ‛࿦)㧚
ᯅᧄජ⛗㧘቟ጟብ๺㧘ᚻႦ⾫⥋㧘ฎᏒ೰ผ (2011) ࠙ࠟࡦ࠳౒๺࿖ߦ߅ߌࠆ᫪ᨋ଻⼔඙๟ㄝߩ࿾ၞ૑᳃ߦࠃࠆ
᫪ᨋ⾗Ḯߩ೑↪ߩታᘒ. ᣣᧄ㔤㐳㘃ቇળ (2011/07, ›ጊᏒ)㧚
᧻ේᐙ (2011) ࠾ࠪࡠ࡯࡜ࡦ࠼ࠧ࡝࡜ߩㆆ߮ⴕേߦਈ߃ࠆኅᣖ᭴ㅧߩᓇ㗀. ᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕേ
ቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓▤ℂቇળวหᄢળ (2011/09, ᘮᔕ⟵Ⴖᄢቇ)㧚
Nackoney J㧘ฎᏒ೰ผ㧘Baraldi A㧘Nolinari G (2011) ࠦࡦࠧ᳃ਥ౒๺࿖ߩᚢᤨਅߩ࡞ࠝ࡯଻⼔඙๟ㄝߩ᫪ᨋߩ
⎕უߣࡏࡁࡏ୘૕⟲ߦ߅ࠃ߷ߔᓇ㗀ߩࡕ࠾࠲࡝ࡦࠣ. ᣣᧄੱ㘃ቇળ (2011/11, ᴒ✽⋵┙ඳ‛㙚࡮⟤ⴚ㙚)㧚
᧖ᶆ⑲᮸㧘ਅጟࠁ߈ሶ㧘ㄞᄢ๺ (2011) ࠾ࡎࡦࠩ࡞㊁↢⟲ߦ߅ߌࠆ⟲ࠇߩಝ㓸ᕈߩ⛽ᜬᯏ᭴. ᣣᧄ㔤㐳㘃ቇળ
(2011/07, ›ጊᏒ)㧚
᧖ᶆ⑲᮸㧘ਅጟࠁ߈ሶ㧘ㄞᄢ๺ (2011) ࠾ࡎࡦࠩ࡞ߪ⟲ࠇߩ߹ߣ߹ࠅࠍ଻ߟߚ߼ߦ㧘ߤߩࠃ߁ߥേ߈ࠍߒߡ޿
ࠆ߆. ᣣᧄູ੃㘃ቇળ (2011/09, ችፒᏒ)㧚
ᓼጊᄹᏔሶ (2011) ࠾ࡎࡦࠩ࡞ߪߥߗੑᰴ᡹᠄ࠍⴕ߁ߩ߆. ᣣᧄ㔤㐳㘃ቇળ (2011/07, ›ጊᏒ)㧚
ㄞᄢ๺ (2011) ㊄⪇ጊፉߦ߅ߌࠆ࠾ࡎࡦࠩ࡞ߩ↢ᘒ⎇ⓥ㧙㐳ᦼ⺞ᩏ߆ࠄߺ߃ߡ߈ߚߎߣ㧙. ᣣᧄູ੃㘃ቇળ
(2011/09, ችፒᏒ)㧚
⻠Ṷ
1)
2)
3)
4)
Huffman MA (2011/04) The evolution of self-medication in the animal kingdom and the origins of traditional medicine
in humans. Department of Wild and Resources Faculty Seminar Utah State University.
Huffman MA (2011/05) The evolution of self-medication in the animal kingdom. Visiting Speaker in Primatology Series,
Department of Anthropology University of Calgary.
Huffman MA (2011/09) Chimpanzee use of plants as medicine and the evolution of self-medication in primates and
other animals, Living Links to Human Biology. Mind and Medicine Lecture Series of the Royal Zoological Society of
Scotland, Edinburgh Zoo.
Huffman MA (2012/01) ࠬ࡯ࡄ࡯ࠨࠗࠛࡦࠬࡂࠗࠬࠢ࡯࡞⻠⟵㧚৻ች㜞ᩞ㧚
㧙㧙
5)
6)
7)
8)
9)
10)
11)
12)
13)
Huffman MA (2011/05) The evolution of self-medication in the animal kingdom: great and small minds ‘think’ alike.
taff Development Colloquium Washington National Zoo㧚
Tsuji Y (2011/08) Feeding ecology of silvered leaf monkeys at Pangandaran Nature Reserve. West Java, Indonesia: a
preliminary report, IPB Bogor, Indonesia.
ฎᏒ೰ผ (2011/10) ࡏࡁࡏߪࡅ࠻ߩ૗ࠍ⺆ࠆ߆. ࡕࡦࠠ࡯ࠞ࡟࠶ࠫ, ›ጊᏒ ᣣᧄࡕࡦࠠ࡯࠮ࡦ࠲࡯㧚
ฎᏒ೰ผ (2011/11) ࡏࡁࡏ㧘ᦨᓟߩ㘃ੱ₎. ╙ 2 ࿁୯ߩ᜚ὐ࠮ࡒ࠽࡯ ੩ㇺᄢቇ᧲੩ࠝࡈࠖࠬ, ᧲੩㧚
ฎᏒ೰ผ (2011/11) ࡏࡁࡏߩ૑߻ࠦࡦࠧ⋆࿾ߩᄢᾲᏪ㔎ᨋ㧦ߘߩ⃻⁁ߣ዁᧪. ࡊ࡝ࡑ࡯࠹ࠬ⎇ⓥળ ᣣᧄࡕࡦ
ࠠ࡯࠮ࡦ࠲࡯, ›ጊᏒ㧚
ฎᏒ೰ผ (2011/05) ࠬ࡯ࡄ࡯ࠨࠗࠛࡦࠬࡂࠗࠬࠢ࡯࡞⻠⟵. ৻ች㜞ᩞ㧚
⪭ว⍮⟤㧘ᛂ⿧ਁ༑ሶ㧘੹੗໪㓶㧘ㇹᐽᐢ㧘⷏᧛೰㧘෹᳗㓷Ꮖ㧘દ⼱ේ৻㧘᧻ᴛື㇢ (2012) ᄢဳ㘃ੱ₎ᖱႎ
ࡀ࠶࠻ࡢ࡯ࠢ㧔GAIN㧕╙ 3 ᦼߩᵴേߦะߌߡ. ߕ࡯ߛ߇߿ (2012/03/20, ฬฎደ)㧚
ㄞᄢ๺ (2011/09) ࠨ࡞ࠍㅢߒߡ↢߈‛ߩߟߥ߇ࠅࠍ⠨߃ࠆ㨪㊄⪇ጊፉߩ࠾ࡎࡦࠩ࡞ߩⷰኤ߆ࠄ㨪. ੩ㇺᄢቇ
㔤㐳㘃⎇ⓥᚲ᧲੩౏㐿⻠ᐳ ᣣᧄ⑼ቇᧂ᧪㙚, ᧲੩㧚
ㄞᄢ๺ (2012/03) ⎇ⓥ⠪ߩ઀੐. ᗲ⍮⋵›ጊᏒ┙ၔ᧲ਛቇᩞ‫ޟ‬௛ߊੱߩ⹤ࠍ⡞ߊળ‫ޠ‬, ›ጊᏒ㧚
ⴕേ␹⚻⎇ⓥㇱ㐷
ᕁ⠨⸒⺆ಽ㊁
㧨⎇ⓥ᭎ⷐ㧪
A) ࠴ࡦࡄࡦࠫ࡯ߩᲧセ⹺⍮⊒㆐⎇ⓥ
᧻ᴛື㇢㧘෹᳗㓷Ꮖ㧘ᨋ⟤㉿㧘દ᧛⍮ሶ㧘⿷┙ᐞ⏴㧔࿖㓙౒หవ┵⎇ⓥ࠮ࡦ࠲࡯㧕㧘᦯ㇱ⵨ሶ㧘ự↰ⓛ㧔ㅴൻᒻᘒ
ಽ㊁㧕㧘⷏᧛೰㧔♽⛔⊒↢ಽ㊁㧕㧘ධ㔕⚐ᴦ㧔࿖㓙౒หవ┵⎇ⓥ࠮ࡦ࠲࡯㧕㧘㋈ᧁ᮸ℂ㧘ችㇱ⾆ሶ㧘೨↰ౖᒾ㧘ᷰㆺ
ᦶ㊁㧘౗ሶ᣿ਭ㧘ᷰㆺ␽ᐔ㧔એ਄㧘ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕㧘ᾢፒᷡೣ㧘⪭ว㧔ᄢᐔ㧕⍮⟤㧘㜞ፉ෹ሶ㧘㈬
੗᦮ሶ㧔੩ㇺᄢ㧕
1 ⟲ 14 ୘૕ߩ࠴ࡦࡄࡦࠫ࡯ߩ߁ߜ‫ ߦ․ޔ‬10-11 ᱦߦߥࠆሶߤ߽ 3 ୘૕ࠍኻ⽎ߣߒߡ‫ޔ‬Ყセ⊒㆐⎇ⓥࠍ✚ว⊛ߦ߅
ߎߥߞߚ‫⹺ޕ‬⍮ᯏ⢻ߩ⸃ᨆߣߒߡ‫⺖࠲࡯ࡘࡇࡦࠦޔ‬㗴߿ኻ⽎ᠲ૞⺖㗴ߥߤฦ⒳⹺⍮⺖㗴ߦ߅޿ߡ࠴ࡦࡄࡦࠫ࡯ߩ߅
ߣߥ୘૕߿‫࠻ࡅޔ‬ᐜఽߣߩᲧセᬌ⸛ࠍ߅ߎߥߞߚ‫ޔߚ߹ޕ‬ቯᦼ⊛ߦ⣖߿り૕ฦㇱߩ⸘᷹߽߅ߎߥߞߡ޿ࠆ‫ޕ‬
B) ࠴ࡦࡄࡦࠫ࡯ߩ⍮ⷡ࡮⹺⍮⢻ജߩᲧセ⹺⍮⑼ቇ⊛⎇ⓥ
᧻ᴛື㇢㧘෹᳗㓷Ꮖ㧘દ᧛⍮ሶ㧘⿷┙ᐞ⏴㧘᦯ㇱ⵨ሶ㧘ධ㔕⚐ᴦ㧘⁚㊁ᢥᶈ㧘౗ሶፄ᣿㧘᧛᧻᣿Ⓞ㧘్࡝࡜㧘ᬀ↰
ᗐ㧘CD Dahl㧘M Kret㧘㜞ፉ෹ሶ㧘᧛੗ජኼሶ㧔₹Ꮉᄢ㧕㧘‐⼱ᥓ৻㧔ජ⪲ᄢ㧕
㧘ᓟ⮮๺ብ㧔੩ㇺᄢ㧕
㧘૒㊁᣿ੱ㧘↰
ਛ↱ᶈ㧘㈬੗ၮⴕ㧔એ਄‫ޔ‬ฬᎿᄢ㧕
࠴ࡦࡄࡦࠫ࡯ߣࡅ࠻ࠍኻ⽎ߦ‫⹺ޔ‬⍮࡮⸒⺆ᯏ⢻ߩᲧセ⎇ⓥࠍ⛮⛯ߒߡ߅ߎߥߞߚ‫ޕ‬ਥߣߒߡ‫ޔ‬1 ୘૕ߩ࠹ࠬ࠻႐
㕙ߢ‫ޔ‬ᢙ♽೉ቇ⠌‫ߣ⦡ޔ‬ᢥሼߩኻᔕ‫ޔ⼂⹺ߩ✢ⷞޔ‬㗻ߩ⍮ⷡ‫ޔ‬ᵈᗧ‫ⷡⷞޔ⼂⹺ࡦ࡯࠲ࡄޔ‬ត⚝‫ޔ⼂⹺࡯࡝ࠧ࠹ࠞޔ‬
‛ℂ⊛੐⽎ߩ⹺⼂‫ⷞޔ‬⡬ⷡ⛔ว‫ޔ‬ᖱേ⹺⍮‫ޔ‬ㆇേ⍮ⷡ‫ⴕޔ‬േߩห⺞ߥߤߩ⎇ⓥࠍ߅ߎߥߞߚ‫ޕ‬
C) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆࠕࠗ࠻࡜࠶ࠞ࡯ࠍ↪޿ߚⷞ✢⸘᷹
෹᳗㓷Ꮖ㧘⁚㊁ᢥᶈ㧘౗ሶፄ᣿㧘ᬀ↰ᗐ
࠴ࡦࡄࡦࠫ࡯ߣࡅ࠻ࠍኻ⽎ߦ‫ޔ‬㕖᜔᧤ဳߩࠕࠗ࠻࡜࠶ࠞ࡯ࠍ↪޿ߡ‫ޔ‬ฦ⒳ߩⷞⷡೝỗឭ␜ᤨ߿⺖㗴ㆀⴕᤨߩⷞ✢
ߩ⸘᷹ࠍ߅ߎߥߞߚ‫ޕ‬
D) ㊁↢࠴ࡦࡄࡦࠫ࡯ߩ㆏ౕ૶↪ߣᢥൻ⊛ᄌ⇣ߣ᫪ᨋౣ↢
᧻ᴛື㇢㧘ᨋ⟤㉿㧘ᄢᯅጪ㧘C Martin㧘ጊ⿧⸒㧔੩ㇺᄢ㧕㧘᫪᧛ᚑ᮸㧘⮮Ỉ㆏ሶ㧔એ਄㧘㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯㧕㧘
ጊᧄ⌀਽㧔ࡏࡁࡏ⎇ⓥㇱ㐷㧕㧘T Humle㧔ࠤࡦ࠻ᄢ㧕
㧘D Biro㧔ࠝ࠶ࠢࠬࡈࠜ࡯࠼ᄢ㧕㧘C Sousa㧔࡝ࠬࡏࡦᣂᄢ㧕㧘K Koops
㧔ࠤࡦࡉ࡝࠶ࠫᄢ㧕㧘K Hockings㧔࡝ࠬࡏࡦᣂᄢ㧕㧘S Carvalho㧔ࠤࡦࡉ࡝࠶ࠫᄢ㧕㧘N Granier㧔࡝ࠛ࡯ࠫࡘᄢ㧕㧘L Martinez
㧔᪸⧎ᅚሶᄢ㧕㧘AG Soumah (IREB)㧘T Tagbino㧔ࠞࡦࠞࡦᄢ㧕
⷏ࠕࡈ࡝ࠞ࡮ࠡ࠾ࠕߩࡏ࠶࠰࠙ߣ‫᧲ޔ‬㓞ߩ࠾ࡦࡃጊߣࠦ࡯࠻ࠫࡏࡢ࡯࡞㗔ౝ‫⷏ޔ‬㓞ߩ࠺ࠖࠛࠤߩ᫪‫ޔ‬ධ㓞ߩ࡝ࡌ
࡝ࠕ㗔ౝߢ‫ޔ‬㊁↢࠴ࡦࡄࡦࠫ࡯ߩⴕേߣ↢ᘒࠍ⺞ᩏߒ‫⸥ޔ‬㍳ߩ⸃ᨆࠍ߅ߎߥߞߚ‫ߩ✛ޟޔߚ߹ޕ‬࿁ᑈ‫ߣޠ‬๭߱᫪ᨋ
ౣ↢⎇ⓥࠍ⹜ߺ‫᧲߁ⷒࠍᧁ⧣ޔ‬ደࠍ⸳⟎ߔࠆᵴേࠍ⛮⛯ߒߚ‫ޕ‬
E) 㘺⢒㔤㐳㘃ߩⅣႺࠛࡦ࡝࠶࠴ࡔࡦ࠻
෹᳗㓷Ꮖ㧘᧻ᴛື㇢㧘ᨋ⟤㉿㧘ᾢፒᷡೣ㧘⪭ว㧔ᄢᐔ㧕⍮⟤㧘ዊୖආବ㧘ጊ᪸⵨⟤㧘ᰞᐸ㓁ሶ㧘Ꮉ਄ᷡᢥ㧔⡛ᔃᅚ
ሶᄢ㧕㧘㋈ᧁ᮸ℂ㧘೨↰ౖᒾ㧘ᷰㆺ␽ᐔ㧔એ਄‫ੱޔ‬㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕
േ‛⑔␩ߩ┙႐߆ࠄⅣႺࠛࡦ࡝࠶࠴ࡔࡦ࠻ߦ㑐ߔࠆ⎇ⓥࠍ߅ߎߥߞߚ‫ޕ‬3 ᰴర᭴▽‛ߩዉ౉߿ᬀ᮸ߩലᨐߩ⹏ଔ‫ޔ‬
ⷞⷡೝỗߩ๒␜ߦࠃࠆࠬ࠻࡟ࠬߩૐᷫ‫ޔ‬୘೎㘺⢒୘૕ߦኻߔࠆേ↹ೝỗឭ␜ߩലᨐ‫⹺ޔ‬⍮ታ㛎߇࠴ࡦࡄࡦࠫ࡯ߩⴕ
േߦ෸߷ߔᓇ㗀ߩ⹏ଔ‫ߩᧁޔ‬ᨑ߿⨲ߥߤࠍ૶ߞߚ⇣Ᏹⴕേߩૐᷫ‫ߩ↪࠻ࡦࡔ࠴࠶࡝ࡦࠛޔ‬ㆆౕߩዉ౉ߥߤߩ⎇ⓥࠍ
߅ߎߥߞߚ‫ޕ‬
㧙㧙
F) ฦ⒳㔤㐳㘃ߩ⹺⍮⊒㆐
෹᳗㓷Ꮖ㧘᧻ᴛື㇢㧘દ᧛⍮ሶ㧔Ყセ⹺⍮⊒㆐㧕㧘᦯ㇱ⵨ሶ㧘౗ሶፄ᣿㧘ᛂ⿧ਁ༑ሶ㧘⊕੗ㅀ㧔ᣂẟᄢ㧕㧘⮮↰๺↢㧘
ᷰㄝഃᄥ㧔એ਄㧘੩ㇺᄢ㧕㧘᧛੗ජኼሶ㧔₹Ꮉᄢ㧕
㧘⛚↰ବ๺㧘⑔቞ᦶ㧘ጊ↰ାብ㧘ᧁ᧛ᄐሶ㧘ዊ⷏స਽㧔એ਄㧘㜞
⍮⋵ߩ޿ߜേ‛౏࿦㧕㧘቟⮮ኼᐽ㧔ᘮᔕᄢ㧕㧘Ꮉ਄ᢥੱ㧔᧲੩ᄢ㧕㧘ጯᧄஜ㧔⡛ᔃᅚሶᄢ㧕
ࠕࠫ࡞࠹࠽ࠟࠩ࡞ߩᐜఽ‫ߩ࡞ࠩࠢࠞࡑޔ‬ᐜఽ‫߮ࠃ߅ޔ‬ᣂ਎⇇ࠩ࡞ฦ⒳ᚑ૕ࠍኻ⽎ߦ‫ޔ‬⒳‫⹺ߩޘ‬⍮⢻ജߣߘߩ⊒㆐
ߦߟ޿ߡᬌ⸛ࠍ߅ߎߥߞߚ‫ޔߦࠄߐޕ‬㜞⍮⋵ߩ޿ߜേ‛౏࿦ߦ߅޿ߡੑෆᕈ෺↢ఽߩ࠴ࡦࡄࡦࠫ࡯ߩⴕേ⊒㆐ࠍ❑
ᢿ⊛ߦⷰኤߒߡ޿ࠆ‫ޕ‬
G) ࡅ࠻ߩሶߤ߽ߩ⹺⍮⊒㆐
ᨋ⟤㉿㧘᦯ㇱ⵨ሶ
›ጊᏒߩᔃり㓚ኂఽ࠺ࠗࠨ࡯ࡆࠬ࠮ࡦ࠲࡯‫ߔ߽ߔߎޟ‬࿦‫⥄ޔߢޠ‬㐽∝‫ޔ∝ࡦ࠙࠳ޔ‬ᐢ᳢ᕈ⊒㆐㓚ኂߥߤ㕖ቯဳ⊒
㆐ఽߩࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦⴕേߩ⊒㆐ߦߟ޿ߡ‫ޔ‬ෳਈⷰኤ⎇ⓥࠍ߅ߎߥߞߚ‫ޕ‬
H) േ‛࿦ߩ࠴ࡦࡄࡦࠫ࡯ߩ⍮ᕈߩ⎇ⓥ
⿷┙ᐞ⏴㧘ᰞᐸ㓁ሶ㧘᧻ᴛື㇢
ฬฎደᏒߩ᧲ጊേ‛࿦ߩ࠴ࡦࡄࡦࠫ࡯1 ⟲ 6 ୘૕ࠍኻ⽎ߦ‫ޔ‬ᣂ⸳ߐࠇߚደᄖㆇേ႐ߢߩ␠ળⴕേࠍⷰኤ⸥㍳ߒߚ‫ޕ‬
߹ߚ‫ߣޠࡏ࡜ࡦࡄޟ‬ฬߠߌࠄࠇߚࡉ࡯ࠬߦ߅޿ߡ‫ޔ‬㆏ౕ૶↪ߣࠦࡦࡇࡘ࡯࠲⺖㗴ߩ 2 ߟߩ஥㕙߆ࠄ⍮ᕈߩ⎇ⓥࠍ߅
ߎߥߞߚ‫ޕ‬
I) ࠴ࡦࡄࡦࠫ࡯2 ୘૕႐㕙ߦ߅ߌࠆ␠ળ⊛⍮ᕈߩ⎇ⓥ
᦯ㇱ⵨ሶ㧘C Martin
࠴ࡦࡄࡦࠫ࡯2 ୘૕ࠍኻ⽎ߣߒ‫ⴕߩ࡯ࠫࡦࡄࡦ࠴ޔ‬േ߇ઁ⠪ߦᓇ㗀ߐࠇࠆ߆ߤ߁߆ࠍᬌ⸛ߒߚ‫ޕ‬2 ߟߩࡕ࠾࠲࡯
ࠍㅢߓߡ 2 ୘૕ߦ৻ㅪߩ⺖㗴ࠍ߅ߎߥࠊߖࠆߣ‫⋧ߪ࡯ࠫࡦࡄࡦ࠴ޔ‬ᚻߩⴕേࠍ⷗ߡ⥄ಽߩⴕേࠍ⺞ᢛߒߚ‫ࠄࠇߎޕ‬
ߩታ㛎߆ࠄ‫ઁࠆߌ߅ߦ࡯ࠫࡦࡄࡦ࠴ޔ‬⠪ℂ⸃ߩ৻㕙߇␜ߐࠇߚ‫ޕ‬
J) 㟗㘃ߣᄢဳ㘃ੱ₎ߩᲧセ⹺⍮⎇ⓥ
෹᳗㓷Ꮖ㧘᧛ጊ⟤Ⓞ㧘᫪㒋ආㅢ㧔㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯㧕㧘ਛේผ↢㧔Ᏹ⏥ᄢ㧕㧘ᢧ⮮⼾㧘਄㊁෹㚅㧘␹↰ᐘม㧘ศ
੗⺈㧘㒙ਭᩮ㓶৻㇢㧘ᣣ⊓ᒄ㧘␲৻⺈㧔એ਄‫ޔ‬ฬฎደ᷼᳓ᣖ㙚㧕
ฬฎደ᷼᳓ᣖ㙚ߣߩ౒ห⎇ⓥߣߒߡ‫ޔ‬㟗㘃ߩ⹺⍮⎇ⓥࠍㅴ߼ߡ޿ࠆ‫⹺ⷡⷞࠆߌ߅ߦࠞ࡞ࠗޔߦߊߣޕ‬⍮‫ࡦࠗࠨޔ‬
ߩℂ⸃‫⊛ⷡⷞޔ‬୘૕⼂೎ߥߤࠍᄢဳ㘃ੱ₎ߣߩᲧセ⎇ⓥߣߒߡㅴ߼ߡ޿ࠆ‫ߩઁߩߘޔߚ߹ޕ‬᳓ᣖ㙚ߣ߽ㅪ៤ߒߡ⎇
ⓥࠍㅴ߼ߡ޿ࠆ‫ޕ‬
K) ᄢဳ㘃ੱ₎ߩᲧセ⹺⍮⎇ⓥ
᧻ᴛື㇢㧘෹᳗㓷Ꮖ㧘ᨋ⟤㉿㧘ጊ᪸⵨⟤㧘ᾢፒᷡೣ㧘ᐘፉม㇢㧘ਭ਎Ớሶ㧘㊄᫪ᦺሶ㧔એ਄‫ޔ‬㊁↢േ‛⎇ⓥ࠮ࡦ࠲
࡯㧕ጊፒᓀᄐ㧔᧲੩ㄘᎿᄢ㧕㧘S Weide㧔ࡗࡗࠨࡦࠨࡃ⽷࿅㧕㧘HA Abdul㧔ࡑ࡟࡯ࠪࠕ࡮ࠨࡃᄢ㧕㧘D Sabapathy㧔ࠝ࡜
ࡦ࠙࡯࠲ࡦፉ⽷࿅㧕㧘D Baskaran㧔ࡊ࡜࠙ࡃࡦ࠺ࠖࡦࠣ⽷࿅㧕㧘M Mansor㧔ࡑ࡟࡯ࠪࠕ⑼ቇᄢቇ㧕
ࡑ࡟࡯ࠪࠕߩࠨࡃᎺߢ㊁↢ࠝ࡜ࡦ࠙࡯࠲ࡦߩ↢ᘒߣⴕേߩ⺞ᩏࠍ߅ߎߥߞߚ‫࡯࡟ࡑޔߚ߹ޕ‬ඨፉߩ㘺⢒ࠝ࡜ࡦ࠙
࡯࠲ࡦߩⅣႺࠛࡦ࡝࠶࠴ࡔࡦ࠻ߣ‫ࠍࡦ࠲࡯࠙ࡦ࡜ࠝޔ‬㊁↢ᓳᏫߐߖࠆ⹜ߺࠍ߅ߎߥߞߡ޿ࠆ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
Dahl CD, Logothetis NK, Bülthoff HH, Wallraven C (2011) Second-order relational manipulations affect both humans
and monkeys. PLoS One 6(10):e25793.
2) Hattori Y, Tomonaga M, Fujita K (2011) Chimpanzees (Pan troglodytes) show more understanding of human attentional
states when they request food in the experimenter’s hand than on the table. Interaction Studies 12:418–429.
3) Inoue S, Matsuzawa T (2011) Correlation between menstrual cycle and cognitive performance in a chimpanzee (Pan
troglodytes). Journal of Comparative Psychology 125:104-110.
4) Kaneko T, Tomonaga M (2011) The perception of self-agency in chimpanzees (Pan troglodytes). Proceedings of the
Royal Society Series B 278:3694-3702. doi:10.1098/rspb.2011.0611.
5) Kano F, Tomonaga M (2011) Species difference in the timing of gaze movement between chimpanzees and humans.
Animal Cognition 14:879-892.
6) Kano F, Tomonaga M (2011) Perceptual mechanism underlying gaze guidance in chimpanzees and humans. Animal
Cognition 14:377-386.
7) Kano F, Hirata S, Call J, Tomonaga M (2011) The visual strategy specific to humans among hominids: A study using
the gap-overlap paradigm. Vision Research 51:2348-2355.
8) Kawakami K, Kawakami F, Tomonaga M, Kishimoto T, Minami T, Takai-Kawakami K (2011) Origins of a theory of
mind. Infant Behavior & Development 34:264-269.
9) Ludwig VU, Adachi I, Matsuzawa T (2011) Visuoauditory mappings betweenhigh luminance and high pitch are shared
by chimpanzees (Pan troglodytes) and humans. PNAS 108(51):20661-20665.
10) Martin CF, Biro D, Matsuzawa T (2011) Chimpanzees’ use of conspecific cues in matching-to-sample tasks: public
information use in a fully automated testing environment. Animal Cognition 14:893-902.
㧙㧙
11) Morimura N, Idani G, Matsuzawa T (2011) The first chimpanzee sanctuary in Japan: an attempt to care for the “surplus”
of biomedical research. American Journal of Primatology 73:226–232.
12) Murai C, Tanaka M, Tomonaga M, Sakagami M (2011) Long-term visual recognition of familiar persons, peers, and
places by young monkeys (Macaca fuscata). Developmental Psychobiology 53:732-737. DOI: 10.1002/dev.20548.
13) Ogura T (2011) Contrafreeloading and the value of control over visual stimuli in Japanese macaques (Macaca fuscata).
Animal Cognition 14(3):427-431.
14) Ohashi G, Matsuzawa T (2011) Deactivation of snares by wild chimpanzees. Primates 52:1-5.
15) Saito A, Hayashi M, Ueno A, Takeshita H (2011) Orientation-indifferent representation in children’s drawings. Japanese
Psychological Research 53:379-390.
16) Sakai T, Mikami A, Tomonaga M, Matsui M, Suzuki J, Hamada Y, Tanaka M, Miyabe-Nishiwaki T, Makishima H,
Nakatsukasa M, Matsuzawa T (2011) Differential prefrontal white matter development in chimpanzees and humans.
Current Biology 21:1397-1402. doi:10.1016/j.cub.2011.07.019.
17) Sugawara T, Go Y, Udono T, Morimura N, Tomonaga M, Hirai H, Imai H (2011) Diversification of bitter taste receptor
gene family in western chimpanzees. Molecular Biology and Evolution 28:921-931.
18) Yamanashi Y, Hayashi M (2011) Assessing the effects of cognitive experiments on the welfare of captive chimpanzees
(Pan troglodytes) by direct comparison of activity budget between wild and captive chimpanzees. American Journal of
Primatology 73:1231-1238.
19) Carvalho S, Biro D, Cunha E, Hockings K, McGrew W, Richmond BG, Matsuzawa T (2012) Chimpanzee carrying
behaviour and the origins of human bipedality. Current Biology 22(6):180-181.
20) Goto K, Imura T, Tomonaga M (2012) Perception of emergent configurations in humans (Homo sapiens) and
chimpanzees (Pan troglodytes). Journal of Experimental Psychology: Animal Behavioral Processes 38:125-138.
21) Hattori Y, Leimgruber K, Fujita K, de Waal FBM (2012) Food-related tolerance in capuchin monkeys (Cebus apella)
varies with knowledge of the partner's previous food-consumption. Behaviour 149:171-185.
22) Hockings K, Anderson J, Matsuzawa T (2012) Socioecological adaptations in chimpanzees, Pan troglodytes verus, in
habiting anthropogenically impacted habitat. Animal Behaviour 83:801-810.
23) Koops K, McGrew W, Matsuzawa T, Knapp L (2012) Terrestrial nest-building by wild chimpanzees (Pan troglodytes):
Implications for the tree-to-ground sleep transition in early hominins. American Journal of Physical Anthropology,
March 28, 2012,DOI:10.1002/ajpa.22056.
24) Koops K, McGrew W, de Vries H, Matsuzawa T (2012) Nest-building by chimpanzees (Pan troglodytes) at Seringbara,
Nimba Mountains: Antepredation, thermoregulation, and antivector hypotheses. International Journal of Primatology
33:356-380.
25) Ogura T, Matsuzawa T (2012) Visual preference assessment and behavioural management of single-caged Japanese
macaques (Macaca fuscata) by movie presentation. Journal of Applied Animal Welfare Science 15(2):101-112.
26) Tsutsumi S, Ushitani T, Tomonaga M, Fujita K (2012) Infant monkeys’ concept of animacy: The role of eyes and
fluffiness. Primates 53:113-119. DOI 10.1007/s10329-011-0289-8.
✚⺑
1)
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5)
ႎ๔
1)
ᨋ⟤㉿ (2011) ࠝ࡜ࡦ࠙࡯࠲ࡦߩߔ߻ፉߢ. ⊒㆐ 127:94-102.
ዊୖආବ㧘ጊፒᓀᄐ㧘ጊ᪸⵨⟤㧘ਃኅ⹞❱ (2011) േ‛⑔␩⎇ⓥߩዷ㐿㨪േ‛ᔃℂቇળ⥄↱㓸ળߩ⼏⺰߆ࠄ. േ
‛ᔃℂቇ⎇ⓥ 61(1):115-124.
෹᳗㓷Ꮖ㧘࡙࡮࡝࡜ (2011) り૕ߢ⍮ⷡߔࠆ‫ޔ‬り૕ࠍ⍮ⷡߔࠆ㧦࠴ࡦࡄࡦࠫ࡯ߩ⹺⍮ߦ߅ߌࠆり૕ߩᓎഀ. Clinical
Neuroscience 29:876-880.
Kano F, Yamanashi Y, Tomonaga M (2012) Emotions as an intervening variable in understanding the cognitive and
social complexity and well-being of chimpanzees. Psychologia 55:9-20.
᧻ᴛື㇢ (2012) ‫⊒߁޿ߣޠࡊ࡯࡞ࠣ࠻࠙ࠕޟ‬ᗐ㧦ࡏࡁࡏ‫ੱࠄ߆࡜࡝ࠧޔࡦ࠲࡯࠙ࡦ࡜ࠝޔ‬㑆ࠍ⠨߃ࠆ. ⊒㆐
129:97-104.
↰ፉ⍮ਯ㧘⼱ญ᥍㚅㧘ጊ᪸⵨⟤㧘ᄢ⼱ᵗ੺㧘ዊᎹ⹞ਫ㧘ᣧᎹථᔒ㧘ᧄㇹፋ (2011) ╙ 23 ࿁࿖㓙㔤㐳㘃ቇળ Student
Affairs Workshop ࠍ⚳߃ߡ. 㔤㐳㘃⎇ⓥ 27:57-62.
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
2)
3)
4)
5)
Hayashi M, Inoue-Nakamura N (2011) From handling stones and nuts to tool-use. (The chimpanzees of Bossou and
Nimba) (ed. Matsuzawa T, Humle T, Sugiyama Y) p.175-182 Springer, Tokyo.
Hirata S, Hayashi M (2011) The emergence of stone-too use in captive chimpanzees. (The chimpanzees of Bossou and
Nimba) (ed. Matsuzawa T, Humle T, Sugiyama Y) p.183-190 Springer, Tokyo.
Matsuzawa T (2011) Education by master-apprenticeship. (The Chimpanzees of Bossou and Nimba) (ed. Matsuzawa T,
et al.) p.201-208 Tokyo: Springer.
Matsuzawa T (2011) Field experiments of tool-use. (The Chimpanzees of Bossou and Nimba) (ed. Matsuzawa T, et al.)
p.157-164 Tokyo: Springer.
Matsuzawa T (2011) Log doll: pretence in wild chimpanzees. (The Chimpanzees of Bossou and Nimba.) (ed.
Matsuzawa T, et al.) p.131-135 Tokyo: Springer.
㧙㧙
6)
7)
8)
9)
Matsuzawa T (2011) Stone tools for nut-cracking. (The Chimpanzees of Bossou and Nimba.) (ed. Matsuzawa T, et al.)
p.73-83 Tokyo: Springer.
Matsuzawa T, Humle T (2011) Bossou: 33 Years. (The Chimpanzees of Bossou and Nimba) (ed. Matsuzawa T, et al.)
p.3-10 Tokyo: Springer.
Matsuzawa T, Ohashi G, Humle T, Granier N, Kourouma M, Soumah AG (2011) Green Corridor Project: Planting in the
savanna between Bossouand Nimba. (The Chimpanzees of Bossouand Nimba) (ed. Matsuzawa T, et al.) p.361-370
Tokyo: Springer.
Yamamoto S, Yamakoshi G, Humle T, Matsuzawa T (2011) Ant fishing in trees: invention and modification of a new
tool-use behavior. (The Chimpanzees of Bossou and Nimba) (ed. Matsuzawa T, et al.) p.123-130 Tokyo: Springer.
✬㓸
1)
Matsuzawa T, Humle T, Sugiyama Y (2011) The Chimpanzees of Bossou and Nimba. p.465 Tokyo, Springer.
ߘߩઁߩၫ╩
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14)
ᨋ⟤㉿ (2011) ࠝ࡜ࡦ࠙࡯࠲ࡦࠍ᫪ߦᏫߔ. ⑼ቇ, 81 p.352-353㧚
ᨋ⟤㉿ (2011) ࠴ࡦࡄࡦࠫ࡯⎇ⓥ⠪߆ࠄߺߚࡏࡁࡏ. ⑼ቇ, 81 p.1126-1127㧚
᧻ᴛື㇢ (2011) ࠕ࠙࠻ࠣ࡞࡯ࡊߣ޿߁⊒ᗐ. ‫⑼ޟ‬ቇ‫ޠ‬12 ᦬ภ㧔╙ 120 ࿁㧕p.1228-1229㧚
᧻ᴛື㇢ (2011) ㊁↢ࡑ࠙ࡦ࠹ࡦࠧ࡝࡜ߩ࿖߆ࠄ. ‫⑼ޟ‬ቇ‫ޠ‬10 ᦬ภ㧔╙ 118 ࿁㧕p.1000-1001㧚
᧻ᴛື㇢㧘ࠢ࡝ࠬ࡮ࡑ࡯࠴ࡦ㧘࠼࡜࡮ࡆࡠ (2011) ⷗ߡ߹ߨࠆ. ‫⑼ޟ‬ቇ‫ޠ‬9 ᦬ภ㧔╙ 117 ࿁㧕p.870-871㧚
᧻ᴛື㇢ (2011) ࡏ࡞ࡀࠝߩ᫪߆ࠄ. ‫⑼ޟ‬ቇ‫ޠ‬8 ᦬ภ㧔╙ 116 ࿁㧕p.742-743㧚
᧛ጊ⟤Ⓞ㧘ᣧ㊁޽ߠߐ㧘෹᳗㓷Ꮖ㧘੹㊁ᤩ༹㧘਄㊁෹㚅㧘ᢧ⮮⼾㧘◉ේᱜౖ (2011) ࠗ࡞ࠞߩᕈᩰࠍㆮવሶ߆
ࠄ⍮ࠆ. ാ㝼, 54 p.11-14㧚
෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯‫ߦࠞ࡞ࠗޔ‬ળ߁. ാ㝼, 54 p.2-10㧚
෹᳗㓷Ꮖ (2011) ߰ߚߏߩߜ߮ߞߎ࠴ࡦࡄࡦࠫ࡯㧔ߜ߮ߞߎ࠴ࡦࡄࡦࠫ࡯ߣખ㑆ߚߜ㧔╙ 114 ࿁㧕㧕. ⑼ቇ, 81
p.516-517㧚
෹᳗㓷Ꮖ㧘౗ሶፄ᣿ (2011) ਎⇇ߦ௛߈߆ߌࠆ‫ޠߒߚࠊޟ‬㧔ߜ߮ߞߎ࠴ࡦࡄࡦࠫ࡯ߣખ㑆ߚߜ㧔╙ 115 ࿁㧕㧕.
⑼ቇ, 81 p.516-517㧚
ᨋ⟤㉿ (2012) ࠴ࡦࡄࡦࠫ࡯ߣߣ߽ߦ㧔ㅪタ㧦⎇ⓥ⠪ߦߥࠆ㧍╙ 37 ࿁㧕. ੩ㇺᄢቇᅚᕈ⎇ⓥ⠪ᡰេ࠮ࡦ࠲࡯
News Letter ߚߜ߫ߥ, 44 p.4㧚
⮮Ỉ㆏ሶ㧘᧻ᴛື㇢ (2012) ㊁↢࠴ࡦࡄࡦࠫ࡯ߩ಴↥. ⑼ቇ, 82 p.264-265㧚
᧻ᴛື㇢ (2012) ੩ㇺᄢቇࡉ࡯࠲ࡦ෹ᅢࡊࡠࠣ࡜ࡓ㧦ߘߩᱧผ⊛ዷᦸ. ࡅࡑ࡜ࡗቇ⹹, 13 ภ p.233-240㧚
෹᳗㓷Ꮖ (2012) ᾢᧄࠨࡦࠢ࠴ࡘࠕ࡝ߦࠃ߁ߎߘ㧔ߜ߮ߞߎ࠴ࡦࡄࡦࠫ࡯ߣખ㑆ߚߜ㧔╙ 121 ࿁㧕㧕. ⑼ቇ, 82
p.38-39㧚
ቇળ⊒⴫
1)
Chin H, Tomonaga M, Nakajima S, Uwano Y, Ogura S (2011) Study of using self-view images in the bottlenose dolphin
(Tursiops truncatus). The 19th Biennial Conference on the Biology of Marine Mammals (2011/11/27-12/02, Tampa,
Florida, USA).
2) Dahl CD, Rasch MJ, Tomonaga M, Adachi I (2012) Nature or Nurture in Face Perception? Looking Within,
Interdisciplinary Approaches to Consciousness. International Conference, National Institute of Advanced Studies
(2012/01/05-07, Bangalore India).
3) Dahl CD (2011) Evaluating factors of innate and experience-based contributions to chimpanzee face perception. The
75th Annual Convention of the Japanese Psychological Association, Nihon University.Workshop "Integrative
approaches toward holistic processing of faces" with Olivier Pascalis and Goedele van Belle (2011/09/15-17, Tokyo,
Japan).
4) Kret ME, Kano F, Tomonaga M, Matsuzawa T (2011) Pupil dilation in response to increasing pupils of male
conspecifics. ᣣᧄ㔤㐳㘃ቇળ╙ 27 ࿁ቇⴚ㓸ળ (2011/07/16-18, ›ጊ࿖㓙ⷰశ࠮ࡦ࠲࡯࡮ࡈࡠࠗ࠺).
5) Ochiai-Ohira T, Uchikoshi M, Imai H, Go Y, Nishimura T, Idani G, Matsuzawa T (2011) Activity of the Great Ape
Information Network (GAIN): a web site promoting animal welfare. International Conference on Environmental
Enrichment (2011/08/14-19, Benson Hotel, Oregon, USA).
6) Ogura T (2011) Visual enrichment using Internet video sharing for Japanese monkeys. International Conference on
Environmental Enrichment 2011 (2011/08/14-19, Benson Hotel, Oregon, USA).
7) Yamanashi Y, Hayashi M (2011) Wild captive comparisons of the behaviors of chimpanzees (Pan troglodytes).
International Conference of Environmental Enrichment (2011/08/14-19, Benson Hotel, Oregon, USA).
8) ⿷┙ᐞ⏴㧘෹᳗㓷Ꮖ㧘᧻ᴛື㇢ (2011) ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ㗻⍮ⷡ᭽ᑼߩ⊒㆐. ᣣᧄ⿒ߜ߾ࠎቇળ╙ 11 ࿁ቇⴚ
㓸ળ (2011/05/07-08, ਛㇱቇ㒮ᄢቇ)㧚
9) ⿷┙ᐞ⏴㧘෹᳗㓷Ꮖ㧘᧻ᴛື㇢ (2011) ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ㗻ో૕ಣℂߩ⊒㆐. ᣣᧄ㔤㐳㘃ቇળ╙ 27 ࿁ቇⴚ㓸
ળ (2011/07/16-18, ›ጊ࿖㓙ⷰశ࠮ࡦ࠲࡯࡮ࡈࡠࠗ࠺)㧚
10) 㒙଻஻ሶ㧘ችਯේ⌀↱㧘੹੗ᅕ㧘ጟᧄ౏ᓆ㧘㥲⮮ᷤ㧘ጊญ⾆ᄩ㧘੗Ꮉ⍮ሶ㧘ዊᎹඅ㧘㊁᧛⟵᣿㧘ችㇱ⾆ሶ㧘
᩶੗଻ሶ㧘⧎↰ାᒄ (2011) ࠴ࡦࡄࡦࠫ࡯ญ⣧↱᧪ߩࡒࡘ࡯࠲ࡦࠬ࡟ࡦࠨ⃿⩶᭽⚦⩶ߦ㑐ߔࠆ⎇ⓥ. 㢬⷗ᄢቇ
ᱤቇળ╙ 73 ࿁଀ળ (2011/06/25, 㢬⷗ᄢቇળ㙚ࡔࠗࡦࡎ࡯࡞)㧚
㧙㧙
11) ⑺ศ↱૫㧘ᐔᩙ᣿ታ㧘᳓㊁૫✜㉿㧘Ꮢ㊁ᖝሶ㧘᦭⾐⩿ᵤ⟤㧘ᷰㆺߺߥߺ㧘ᨋ⟤㉿ (2011) ‫⊒ߩ࡯ࠫࡦࡄࡦ࠴ޟ‬
㆐ߦ઻߁␠ળ㑐ଥߩᄌൻ㨪ᦨㄭធ୘૕ߩ❑ᢿ⊛⸃ᨆ㨪‫ޠ‬. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒ
േᬀ‛࿦)㧚
12) ቟⮮ኼᐽ㧘෹᳗㓷Ꮖ㧘⑔቞ᦶ㧘ጊ↰ାብ㧘ᧁ᧛ᄐሶ㧘⛚↰ବ๺ (2012) ࠴ࡦࡄࡦࠫ࡯ߩ߈ࠂ߁ߛ޿㑐ଥ㧙Ყセ
෺↢ఽቇߩ⹜ߺ (2). ╙ 26 ࿁ᣣᧄ෺↢ఽ⎇ⓥቇળቇⴚ⻠Ṷળ (2012/01/28, ᧲੩)㧚
13) ᦭⾐⩿ᵤ⟤㧘ጊ᪸⵨⟤㧘ᨋ⟤㉿ (2011) ‫ޟ‬ᡷୃᎿ੐߇࠴ࡦࡄࡦࠫ࡯ߩⴕേߦਈ߃ࠆᓇ㗀⹏ଔ‫ޠ‬. ╙ 14 ࿁ SAGA
ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
14) 㒸㚅⚐㧘෹᳗㓷Ꮖ㧘ਛፉቯᒾ㧘਄㊁෹㚅㧘ዊୖੳ (2011) ࡃࡦ࠼࠙ࠗ࡞ࠞߦ߅ߌࠆ⥄Ꮖᤋ௝ࠍ↪޿ߚ⥄Ꮖ⹺⍮
ߩᬌ⸛. Animal 2011㧔2011 ᐕᐲᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕേቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓▤ℂቇળ
วหᄢળ㧕 (2011/09/08-11, ᘮᔕᄢቇ)㧚
15) Dahl CD, Tomonaga M, Adachi I (2011) Evaluating factors of innate and experience-based contributions to chimpanzee
face perception? ᣣᧄ㔤㐳㘃ቇળ╙ 27 ࿁ቇⴚ㓸ળ (2011/07/16-18, ›ጊ࿖㓙ⷰశ࠮ࡦ࠲࡯࡮ࡈࡠࠗ࠺)㧚
16) ⮮᫪໑㧘ᨋ⟤㉿ (2011) ‫ࠆߌ߅ߦ࡞ࠩࠠࡑࠝޟ‬ⅣႺࠛࡦ࡝࠶࠴ࡔࡦ࠻ߣߘߩലᨐ‫ޠ‬. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ
࠙ࡓ (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
17) ᦯ㇱ⵨ሶ㧘෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆઁ⠪ߩⴕേߦኻߔࠆห⺞௑ะ. ᣣᧄ⿒ߜ߾ࠎቇળ╙ 11 ࿁ቇ
ⴚ㓸ળ (2011/05/07-08, ਛㇱቇ㒮ᄢቇ)㧚
18) ᦯ㇱ⵨ሶ㧘෹᳗㓷Ꮖ (2011) ⇣ߥࠆᗧ࿑ࠍᜬߟേ૞ߦኻߔࠆ࠴ࡦࡄࡦࠫ࡯ߩᮨ୮௑ะ㧦⷗ᧄวࠊߖ⺖㗴ࠍ↪޿
ߡ. Animal 2011㧔2011 ᐕᐲᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕേቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓▤ℂቇળวห
ᄢળ㧕 (2011/09/08-11, ᘮᔕᄢቇ)㧚
19) ᨋ⟤㉿ (2011) ⿒ߜ߾ࠎቇߩዷ㐿㧦Ყセ⹺⍮⊒㆐ߩⷞὐ߆ࠄ. ᣣᧄ⿒ߜ߾ࠎቇળ (2011/05/07, ਛㇱቇ㒮ᄢቇ)㧚
20) ᨋ⟤㉿㧘ᄢᯅጪ㧘ᩉ⥝㎽ (2011) ⟂ߦ߆߆ߞߚേ‛ߦኻߔࠆࡏࡁࡏߩⴕേ̆ࡢࡦࡃߢߩࠤ࡯ࠬ࡟ࡐ࡯࠻̆.
SAGA14 (2011/11/12, ᾢᧄᏒ)㧚
21) ᨋ⟤㉿㧘┻ਅ⑲ሶ (2011) ᄢဳ㘃ੱ₎ߣࡅ࠻ᐜఽߩⓍᧁߩᠲ૞ߦߺࠆ‛ℂ⊛ߥ․ᕈߩℂ⸃. ᣣᧄ㔤㐳㘃ቇળ╙
27 ࿁ቇⴚ㓸ળ (2011/07/17, ›ጊᏒ)㧚
22) Ꮢ㊁ᖝሶ㧘⮮᫪໑㧘ᧁ᧛రᄢ㧘⑔቞ᦶ㧘ዊ⷏స਽㧘ጊ↰ାብ㧘ᧁ᧛ᄐሶ㧘⛚↰ବ๺㧘෹᳗㓷Ꮖ (2011) ࠴ࡦࡄ
ࡦࠫ࡯ߩሶߤ߽ߩᚑ㐳ߦ઻߁␠ળ㑐ଥߩᄌൻ. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
23) ੹੗໪㓶㧘⪭ว㧔ᄢᐔ㧕⍮⟤㧘ㇹᐽᐢ㧘⷏᧛೰㧘દ⼱ේ৻㧘ᛂ⿧ਁ༑ሶ㧘᧻ᴛື㇢ (2011) NBRP‫ޟ‬GAIN㧔ᄢ
ဳ㘃ੱ₎ᖱႎࡀ࠶࠻ࡢ࡯ࠢ㧕‫ޠ‬㧦ᄢဳ㘃ੱ₎ᖱႎࡀ࠶࠻ࡢ࡯ࠢ㧔GAIN㧕ߩᵴേ⚫੺. ╙ 34 ࿁ᣣᧄಽሶ↢‛ቇ
ળᐕળ 2011 ᐕ․೎ડ↹ NBRP ዷ␜ (2011/12/13-16, ࡄࠪࡈࠖࠦᮮᵿ)㧚
24) ౗ሶፄ᣿㧘෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߣࡅ࠻ߦ߅ߌࠆⷞⷡ㧙ㆇേ㓐઻ᕈߩᄌൻߦኻߔࠆ᳇ߠ߈ߣ⥄േ⊛ߥ
ㆇേ⵬ᱜ. Animal 2011㧔2011 ᐕᐲᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕേቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓▤ℂቇ
ળวหᄢળ㧕 (2011/09/08-11, ᘮᔕᄢቇ)㧚
25) ౗ሶፄ᣿㧘෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߣࡅ࠻ߦ߅ߌࠆ⥄Ꮖߩ㓐ᗧㆇേߩ⍮ⷡ㧦ⷞⷡ㧙ㆇേ㓐઻ᕈߩᄌൻߦ
ኻߔࠆⷞ✢ⴕേߣ਄⢇ㆇേߩ⺞▵. ᣣᧄ⿒ߜ߾ࠎቇળ╙ 11 ࿁ቇⴚ㓸ળ (2011/05/07-08, ਛㇱቇ㒮ᄢቇ)㧚
26) ⁚㊁ᢥᶈ㧘Call J㧘෹᳗㓷Ꮖ (2011) ࡅ࠻⑼ 4 ዻߦ߅ߌࠆ㗻ߩ⷗ᣇ㧦Ყセࠕࠗ࡮࠻࡜࠶ࠠࡦࠣ⎇ⓥ. Animal 2011
㧔2011 ᐕᐲᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕേቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓▤ℂቇળวหᄢળ㧕
(2011/09/08-11, ᘮᔕᄢቇ)㧚
27) ⁚㊁ᢥᶈ㧘Call J㧘෹᳗㓷Ꮖ (2011) ࠕࠗ࡮࠻࡜࠶ࠠࡦࠣߩᚻᴺࠍ↪޿ߚࡅ࠻⑼ 4 ዻߦ߅ߌࠆⷞⷡᚢ⇛ߩᬌ⸛.
Animal 2011㧔2011 ᐕᐲᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕേቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓▤ℂቇળวหᄢ
ળ㧕 (2011/09/08-11, ᘮᔕᄢቇ)㧚
28) ⁚㊁ᢥᶈ㧘ᐔ↰⡡㧘Call J㧘෹᳗㓷Ꮖ (2011) ࡅ࠻⑼ 4 ዻߦ߅ߌࠆᲧセࠕࠗ࡮࠻࡜࠶ࠠࡦࠣ⎇ⓥ߆ࠄ᣿ࠄ߆ߦߥ
ߞߚࡅ࠻․᦭ߩⷞⷡᚢ⇛. ᣣᧄ㔤㐳㘃ቇળ╙ 27 ࿁ቇⴚ㓸ળ (2011/07/16-18, ›ጊ࿖㓙ⷰశ࠮ࡦ࠲࡯࡮ࡈࡠࠗ࠺)㧚
29) ⁚㊁ᢥᶈ㧘ᐔ↰⡡㧘෹᳗㓷Ꮖ (2011) 㘃ੱ₎ 3 ⒳ߣࡅ࠻ߦ߅ߌࠆࠡࡖ࠶ࡊലᨐ. ᣣᧄ⿒ߜ߾ࠎቇળ╙ 11 ࿁ቇⴚ
㓸ળ (2011/05/07-08, ਛㇱቇ㒮ᄢቇ)㧚
30) Kim Y, Lee W, Martinez L, Kim S, Choe JC, Adachi I, Tomonaga M (2011) Ape research project in Seoul Zoo, Korea.
SAGA (Support for African/Asian Great Apes) (2011/11/12-13, Kumamoto City Zoo and Botanical Garden, Japan)㧚
31) ᧁ᧛రᄢ㧘ᰞᐸ㓁ሶ㧘Ꮢ㊁ᖝሶ㧘ፉ↰߆ߥ߃㧘㋈ᧁஜᄥ㧘ᷰㆺߺߥߺ㧘ㄭ⮮⵨ᴦ㧘ᧁ᧛ᐘ৻㧘⿷┙ᐞ⏴ (2011)
‫᧲ޟ‬ጊേ‛࿦ߩ࠴ࡦࡄࡦࠫ࡯࠲ࡢ࡯೑↪⁁ᴫߩ⛮⛯⺞ᩏ‫ޠ‬. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄ
Ꮢേᬀ‛࿦)㧚
32) ችਯේ⌀↱㧘੹੗ᅕ㧘㥲⮮ᷤ㧘ጊญ⾆ᄩ㧘㒙଻஻ሶ㧘㊁᧛⟵᣿㧘᩶੗଻ሶ㧘⧎↰ାᒄ (2011) ࠴ࡦࡄࡦࠫ࡯ญ
⣧↱᧪ߩ࡟ࡦࠨ⃿⩶ߩᕈ⁁ߦ㑐ߔࠆ⎇ⓥ. ╙ 60 ࿁ᣣᧄญ⣧ⴡ↢ቇળ (2011/05/19-21, ජ⪲Ꮢᢥൻ੤ᵹࡊ࡜ࠩ)㧚
33) ᫪ߎߣߩ㧘ਛጊ߰߁ߎ㧘ᰞᐸ㓁ሶ㧘᧻ᴛື㇢ (2011) ‫⥄ޟ‬ὼቇࡐࠤ࠶࠻࠯ࡒ࠽࡯࡞ᵴേ⚫੺‫ޠ‬. ╙ 14 ࿁ SAGA
ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
34) ᫪᧛ᚑ᮸㧘਄ဈඳ੺㧘㇊㗇๺ઍ㧘ᐔ↰⡡㧘෹᳗㓷Ꮖ (2011) ᾢᧄࠨࡦࠢ࠴ࡘࠕ࡝ߩ⸳┙. ╙ 14 ࿁ SAGA ࠪࡦ
ࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
㧙㧙
35) ᧛੗ජኼሶ㧘෹᳗㓷Ꮖ (2011) ߎߞߜࠍ⷗ߡࠆ? ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆઁ⠪ߩᵈᗧ⁁ᘒߩ⹺⼂. Animal 2011㧔2011
ᐕᐲᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕേቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓▤ℂቇળวหᄢળ㧕 (2011/09/08-11,
ᘮᔕᄢቇ)㧚
36) ᄐ⋡ዅᅢ㧘㗇↰⋥ሶ㧘ᨋ⟤㉿ (2011) ‫ࠆߌ߅ߦ߁߷ࠎ߆޽ߩ࡞ࠩࡦࡎ࠾ޟ‬࿕ᒻ㘺ᢱᵞ޿ⴕേ₪ᓧߩㆊ⒟‫ޠ‬. 14
࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒേ‛࿦)㧚
37) ⪭ว㧔ᄢᐔ㧕⍮⟤㧘ᛂ⿧ਁ༑ሶ㧘੹੗໪㓶㧘ㇹᐽᐢ㧘⷏᧛೰㧘દ⼱ේ৻㧘᧻ᴛື㇢ (2011) ᄢဳ㘃ੱ₎ᖱႎࡀ
࠶࠻ࡢ࡯ࠢ (GAIN) ߩᵴേ㧦࠙ࠚࡉࠨࠗ࠻ߢߩᖱႎ▤ℂ. ᣣᧄ㔤㐳㘃ቇળ╙ 27 ࿁ቇⴚ㓸ળ (2011/07/16-18, ›
ጊ࿖㓙ⷰశ࠮ࡦ࠲࡯࡮ࡈࡠࠗ࠺)㧚
38) ⪭ว⍮⟤㧘ᛂ⿧ਁ༑ሶ㧘੹੗໪㓶㧘ㇹᐽᐢ㧘⷏᧛೰㧘દ⼱ේ৻㧘᧻ᴛື㇢ (2011) GAIN ࠙ࠚࡉࠨࠗ࠻ߩ⚫੺㧦
㘃ੱ₎ߦ㑐ߔࠆᖱႎ෼㓸. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
39) ዊୖආବ (2011) േ↹๒␜ߦࠃࠆࠤ࡯ࠫ㘺⢒࠾ࡎࡦࠩ࡞ߩ⇣Ᏹⴕേߩシᷫߣᣂᄸᕈ࡮ౝኈ࡮ᠲ૞ᕈߩലᨐ. ᣣ
ᧄ㔤㐳㘃ቇળ╙ 27 ࿁ᄢળ (2011/07/16-18, ᗲ⍮⋵›ጊ࿖㓙ⷰశ࠮ࡦ࠲࡯ࡈࡠࠗ࠺)㧚
40) ዊୖආବ (2011) േ↹๒␜ߦࠃࠆ㘺⢒࠾ࡎࡦࠩ࡞ߩ⇣Ᏹⴕേߩシᷫߣേ↹๒␜ⷐ࿃ߩᬌ⸛. Animal 2011㧦ᣣᧄ
േ‛ᔃℂቇળ㧔╙ 71 ࿁㧕࡮ᣣᧄേ‛ⴕേቇળ㧔╙ 30 ࿁㧕࡮ᔕ↪േ‛ⴕേቇળ㧛ᣣᧄኅ⇓▤ℂቇળ㧔2011 ᐕ
ᐲ㧕วหᄢળ (2011/09/08-11, ᧲੩ㇺᘮᙥ⟵Ⴖᄢቇਃ↰ࠠࡖࡦࡄࠬ)㧚
41) ዊୖආବ (2011) You Tube േ↹๒␜ߦࠃࠆ࠾ࡎࡦࠩ࡞ߩ⇣Ᏹⴕേߩシᷫ. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ
(2011/11, ᾢᧄ)㧚
42) ㈬੗᦮ሶ㧘᧻੗ਃᨑ㧘Ludize M㧘ਃ਄┨మ㧘ਛോ⌀ੱ㧘෹᳗㓷Ꮖ㧘㋈ᧁ᮸ℂ㧘ự↰ⓛ㧘↰ਛᱜਯ㧘ችㇱ㧙⷏⣁
⾆ሶ㧘Ꮞፉ⟤ᐘ㧘᧻ᴛື㇢ (2011) ࠴ࡦࡄࡦࠫ࡯ߩᄢ⣖⚵❱ߩ⊒㆐ㆊ⒟ߪࡅ࠻ߩ⣖ㅴൻࠍℂ⸃ߔࠆ਄ߢߩᣂߚ
ߥ⷗⼂ࠍਈ߃ࠆ. ╙ 65 ࿁ᣣᧄੱ㘃ቇળᄢળ (2011/11/04, ᴒ✽⋵┙ඳ‛㙚࡮⟤ⴚ㙚)㧚
43) ㈬੗᦮ሶ㧘᧻੗ਃᨑ㧘ਛോ⌀ੱ㧘෹᳗㓷Ꮖ㧘ਃ਄┨మ㧘㋈ᧁ᮸ℂ㧘ự↰ⓛ㧘↰ਛᱜਯ㧘ችㇱ⾆ሶ㧘Ꮞፉ⟤ᐘ㧘
᧻ᴛື㇢ (2011) ࠴ࡦࡄࡦࠫ࡯ߣࡅ࠻ߦ߅ߌࠆᄢ⣖ߩ⊒㆐ㆊ⒟㧦ࡅ࠻ߩ⣖ߩㅴൻ⊛ၮ⋚ߩℂ⸃ߦะߌߡ. ᣣᧄ
⿒ߜ߾ࠎቇળ╙ 11 ࿁ቇⴚ㓸ળ (2011/05/07-08, ਛㇱቇ㒮ᄢቇ)㧚
44) ᰞᐸ㓁ሶ㧘ᨋ⟤㉿ (2011) ‫ޟ‬㘺⢒ⅣႺߩᄌൻ߇྾⢇㤗∽ࠍᖚߞߚ࠴ࡦࡄࡦࠫ࡯㧔Pan troglodytes㧕ߩⴕേߦ߽
ߚࠄߔᓇ㗀ߣ੹ᓟߩ࡝ࡂࡆ࡝⸘↹෸߮⹏ଔᣇᴺߩᬌ⸛‫ޠ‬. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/16-18, ›ጊ)㧚
45) ᰞᐸ㓁ሶ㧘ᨋ⟤㉿ (2011) ‫ޟ‬⣄㜑Ἳࠍ⊒∝ߒߚ࠴ࡦࡄࡦࠫ࡯ߩ 5 ᐕ㑆ߣߎࠇ߆ࠄ‫ޠ‬. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ
࠙ࡓ (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
46) ᰞᐸ㓁ሶ㧘Ꮢ㊁ᖝሶ㧘ᧁ᧛రᄢ㧘ፉ↰߆ߥ߃㧘㋈ᧁஜᄥ㧘ᑝỈ㤗㉿㧘ㄭ⮮⵨ᴦ㧘ጊᧄశ㓁㧘᧻᧛⑲৻㧘⿷┙
ᐞ⏴ (2011) ‫‛ޟ‬ℂ⊛෸߮␠ળ⊛㘺⢒ⅣႺߩᄌൻ߇࠴ࡦࡄࡦࠫ࡯ߩⴕേߦ߽ߚࠄߔᓇ㗀‫ޠ‬. Animal 2011
(2011/09/08-11, ᧲੩)㧚
47) ፉ↰߆ߥ߃㧘ᰞᐸ㓁ሶ㧘Ꮢ㊁ᖝሶ㧘ᧁ᧛రᄢ㧘㋈ᧁஜᄥ㧘ᷰㆺߺߥߺ㧘ㄭ⮮␭ᴦ㧘ᧁ᧛ᐘ৻㧘⿷┙ᐞ⏴ (2011)
‫ޟ‬ዊဳዷ␜ᣉ⸳̌ࡄࡦ࡜ࡏ̍ߩ⹏ଔ㧙࠴ࡦࡄࡦࠫ࡯ߩ೑↪₸࡮ⴕേߩᄌൻࠍㅢߒߡ㧙‫ޠ‬. ╙ 14 ࿁ SAGA ࠪࡦ
ࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
48) ㋈ᧁஜᄥ㧘ᰞᐸ㓁ሶ㧘Ꮢ㊁ᖝሶ㧘ᧁ᧛రᄢ㧘ፉ↰߆ߥ߃㧘ᷰㆺߺߥߺ㧘ㄭ⮮⵨ᴦ㧘ᧁ᧛ᐘ৻㧘⿷┙ᐞ⏴ (2011)
‫᧲ޟ‬ጊേ‛࿦ߩ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ⹺⍮ታ㛎ෳട₸ߩᄌേ‫ޠ‬. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13,
ᾢᧄᏒേᬀ‛࿦)㧚
49) ෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ‫ߩޠ✢࠼࡯ࡇࠬޟ‬⍮ⷡ. ᣣᧄᔃℂቇળ╙ 75 ࿁ᄢળ (2011/09/15-17, ᣣ
ᧄᄢቇ)㧚
50) ෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ‫ߩޠ✢࠼࡯ࡇࠬޟ‬⍮ⷡ. ᣣᧄ㔤㐳㘃ቇળ╙ 27 ࿁ቇⴚ㓸ળ (2011/07/16-18,
›ጊ࿖㓙ⷰశ࠮ࡦ࠲࡯࡮ࡈࡠࠗ࠺)㧚
51) ෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ⥄Ꮖ㗻ߩ⍮ⷡ. Animal 2011㧔2011 ᐕᐲᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕ
േቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓▤ℂቇળวหᄢળ㧕 (2011/09/08-11, ᘮᔕᄢቇ)㧚
52) ෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ⥄Ꮖ㗻ߩ⍮ⷡ. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒേ
ᬀ‛࿦)㧚
53) ෹᳗㓷Ꮖ㧘౗ሶፄ᣿ (2011) ߐߞ߈૗ࠍㆬࠎߛߩ㧫࠴ࡦࡄࡦࠫ࡯ߪ⋥೨ߩ⥄ಽߩⴕേࠍ⸥ᙘߒߡ޿ࠆ߆. ᣣᧄ
⿒ߜ߾ࠎቇળ╙ 11 ࿁ቇⴚ㓸ળ (2011/05/07-08, ਛㇱቇ㒮ᄢቇ)㧚
54) ਄㊁෹㚅㧘ᢧ⮮⼾㧘૒⮮⌀ᄹ⟤㧘ේഞᰴ㇢㧘෹᳗㓷Ꮖ (2011) ࡃࡦ࠼࠙ࠗ࡞ࠞߦ߅ߌࠆኻ⒓ᕈߩᚑ┙ߦߟ޿ߡ.
Animal 2011㧔2011 ᐕᐲᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕേቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓▤ℂቇળวหᄢ
ળ㧕 (2011/09/08-11, ᘮᔕᄢቇ)㧚
55) ᬀ↰ᗐ㧘෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦࠃࠆ⦡ߣ࿑ᒻߩ⽎ᓽ⷗ᧄวࠊߖߩ㐳ᦼ଻ᜬ. Animal 2011㧔2011 ᐕᐲ
ᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕേቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓▤ℂቇળวหᄢળ㧕 (2011/09/08-11, ᘮ
ᔕᄢቇ)㧚
56) ᬀ↰ᗐ㧘෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦࠃࠆ⦡ߣ࿑ᒻߩ⽎ᓽ⷗ᧄวࠊߖߩ㐳ᦼ⸥ᙘ. ╙ 14 ࿁ SAGA ࠪࡦࡐ
ࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
57) ᬀ↰ᗐ㧘෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦࠃࠆ⦡ߣ࿑ᒻߩ⽎ᓽ⷗ᧄวࠊߖߩ㐳ᦼ଻ᜬ. ᣣᧄ㔤㐳㘃ቇળ╙ 27
࿁ቇⴚ㓸ળ (2011/07/16-18, ›ጊ࿖㓙ⷰశ࠮ࡦ࠲࡯࡮ࡈࡠࠗ࠺)㧚
㧙㧙
58) ᷰㆺߺߥߺ㧘ᰞᐸ㓁ሶ㧘Ꮢ㊁ᖝሶ㧘ᧁ᧛రᄢ㧘ፉ↰߆ߥ߃㧘㋈ᧁஜᄥ㧘ㄭ⮮⵨ᴦ㧘ᧁ᧛ᐘ৻㧘⿷┙ᐞ⏴ (2011)
‫᧲ޟ‬ጊേ‛࿦ߢߩ࠴ࡦࡄࡦࠫ࡯ߩ⍮ᕈዷ␜‫ޠ‬. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
59) ጊ ᪸ ⵨ ⟤ 㧘 ᨋ ⟤ ㉿ (2011) ㊁ ↢ ߩ ណ 㘩 ᤨ 㑆 ߪ ࠴ ࡦ ࡄ ࡦ ࠫ ࡯ ߩ ⇣ Ᏹ ⴕ േ ࠍ ᷫ ዋ ߐ ߖ ࠆ ߆ . Animal 2011
(2011/09/08-11, ᧲੩)㧚
60) ጊ᪸⵨⟤㧘᫪᧛ᚑ᮸㧘᫪⵨੺㧘ᨋ⟤㉿㧘㋈ᧁ᮸ℂ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆᲫਛࠦ࡞࠴࠱࡞᷹ቯ. ╙ 14
࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ (2011/11/12-13, ᾢᧄ)㧚
61) ࡙࡮࡝࡜㧘෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ⥄⊒⊛ߥห⺞ⴕേ㧔Spontaneous interpersonal synchrony in
chimpanzees㧕. Animal 2011㧔2011 ᐕᐲᣣᧄേ‛ᔃℂቇળ࡮ᣣᧄേ‛ⴕേቇળ࡮ᔕ↪േ‛ⴕേቇળ࡮ᣣᧄኅ⇓
▤ℂቇળวหᄢળ㧕 (2011/09/08-11, ᘮᔕᄢቇ)㧚
62) ࡙࡮࡝࡜㧘෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆห⺞ⴕേ㧙࠲࠶ࡇࡦࠣ⺖㗴ࠍ↪޿ߡ. ᐔᚑ 23 ᐕᐲ↢ℂ⎇⎇
ⓥળ (2011/10/06-07, ↢ℂቇ⎇ⓥᚲ)㧚
63) ࡙࡮࡝࡜㧘෹᳗㓷Ꮖ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆήᗧ࿑⊛ߥห⺞ⴕേߩታ㛎⊛ⷰኤ㧦࠲࠶ࡇࡦࠣࡄ࡜࠳ࠗࡓ
ࠍ↪޿ߡ. ᣣᧄ㔤㐳㘃ቇળ╙ 27 ࿁ቇⴚ㓸ળ (2011/07/16-18, ›ጊ࿖㓙ⷰశ࠮ࡦ࠲࡯࡮ࡈࡠࠗ࠺)㧚
64) Dahl CD (2012) Multi-disciplinary approach on face perception to disentangle developmental characteristics.
International Institute for Advanced Studies, "The Origin of Mind", 3rd Conference (2012/01/28-29, Kyoto).
65) ⪭ว⍮⟤㧘ᛂ⿧ਁ༑ሶ㧘੹੗໪㓶㧘ㇹᐽᐢ㧘⷏᧛೰㧘෹᳗㓷Ꮖ㧘દ⼱ේ৻㧘᧻ᴛື㇢ (2012) ᄢဳ㘃ੱ₎ᖱႎ
ࡀ࠶࠻ࡢ࡯ࠢ㧔GAIN㧕╙ 3 ᦼߩᵴേߦะߌߡ. ߕ࡯ߛ߇߿ (2012/03/20, ฬฎደ)㧚
66) ෹᳗㓷Ꮖ (2012) Looking for Myself ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ⥄Ꮖ㗻ߩ⍮ⷡ. ᣣᧄ⊒㆐ᔃℂቇળ╙ 23 ࿁ᄢળ
(2012/03/09-11, ฬฎደ࿖㓙ળ⼏႐)㧚
67) ෹᳗㓷Ꮖ (2012) ࠗ࡞ࠞ߆ࠄ⷗ߚ਎⇇. ╙ 7 ࿁›ጊᲧセ␠ળ⹺⍮ࠪࡦࡐࠫ࠙ࡓ㧔iCS2-7㧕 (2012/03/24-25, ੩ㇺ
ᄢቇ㔤㐳㘃⎇ⓥᚲ)㧚
⻠Ṷ
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Hayashi M (2011/08/12) Experimental approach to study cognitive development in captive and wild chimpanzees. AA
Platform Workshop "Comparative study of adaptation to environment in apes" Mabali, D.R. Congo.
Matsuzawa T (2011/04/27) What is uniquely human? An answer from the study of chimpanzees. Harvard University
Anthropology-Psychlogy Joint Special Seminar, Harvard University, USA.
Matsuzawa T (2011/04/28) What is uniquely human? An answer from the study of chimpanzees. New York Colloquium
of Primatology. Columbia University, USA.
Matsuzawa T (2011/04/29) Cognitive development in chimpanzees. Special lecture in Hunter college, City University of
New York. City University of New York, USA.
Matsuzawa T (2011/05/02) What is uniquely human? An answer from the study of chimpanzees. The Colloquium at
IRCS. University of Pennsylvania, USA.
Matsuzawa T (2011/05/17) What is uniquely human? An answer from the study of chimpanzees. Special lecture in
Institute of Health. University College London, UK.
Matsuzawa T (2011/05/18) What is uniquely human? An answer from the study of chimpanzees. Special lecture in the
Psychology and Anthropology joint colloquium. Cambridge University, UK.
Matsuzawa T (2011/06/29) Potentials for ICCA as research site for primates by Kyoto University. ICCA stakeholders
workshop, Kota Kinabalu, Malaysia.
Matsuzawa T (2011/08/16) Social evolution of the chimpanzee mind. Wellcome Trust school on biology of social
cognition. Cambridge, UK.
Matsuzawa T (2011/12/05) What is uniquely human? An answer from the study of chimpanzees. IIAS Lecture 2011 on
"Frontiers in neuroscience: from brain to mind". Tokyo.
Matsuzawa T (2011/12/09) Outgroup: A new paradigm of thinking about the evolutionary basis of human mind. IIAS
research conference 2011 on "Frontiers in neuroscience: from brain to mind". Kyoto.
Matsuzawa T (2011/12/12) What is uniquely human? An answer from the study of chimpanzees. Special lecture for the
Philosophy section of ENS. Ecole Normale Superieure, Paris.
Matsuzawa T (2011/08/06) Stone tool use of wild chimpanzees in Bossou, Guinea: Overviewofthe field experiment.
International Workshop on Primate Archaeology: Stone tool use in fossil hominids and nonhuman primates. Nairobi,
Kenya.
Tomonaga M (2011/04/11-12) How the chimpanzees see the social world: Comparative cognitive approach. 2011
International Conference on Social Cognition and Neuroscience in Hangzhou, China. Hangzhou, China.
ᨋ⟤㉿ (2011/05/09) ࠴ࡦࡄࡦࠫ࡯ߩ⎇ⓥ߆ࠄ. ฬฎደᏒ┙ᳯ⷏ዊቇᩞ㧚
ᨋ⟤㉿ (2011/07/15) ੱߣࡅ࠻‫ޔ‬หߓߎߣߣ㆑ߞߡ޿ࠆߎߣ. 2011 ᐕᐲ߈ࠂ߁ߐࠇࠎ቟ዬᭉᬺ࠯ࡒ࠽࡯࡞ߊࠄߒ.
ฬฎደ᷼㧚
᧻ᴛື㇢ (2011/05/21) ੱ㑆ߣߪ૗߆㧦࠴ࡦࡄࡦࠫ࡯⎇ⓥ߆ࠄߺ߃ߡ߈ߚߎߣ. ╙ 62 ࿁ᜰቯㇺᏒቇᩞ଻ஜද⼏
ળ‫ޟ‬ౝ⑼࡮⌒⑼࡮⡊㥦ຜ༄⑼วห⎇ୃળ‫ޠ‬. ੩ㇺ㧚
᧻ᴛື㇢ (2011/05/21) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ᄢ㒋ᢎ⢒ᄢቇઃዻ㜞╬ቇᩞ.
ᄢ㒋㧚
㧙㧙
19) ᧻ᴛື㇢ (2011/05/26) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ᣿๺㜞╬ቇᩞ SSH ၮ⺞⻠Ṷ.
›ጊ㧚
20) ᧻ᴛື㇢ (2011/06/04) ሶ⢒ߡߩ↱᧪㧦࠴ࡦࡄࡦࠫ࡯⎇ⓥ߆ࠄߺ߃ߡ߈ߚߎߣ. ᧲੩ YMCA ㊁ዥࠠࡖࡦࡊ 80
๟ᐕ⸥ᔨ⻠Ṷ. ᧲੩㧚
21) ᧻ᴛື㇢ (2011/06/04) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ࠫࡘࡦࠢၴᦠᐫ. ᧲੩㧚
22) ᧻ᴛື㇢ (2011/06/04) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ࡇ࡯ࠛࠬᩣᑼળ␠ࡈࠜ࡯࡜
ࡓ. ᧲੩㧚
23) ᧻ᴛື㇢ (2011/07/09) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ਔ࿖㜞╬ቇᩞഃ┙⊖ච๟ᐕ
⸥ᔨ⻠Ṷ. ᧲੩㧚
24) ᧻ᴛື㇢ (2011/10/14) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ㄭ⇰∛ᱧ▤ℂ࠮ࡒ࠽࡯100
࿁⸥ᔨ࠮ࡒ࠽࡯. ᄢ㒋㧚
25) ᧻ᴛື㇢ (2011/10/20) ੱ㑆ߣߪ૗߆㧦࠴ࡦࡄࡦࠫ࡯⎇ⓥ߆ࠄߺ߃ߡ߈ߚߎߣ. ╙ 47 ࿁ᣣᧄ⿒චሼ␠කቇળ✚
ળ. ⑔੗㧚
26) ᧻ᴛື㇢ (2011/10/22) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ╙ 10 ᦼ‫ޟ‬The challenge of
leadership program‫ޠ‬. ੩ㇺ㧚
27) ᧻ᴛື㇢ (2011/11/05) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ޽޿ߜࠨࠗࠛࡦࠬࡈࠚࠬ࠹
ࠖࡃ࡞ 2011. ฬฎደ㧚
28) ᧻ᴛື㇢ (2011/11/25) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ᷦ⼱ᢎ⢒᐀ᒛਛቇ࡮㜞╬ቇᩞ
ㅴ〝⻠Ṷળ. ජ⪲㧚
29) ᧻ᴛື㇢ (2011/12/08) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ᗲ⍮⋵ක⮎ຠᎿᬺදળ. ฬ
ฎደ㧚
30) ᧻ᴛື㇢ (2011/07/10) ࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩⷫሶ㑐ଥߣሶ⢒ߡ. ╙ 47 ࿁ᣣᧄ๟↥ᦼ࡮ᣂ↢ఽක
ቇળቇⴚ㓸ળ. ᧅᏻ㧚
31) ෹᳗㓷Ꮖ (2011/04/17) ࠴ࡦࡄࡦࠫ࡯ߩ⿒ߜ߾ࠎߣ߅Უߐࠎ㧦ᦨೋߩ 2 ᐕ㑆. ߰ߚߏߩ࠴ࡦࡄࡦࠫ࡯࠳ࠗࡗߣࠨ
ࠢ࡜ 2 ᱦ⺀↢ળ౏㐿⻠ᐳ. 㜞⍮⋵┙ߩ޿ߜേ‛౏࿦㧚
32) ෹᳗㓷Ꮖ (2011/07/21) ᫪ߩߎߎࠈ‫ޔ‬ᶏߩߎߎࠈ㧙ߎߎࠈߩㅴൻߦ߅ߌࠆ 2 ߟߩ೙⚂㧙. 㑐⷏ቇ㒮ᄢቇᄢቇ㒮
GP㧔⚵❱⊛ߥᄢቇ㒮ᢎ⢒ᡷ㕟ផㅴࡊࡠࠣ࡜ࡓ㧕‫ޟ‬࿖㓙ൻ␠ળߦ⽸₂ߔࠆᔃℂ⑼ቇታ〣ኅߩ㙃ᚑ‫⻠ޠ‬Ṷળ. 㑐
⷏ቇ㒮ᄢቇ㧚
33) ෹᳗㓷Ꮖ (2011/07/30) ࡅ࠻ߩߎߎࠈࠍ࠴ࡦࡄࡦࠫ࡯߆ࠄ⌑߼ࠆ. ᣣᧄቇⴚળ⼏ᔃߩవ┵⎇ⓥߣᔃℂቇኾ㐷ᢎ
⢒ಽ⑼ળ౏㐿ࠪࡦࡐࠫ࠙ࡓ ‫ޟ‬ᔃߩవ┵⎇ⓥ߳ߩᚺ‫ޠ‬. ᾢᧄᄢቇ㧚
34) ෹᳗㓷Ꮖ (2011/08/05) ࠴ࡦࡄࡦࠫ࡯ߩߎߎࠈߩ਎⇇. ᣣᧄ↢‛ᢎ⢒ળ╙ 66 ࿁ో࿖ᄢળᗲ⍮ᄢળ⎇ୃ⻠ᐳ. ੩
ㇺᄢቇ㔤㐳㘃⎇ⓥᚲ㧚
35) ෹᳗㓷Ꮖ (2011/10/06-07) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ␠ળ⊛⹺⍮. ↢ℂቇ⎇ⓥᚲ⎇ⓥળ␠ળ␹⚻⑼ቇ⎇ⓥળ‫ޔ੹ޟ‬
␠ળ␹⚻⑼ቇ⎇ⓥߦ᳞߼ࠄࠇߡ޿ࠆߎߣ‫ޠ‬. ⥄ὼ⑼ቇ⎇ⓥᯏ᭴ጟፒࠦࡦࡈࠚ࡟ࡦࠬ࠮ࡦ࠲࡯㧚
36) ෹᳗㓷Ꮖ (2011/10/13) 3 ߟߩᤨ㑆ࠍ↢߈ࠆ࠴ࡦࡄࡦࠫ࡯㧙ㅴൻ‫⊒ޔ‬㆐‫ޔ‬ᢥൻ㧙. ᣣᧄേ‛ᔃℂቇળ╙ 156 ࿁଀
ળ‫₹ޔ‬Ꮉᄢቇࠣࡠ࡯ࡃ࡞ COE ․೎ࡢ࡯࡚ࠢࠪ࠶ࡊ㧔౒௅㧦ၮ⋚⎇ⓥ(S)‫ޟ‬ᶏߩߎߎࠈ‫ޔ‬᫪ߩߎߎࠈ‫ޠ‬㧕. ₹Ꮉ
ᄢቇ㧚
37) Dahl CD (2012/01/17) Primate face perception. Graduate school of education, Kyoto University, Myowa-Yamakoshi lab,
Japan㧚
38) ᨋ⟤㉿ (2012/01/28) ࠴ࡦࡄࡦࠫ࡯ߣੱߩ⊒㆐࡮⢒ఽᡰេ. ⷏ችᏒ᳃㑆଻⢒ᚲද⼏ળ⡯ຬ⎇ୃળ. ቲႦ㧚
39) ᧻ᴛື㇢ (2012/01/20) ᗐ௝ߔࠆߜ߆ࠄ. ‫ࠍ᧪ᧂޟ‬ត⍮ߔࠆ࿾ߩࡃ࠻ࡦ࡝࡟࡯‫ ╙ޠ‬1 ࿁ੱ㘃ߩᧂ᧪. ᣣᧄ⑼ቇᧂ
᧪㙚. ᧲੩㧚
40) ᧻ᴛື㇢ (2012/01/21) ᗐ௝ߔࠆߜ߆ࠄ. ᧲ᄢ࡮੩ᄢ࡚ࠫࠗࡦ࠻৻⥸౏㐿ࠪࡦࡐࠫ࠙ࡓ‫ߩ‛߈↢ޟ‬୘ᕈࠍ⚌ߣ
ߊ㧦߆ߚߜ㧘߆߆ࠊࠅ㧘ߎߎࠈ‫ޠ‬. ᧲੩ᄢቇ㧚
41) ᧻ᴛື㇢ (2012/02/12) ࡅ࠻⑼ 4 ዻߩᲧセ߆ࠄߺߚੱ㑆ߩᔃߩ↱᧪. ᣣᧄቇⴚળ⼏౏㐿ࠪࡦࡐࠫ࠙ࡓ㧘ࡢࠗ࡞࠼
ࠨࠗࠛࡦࠬ‫ޟ‬᫪㧘ੱ㧘ᔃߩ↱᧪ࠍ߼ߋߞߡ‫ޠ‬. ደਭፉ㧚
42) ᧻ᴛື㇢ (2012/02/17) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ᗲ⍮⋵ᐔᚑ 23 ᐕᐲࠝ࡯ࡊࡦ
࠮ࡒ࠽࡯㧔╙ 6 ࿁㧕. ฬฎደ㧚
43) ᧻ᴛື㇢ (2012/02/21) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ⡛Უቇ㒮ዊቇᩞᢎ⢒⻠Ṷળ.
੩ㇺ㧚
44) ᧻ᴛື㇢ (2012/03/02) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ࿖㓙࠰ࡠࡊ࠴ࡒࠬ࠻ᄢᵤ 40
๟ᐕ⸥ᔨ⻠Ṷ. ᄢᵤ㧚
45) ᧻ᴛື㇢ (2012/03/13) ᗐ௝ߔࠆߜ߆ࠄ㧙࠴ࡦࡄࡦࠫ࡯߇ᢎ߃ߡߊࠇߚੱ㑆ߩᔃ. ੩ㇺᄢቇ࠙ࠗ࡞ࠬ⎇ⓥᚲ࠮
ࡒ࠽࡯. ੩ㇺᄢቇ࠙ࠗ࡞ࠬ⎇ⓥᚲ㧚
46) ෹᳗㓷Ꮖ (2012/01/19) ࠴ࡦࡄࡦࠫ࡯߆ࠄ⷗ߚᲣሶ㑐ଥߩ⊒㆐. ╙ 45 ࿁ᴒ✽⋵Უሶ଻ஜᄢળ․೎⻠Ṷ. ቱ㊁ḧ
Ꮢ᳃ળ㙚㧚
㧙㧙
47) ෹᳗㓷Ꮖ (2012/01/28-29) Factors affecting motion direction judgment in chimpanzees. ࿖㓙㜞╬⎇ⓥᚲ⎇ⓥࡊࡠࠫ
ࠚࠢ࠻‫ޟ‬ᔃߩ⿠Ḯ‫ޠ‬2011 ᐕᐲ╙ 3 ࿁⎇ⓥળ. ࿖㓙㜞╬⎇ⓥᚲ㧚
48) ෹᳗㓷Ꮖ (2012/02/27-28) ᶏߩߎߎࠈ‫ޔ‬᫪ߩߎߎࠈ. ╙ 2 ࿁࠴ࡦࡄࡦࠫ࡯㘺⢒⠪⎇ୃળ⻠Ṷ. ጟጊ㧚
⹺⍮ቇ⠌ಽ㊁
㧨⎇ⓥ᭎ⷐ㧪
A) ࡅ࠻ߣ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ⹺⍮ᯏ⢻ߩട㦂ᄌൻߦߟ޿ߡߩታ㛎⊛Ყセ⎇ⓥ
ᱜ㜞ା↵㧘ศᎹᏀ♿ሶ㧔੩ᄢ࡮ߎߎࠈߩᧂ᧪⎇ⓥ࠮ࡦ࠲࡯㧕㧘Ꮉวિᐘ㧔ฬฎደᄢቇ㧕㧘ਭ଻ධᶏሶ㧔ᗲ⍮ᶻᓼᄢቇ㧕
⹺⍮ᯏ⢻ߩട㦂ߦߣ߽ߥ߁ᄌൻࠍੱ㑆ߣ࠾ࡎࡦࠩ࡞ߢᲧセࠍ߅ߎߥߞߚ‫ޕ‬
B) ⊒㆐㓚ኂߩᒝߺ㧔strength㧕ߩታ㛎⊛ᬌ⸽
ᱜ㜞ା↵
ⷞⷡត⚝⺖㗴ࠍታ㛎ࡄ࡜࠳ࠗࡓߣߒߡ↪޿ߚታ㛎ࠍቯဳ⊒㆐ఽ 20 ฬߦ߅ߎߥ޿‫⚿ޔ‬ᨐࠍ㓚ኂఽߣᲧセߒߚ‫ޕ‬
C) 㔤㐳㘃ߩࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߩㅴൻߦ㑐ߔࠆ⎇ⓥ
㚅↰໪⾆㧘ᩊፒోᒄ㧘૒⮮᧙ᄹ㧘H Bouchet㧔ᣣᧄቇⴚᝄ⥝ળᶏᄖ․೎⎇ⓥຬ㧔᰷☨⍴ᦼ㧕
㧕
㧘Ꮉวિᐘ㧔ฬฎደᄢ
ቇ㧕
㧘ᣧᎹ␽ሶ㧘ട⮮ᧇ⟤㧘࿡ᨑඅ㧘⍹↰ᕺሶ㧘ධ㔕⚐ᴦ㧔㔤㐳⎇࡮࿖㓙౒หవ┵⎇ⓥ࠮ࡦ࠲࡯㧕㧘ᱜ㜞ା↵
࠾ࡎࡦࠩ࡞‫ࠍߤߥ࡞ࠩࠟ࠽࠹ޔࡦࡁࠛࠣޔ‬ኻ⽎ߦ‫ޔ‬㔤㐳㘃ߩⷞ⡬ⷡࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦ߇ߤߩࠃ߁ߦㅴൻߒߡ߈
ߚߩ߆ࠍ‫ޔ‬ታ㛎ቶ‫ޔ‬㊁↢ਅߩਔ⠪ߦ߅޿ߡ‫⎇࠼࡞࡯ࠖࡈޔ‬ⓥߣታ㛎⎇ⓥߩਔ㕙߆ࠄ⎇ⓥࠍⴕߞߡ޿ࠆ‫ޕ‬
D) 㔤㐳㘃ߩࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߩㅴൻߦ㑐ߔࠆ࿖㓙౒ห⎇ⓥ
㚅↰໪⾆㧘K Zuberbuhler (University of St Andrews)
⇣⒳㑆ࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߩᚑ┙ߩㆊ⒟ߦߟ޿ߡ‫ޔ‬㐳ᦼ࿖㓙౒ห⎇ⓥࠍ㐿ᆎߒߚ‫ޕ‬University of St Andrews ߢᵴ
േࠍⴕߞߡ޿ࠆ‫ޕ‬
E) ደਭፉߦ߅ߌࠆ㊁↢࠾ࡎࡦࠩ࡞ߩⴕേ࡮↢ᘒቇ⊛⎇ⓥ
H Bouchet㧘㚅↰໪⾆
ደਭፉߩ㊁↢࠾ࡎࡦࠩ࡞ࠍኻ⽎ߣߒߡ‫⊒ޔ‬ᖱᤨߩࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߩᬌ⸛ࠍⴕߞߚ‫ޕ‬
F) ࠹ࡦࠣࠩ࡞ߩㆮવ࡮ⴕേ࡮␠ળߩ⎇ⓥ
᧛੗഼⵨㧘ᣧᎹ␽ሶ㧘㚅↰໪⾆
ࠗࡦ࠼ࡀࠪࠕ࡮ࠬ࡜ࡃࡗേ‛࿦ߦዉ౉ߐࠇߚᲧセ⊛ᄢⷙᮨߥ࠹ࡦࠣࠩ࡞⟲ࠍኻ⽎ߣߒ‫ޔ‬ㆮવቇ࡮ⴕേቇ࡮␠ળቇ
⊛⎇ⓥࠍⴕߞߚ‫ޕ‬
G) ࠾ࡎࡦࠩ࡞ߩᤨ㑆⍮ⷡߦ㑐ߔࠆ⎇ⓥ
ᩊፒోᒄ㧘㚅↰໪⾆㧘ᱜ㜞ା↵
࠾ࡎࡦࠩ࡞ࠍⵍ㛎૕ߣߒߡ‫ޔ‬㑆㓒ੑಽᴺߦࠃࠆᤨ㑆㐳ᑯ೎ߩታ㛎ࠍⴕߥ޿‫ᤨޔ‬㑆㐳ߩ್ᢿߦߺࠄࠇࠆ․ᓽ߿‫ᤨޔ‬
㑆⍮ⷡߦ෸߷ߔ⻉ⷐ࿃ߦߟ޿ߡᬌ⸛ߒߚ‫ޕ‬
H) ⥄Ꮖᖱႎಣℂߦ㑐ߔࠆ੐⽎㑐ㅪ㔚૏⎇ⓥ
Ỉ↰⃍ሶ㧘ᱜ㜞ା↵
⥄Ꮖᖱႎಣℂߩᤨ㑆․ᕈࠍ⺞ߴࠆߚ߼ߦ‫ޔ‬ᚑੱࠍኻ⽎ߦ‫⥄ޔ‬Ꮖ㑐ㅪೝỗ‫ઁޔ‬⠪㑐ㅪೝỗࠍⷰኤਛߩ⣖ᵄࠍ⸘᷹ߒ
ߚ‫ޕ‬
I) ⥄㐽∝ఽߦ߅ߌࠆ⍮⼂࡮⸥ᙘߩ᳢ൻߩ⧰ᚻߐߦߟ޿ߡߩ⹺⍮ታ㛎߅ࠃ߮ࡆ࠺ࠝߦࠃࠆⴕേⷰኤ
દ⮮␭ᐽ㧘ᱜ㜞ା↵
⊒㆐㓚ኂఽࠍኻ⽎ߦ⺰ℂṶ▚ࠍℂ⸃ߢ߈ࠆ߆ߦߟ޿ߡߩ⹺⍮ታ㛎ࠍⴕߞߚ‫ޕ‬
J) ⊒㆐㓚ኂఽ߳ߩቇ⠌ᡰេߣߘߩ⹏ଔ⎇ⓥ
દ⮮␭ᐽ㧘ዊᎹ⹞ਫ㧘੗↰⟤ᴕሶ㧘⏷᧛᦮ሶ㧘↰ਛ⟤ㇺ㧘↰᧛✍⩿㧔ᤘ๺ᅚሶᄢቇ㧕㧘⑔ፉ⟤๺㧔᧲੩ᄢቇ㧕㧘⦁ᦜ
ᐽሶ㧔੩ᄢ࡮කቇㇱ㧕㧘㐳ጟජ⾐㧔੩ᄢ࡮ߎߎࠈߩᧂ᧪⎇ⓥ࠮ࡦ࠲࡯㧕㧘᫪ፒ␞ሶ㧔੩ᄢ࡮ߎߎࠈߩᧂ᧪⎇ⓥ࠮ࡦ࠲
࡯㧕
㧘ศᎹᏀ♿ሶ㧔੩ᄢ࡮ߎߎࠈߩᧂ᧪⎇ⓥ࠮ࡦ࠲࡯㧕㧘ᱜ㜞ା↵
੩ㇺᄢቇߎߎࠈߩᧂ᧪⎇ⓥ࠮ࡦ࠲࡯ߦ߅޿ߡ‫ ⚂ޔ‬30 ฬߩቇ⠌ߦ࿎㔍ࠍ␜ߔ⊒㆐㓚ኂఽࠍኻ⽎ߦ⺒ߺᦠ߈ࠍਛᔃ
ߣߒߚቇ⠌ᡰេࠍⴕߞߚ‫ޕ‬
K) e ࡜࡯࠾ࡦࠣࠍᩭߣߔࠆᄙ᭽ߥቇ⠌࿎㔍ߦኻᔕߒߚኅᐸቇ⠌ᡰេᣇᴺߩ᭴▽
ዊᎹ⹞ਫ㧘⑔ፉ⟤๺㧔᧲੩ᄢቇ㧕㧘ਭ଻ධᶏሶ㧔ᗲ⍮ᶻᓼᄢቇ㧕㧘ᱜ㜞ା↵
⚂ 10 ฬߩ⺒ߺᦠ߈ߦ࿎㔍ࠍ␜ߔቇ⠌㓚ኂߩఽ┬ߦ‫࡮࠽ࠞ࠲ࠞ࡮ߥ߇ࠄ߭ޔ‬ṽሼߩ⺒ߺᦠ߈ቇ⠌ߩߚ߼ߦ㐿⊒ߒ
ߚࡄ࠰ࠦࡦ↪ᢎ᧚࠰ࡈ࠻ࠍឭଏߒ‫ߩߘޔ‬ലᨐࠍᬌ⸛ߒߚ‫ޕ‬
㧙㧙
L) ቇᩞ⃻႐ߦ߅ߌࠆ⊒㆐㓚ኂఽ߳ߩ IT ᢎ᧚ࠍᩭߣߒߚቇ⠌ᡰេߩታ〣⎇ⓥ
દ⮮␭ᐽ㧘ዊᎹ⹞ਫ㧘ᷡ㐳⼾㧘੗↰⟤ᴕሶ㧘↰ਛ⟤ㇺ㧘ጊ↰ᥓሶ㧘ᱜ㜞ା↵
ቇᩞ⃻႐ߦ߅޿ߡᣣᧄ⺆ߩ߭ࠄ߇ߥ࡮ࠞ࠲ࠞ࠽ߩ⺒ߺᦠ߈ቇ⠌ߩߚ߼ߦ㐿⊒ߒߚࡄ࠰ࠦࡦ↪ᢎ᧚࠰ࡈ࠻ࠍឭଏߒ‫ޔ‬
ቇᩞ⃻႐ߢߩᵴ↪ߦߟ޿ߡߩᬌ⸽߿ߘߩലᨐࠍᬌ⸛ߒߚ‫ޕ‬
M) ᄖ࿖ੱఽ┬↢ᓤߩ㖸㖿ᗧ⼂⊒㆐
ᷡ㐳⼾㧘ᱜ㜞ା↵
ጘ㒂⋵นఽᏒߩዊቇ৻ᐕ↢ߩᄖ࿖ੱఽ┬ࠍኻ⽎ߦ㖸ჿ⍮ⷡታ㛎ࠍታᣉߒߚ‫⚿ߩߘޕ‬ᨐ‫ߥ߆ޔ‬ᢥሼ⠌ᓧߦ߅޿ߡ㖸
㖿ᗧ⼂ߩ⊒㆐߇߭ࠄ߇ߥ⺒ߺᦠ߈ߩၮ␆⊛᧦ઙߢ޽ࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬
N) ⥄㐽∝ఽߦ߅ߌࠆⷞⷡត⚝⎇ⓥ
⏷᧛᦮ሶ㧘દ⮮␭ᐽ㧘ᱜ㜞ା↵
⥄㐽∝ఽߦ߅ߌࠆ⴫ᖱ⹺⍮߿ᖱേಣℂߦ㑐ࠊࠆ⹺⍮․ᕈࠍ‫ⷡⷞߚ޿↪ࠍ࡞ࡀࡄ࠴࠶࠲࠲࡯ࡘࡇࡦࠦޔ‬ត⚝⺖㗴ߦ
ࠃߞߡᬌ⸛ߒߚ‫ޕ‬
O) 㔤㐳㘃ߦ߅ߌࠆ੃ఽ↹௝߳ߩㆬᅢᕈߩᬌ⸛
૒⮮᧙ᄹ㧘㚅↰໪⾆㧘ධ㔕⚐ᴦ㧔㔤㐳⎇࡮࿖㓙౒หవ┵⎇ⓥ࠮ࡦ࠲࡯㧕㧘A Lemasson (University of Rennes 1)㧘ᱜ㜞
ା↵
ᣥ਎⇇ࠩ࡞ 2 ⒳ࠍኻ⽎ߦ‫↹ߩఽ੃࡞ࠩࡦࡎ࠾ޔ‬௝ࠍೝỗߣߒߡ↪޿‫↹ఽ੃ޔ‬௝߳ߩㆬᅢᕈߩ᦭ήࠍታ㛎ᔃℂቇ⊛
ߦᬌ⸛ߒߚ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
2)
3)
4)
✚⺑
1)
2)
Hayakawa S, Kawai N, Masataka N (2011) The influence of color on the snake detection in visual search in young
children. Scientific Reports 1:80.
Lemasson A, Glas L, Barbu S, Lacroix A, Guilloux M, Remeuf K, Koda H (2011) Youngsters do not pay attention to
conversational rules: is this so for nonhuman primates? Scientific Reports 1, Article number: 22.
Masataka N (2011) Enhancement of speech-relevant auditory acuity in absolute pitch possessors. Front. Psychology
2:101.
Masataka N, Shibasaki M (2012) Premenstrual enhancement of snake detection in visual search in healthy women.
Scientific Reports 2:307.
⏷᧛᦮ሶ㧘ᱜ㜞ା↵ (2011) ⣖ߩน႟ᕈߣቇ⠌߅ࠃ߮⊒㆐㓚ኂ. ૞ᬺ≮ᴺࠫࡖ࡯࠽࡞ 45(7):656-660.
૒⮮᧙ᄹ㧘ᱜ㜞ା↵ (2011) ჿߪ޿߆ߦߒߡ⹤ߒߎߣ߫ߦߥߞߚ߆. Journal of Otolaryngology, Head and Neck
Surgery 27(8):1169-1174.
ߘߩઁߩၫ╩
1)
ศᎹᏀ♿ሶ㧘ዊᎹ⹞ਫ (2011) ⊒㆐㓚ኂߣ⺒ߺᦠ߈ᡰេ. ቇⴚᐢႎ⹹‫╙ޠ᧪ᧂߩࠈߎߎޟ‬㧣ภ p.38 ߎߎࠈߩ
ᧂ᧪⎇ⓥ࠮ࡦ࠲࡯⊒ⴕ㧚
ቇળ⊒⴫
1)
2)
3)
4)
5)
6)
7)
Fukushima M, Ogawa S, Tamura A, Ito H, Masataka N (2011) Changes in Reading and Writing Aspects of Japanese
Children with Learning Difficulties through Individual Learning Support. 15th European Conference in Developmental
Psychology (2011/08/23-27, Norway).
Tamura A, Ogawa S, Ito H, Yoshikawa S, Masataka N (2011) The response patterns of children with developmental
disorders in interpersonal conflict situations. 15th European Conference in Developmental Psychology (2011/08/23-27,
Norway).
⏷᧛᦮ሶ㧘દ⮮␭ᐽ㧘ዊᎹ⹞ਫ㧘ᱜ㜞ା↵ (2011) ⥄㐽∝ఽߩᖱേಣℂㆊ⒟ߦ߅ߌࠆᵈᗧ㚟േ᭽ᑼߩಽᨆ. 2011
ᐕᐲ൮᜝⣖ࡀ࠶࠻ࡢ࡯ࠢᄐߩࡢ࡯࡚ࠢࠪ࠶ࡊ (2011/08/21, ౓ᐶ)㧚
⏷᧛᦮ሶ㧘દ⮮␭ᐽ㧘ዊᎹ⹞ਫ㧘ᱜ㜞ା↵ (2011) ⥄㐽∝ఽߦ߅ߌࠆⷞⷡត⚝⎇ⓥ—ᖱേߣࠣࡠ࡯ࡃ࡞ࠠࡘ࡯࡮
ࡠ࡯ࠞ࡞ࠠࡘ࡯. ╙ 35 ࿁ᣣᧄ␹⚻ᔃℂቇળ✚ળ (2011/09/15, ᩔᧁ)㧚
દ⮮␭ᐽ (2011) ߆ߥߩ⺒ߺᦠ߈࿎㔍ࠍਥ⸷ߣߔࠆఽ┬߳ߩᡰេ⺖㗴ߩ⚫੺.⺒ߺᦠ߈࿎㔍ߦ㑐ࠊࠆ⹺⍮ᯏ⢻
ߩ⹏ଔᣇᴺߣߘߩታ〣㨪․ᓽߦᔕߓߚᡰេࠍ⠨߃ࠆ㨪⥄ਥࠪࡦࡐࠫ࠙ࡓ㧔⹤㗴ឭଏ⠪㧕. ᣣᧄ LD ቇળ╙ 20
࿁ᄢળ (2011/09/18, ᧲੩)㧚
ᷡ㐳⼾ (2011) ⺒ߺᦠ߈ߦ㑐ࠊࠆ⹺⍮⢻ജߩ⹏ଔ㧔1㧕㖸㖿ᖱႎಣℂㆊ⒟㧚⺒ߺᦠ߈࿎㔍ߦ㑐ࠊࠆ⹺⍮ᯏ⢻ߩ
⹏ଔᣇᴺߣߘߩታ〣㨪․ᓽߦᔕߓߚᡰេࠍ⠨߃ࠆ㨪⥄ਥࠪࡦࡐࠫ࠙ࡓ㧔⹤㗴ឭଏ⠪㧕. ᣣᧄ LD ቇળ╙ 20 ࿁
ᄢળ (2011/09/18, ᧲੩)㧚
ᷡ㐳⼾㧘ᱜ㜞ା↵ (2011) ᄖ࿖ੱఽ┬↢ᓤߩ㖸㖿ᗧ⼂⊒㆐. ᣣᧄ LD ቇળ╙ 20 ࿁ᄢળ (2011/09/19, ᧲੩)㧚
㧙㧙
8)
9)
10)
11)
12)
13)
14)
15)
⻠Ṷ
1)
ዊᎹ⹞ਫ (2011) ⺒ߺᦠ߈ߦ㑐ࠊࠆ⹺⍮⢻ജߩ⹏ଔ㧔2㧕ⷞⷡᖱႎಣℂㆊ⒟㧚⺒ߺᦠ߈࿎㔍ߦ㑐ࠊࠆ⹺⍮ᯏ⢻
ߩ⹏ଔᣇᴺߣߘߩታ〣 㨪․ᓽߦᔕߓߚᡰេࠍ⠨߃ࠆ㨪⥄ਥࠪࡦࡐࠫ࠙ࡓ㧔⹤㗴ឭଏ⠪㧕. ᣣᧄ LD ቇળ╙ 20
࿁ᄢળ (2011/09/18, ᧲੩)㧚
ዊᎹ⹞ਫ㧘⑔ፉ⟤๺㧘ᱜ㜞ା↵ (2011) ⊒㆐㓚ኂఽߦ߅ߌࠆ⺒ߺᦠ߈࿎㔍․ᕈߩ߫ࠄߟ߈. ᣣᧄ LD ቇળ╙ 20
࿁ᄢળ (2011/09/17, ᧲੩)㧚
૒⮮᧙ᄹ㧘ട⮮ᧇ⟤㧘㚅↰໪⾆ (2011) ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ੃ఽ㗻ೝỗ߳ߩⷞⷡ⊛ᵈᗧ. Animal 2011 ᣣᧄേ‛
ⴕേቇળ╙ 30 ࿁ᄢળ (2011/09/09-10, ᧲੩)㧚
૒⮮᧙ᄹ㧘ട⮮ᧇ⟤㧘㚅↰໪⾆ (2011) ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆᐜఽ࿑ᑼ߳ߩㆬᛯ⊛ᵈᗧ. ᣣᧄ㔤㐳㘃ቇળ╙ 27
࿁ቇⴚᄢળ (2011/07/16-18, ᗲ⍮)㧚
ᩊፒోᒄ㧘⦁ᯅᣂᄥ㇢㧘࿡ᨑඅ㧘㚅↰໪⾆㧘ᱜ㜞ା↵ (2011) ࠕࠞࠥࠩ࡞ߦ߅ߌࠆ࠮࡞ࡈࠦࡦ࠻ࡠ࡯࡞⺖㗴ߩ
ᬌ⸛. ᣣᧄⴕേಽᨆቇળ╙ 29 ࿁ᄢળ (2011/09/18, ᧲੩)㧚
ᩊፒోᒄ㧘㚅↰໪⾆㧘ᱜ㜞ା↵ (2011) 㖸ჿࡊ࡜ࠗࡓೝỗ߇࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ੃ఽ㗻ೝỗ߳ߩⷞⷡ⊛ᵈᗧ. ᣣ
ᧄേ‛ᔃℂቇળ╙ 71 ࿁ᄢળ (2011/09/09, ᧲੩)㧚
੗↰⟤ᴕሶ (2012) ␠ળᕈᡰេߩታ〣ႎ๔㧚࡜࠙ࡦ࠼࠹࡯ࡉ࡞‫⊒ޟ‬㆐㓚ኂఽ߳ߩ⊒㆐ᡰេ߇߽ߚࠄߔലᨐߩᄙ
ⷺ⊛ᬌ⸛‫ޠ‬㧔⹤㗴ឭଏ⠪㧕. ╙ 23 ࿁ᣣᧄ⊒㆐ᔃℂቇળ (2012/03/10, ᗲ⍮)㧚
ዊᎹ⹞ਫ㧘᫪ፒ␞ሶ (2012) ᡰេ⠪ߩ௛߈߆ߌߦ㑐ߔࠆᬌ⸛㧚࡜࠙ࡦ࠼࠹࡯ࡉ࡞‫⊒ޟ‬㆐㓚ኂఽ߳ߩ⊒㆐ᡰេ߇
߽ߚࠄߔലᨐߩᄙⷺ⊛ᬌ⸛‫ޠ‬㧔⹤㗴ឭଏ⠪㧕. ╙ 23 ࿁ᣣᧄ⊒㆐ᔃℂቇળ (2012/03/10, ᗲ⍮)㧚
Ỉ↰⃍ሶ (2011/05/21) ࠬ࡯ࡄ࡯ࠨࠗࠛࡦࠬࡂࠗࠬࠢ࡯࡞⻠Ṷ. ᱞᐶᎹᅚሶᄢቇઃዻਛቇ࡮㜞╬ቇᩞ, ⷏ችᏒ,
ᄢቇ㒮ߢቇ߱ߣ޿߁ߎߣቇ໧ࠍㅢߓߡࡅ࠻ࠍ⍮ࠆ㧚
㜞ᰴ⣖ᯏ⢻ಽ㊁
㧨⎇ⓥ᭎ⷐ㧪
A) 㗻⴫ᖱߩᄌൻߦ઻߁ᚳ᩶૕࠾ࡘ࡯ࡠࡦᔕ╵
ୖጟᐽᴦ㧘ਛ᧛స᮸
␠ળ⊛ᖱႎߩಣℂߦ㑐ࠊࠆ⣖ౝᯏᐨࠍ⸃᣿ߔࠆߎߣࠍ⋡⊛ߦ‫ࠍ࡞ࠩࠥࠞࠕޔ‬ኻ⽎ߣߒߡ‫ઁޔ‬୘૕ߩ㗻⴫ᖱࠍᑯ೎
ߔࠆ⺖㗴ࠍ⸠✵ߒߚ‫⴫ߦࠄߐޕ‬ᖱ㑆ࠍㅪ⛯⊛ߦᄌൻߐߖࠆೝỗ↹௝ࠍ૞ᚑߒ‫ߩࠄࠇߘޔ‬㗻ೝỗࠍ⷗ߡ޿ࠆߣ߈ߦࠨ
࡞ᚳ᩶૕ࠃࠅන৻࠾ࡘ࡯ࡠࡦᔕ╵ࠍ⸥㍳ߒߚ‫ޕ‬
B) ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ߩ⹺⍮ᯏ⢻⸘᷹
ਛ᧛స᮸㧘┻ᧄ◊ผ㧘ᧁ႐␞ሶ㧘ጊญᥓᕺሶ㧘ਃベ⟤᮸㧘ᴰ᣿ብ㧘ၳ↰⧷⩐㧘ᷰㄝᥓሶ
ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ߩ⹺⍮ᯏ⢻㧔⍮ⷡ࡮⸥ᙘ╬㧕ࠍ⺞ߴࠆߚ߼ߦ‫⟎ⵝߩߘޔ‬㐿⊒ࠍ฽߼ᣇᴺߩ⏕┙ࠍ⋡ᜰߒߚ⎇
ⓥࠍታᣉߒߡ޿ࠆ‫ޕ‬ዊဳߩ᳢↪⹺⍮ᯏ⢻ታ㛎ⵝ⟎ࠍ㐿⊒ߒ‫ⷡⷞޔ‬ᑯ೎⺖㗴࡮ㅒォቇ⠌⺖㗴࡮ㆃᑧ⷗ᧄวߖ⺖㗴࡮㗅
ᐨቇ⠌⺖㗴╬ࠍ⸠✵ߒ‫ߢ࠻࠶࠮ࡕ࡯ࡑࡦࡕࠦޔ‬ㆀⴕน⢻ߥߎߣࠍ᣿ࠄ߆ߦߒߚ‫╭ޔߚ߹ޕ‬ജ᷹ቯߩ◲ଢߥⵝ⟎ࠍ㐿
⊒ߒ‫ߩ࠻࠶࠮ࡕ࡯ࡑࡦࡕࠦޔ‬೨⢇ߩ╭ജࠍ᷹ቯߒߚ‫ޕ‬
C) ੃ᐜఽߩⷞ✢⸘᷹ߦၮߠߊേ૞ℂ⸃ߩ⊒㆐⎇ⓥ
ਛ᧛స᮸㧘ਛ᧛ᓼሶ㧔ᤘ๺ᅚሶᄢቇ㧕㧘૒‫ਂᧁޘ‬ᄦ㧔ᣣᧄ౏ᢥᢎ⢒⎇ⓥળ㧕
ஜᏱఽߣ⊒㆐㓚ኂఽߩേ૞ℂ⸃⢻ജࠍᲧセ࡮ᬌ⸛ߔࠆߚ߼ߦ‫ޔ‬㕖ଚⷅ⊛ߦⷞ✢ࠍ⸘᷹ߔࠆኾ↪ⵝ⟎ࠍ↪޿ⷞⷡೝ
ỗߦኻߔࠆᵈⷞࡄ࠲࡯ࡦࠍ⺞ߴߡ޿ࠆ‫ޕ‬
D) ࡅ࠻ߩ࡝࠭ࡓ೙ᓮߩ␹⚻ࡔࠞ࠾࠭ࡓ⸃᣿
㡨ᳰ⩿଻㧘ୖጟᐽಃ㧘ች࿾㊀ᒄ㧘᧖ᶆర੫㧔᧲ർᄢቇ㧕㧘Ꮉፉ㓉ᄥ㧔᧲ർᄢቇ㧕㧘ਛ᧛స᮸
࡝࠭ࡓ⸥ᙘߩ⣖ౝᯏ᭴ࠍ᣿ࠄ߆ߦߔࠆߚ߼‫ޔ‬ஜᏱᚑੱࠍኻ⽎ߣߒߡ࡝࠭ࡓ⸥ᙘ࡮ౣ↢⺖㗴ㆀⴕਛߩ⣖ᵴേࠍ‫ޔ‬ᯏ
⢻⊛ MRI ࠍ↪޿ߡ⸘᷹ߒߚ‫ࡓ࠭࡝ޕ‬ᖱႎߩ⸥ฬ‫ޔ‬ౣ↢ߦ㑐ㅪߒߡਅ೨㗡࿁‫ޔ‬ਅ㗡㗂ዊ⪲‫⿷⵬ޔ‬ㆇേ㊁‫ޔ‬ዊ⣖߆ࠄ
ߥࠆࡀ࠶࠻ࡢ࡯ࠢ߇⾮ᵴߐࠇߚ‫ޔߚ߹ޕ‬ਅ೨㗡࿁ࠍ㒰ߊ㗔ၞߪ࡝࠭ࡓᖱႎ଻ᜬߩᦼ㑆ߦ߽⾮ᵴߒߡ޿ࠆߎߣ߇ಽ߆
ߞߚ‫ߩࠄࠇߎޕ‬㗔ၞߩ⾮ᵴࡄ࠲࡯ࡦߪ‫ߦ⊛ⷡⷞ߇ࡓ࠭࡝ޔ‬ឭ␜ߐࠇߡ߽⡬ⷡ⊛ߦឭ␜ߐࠇߡ߽Ꮕ߇ߥߊ‫ޔ‬ᗵⷡࡕ࠳
࡝࠹ࠖࠍ⿥߃ߡಣℂߐࠇߡ޿ࠆߣ⠨߃ࠄࠇߚ‫ޕ‬
E) ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆᕈߩ⹺⍮ߣࡎ࡞ࡕࡦߩ㑐ㅪᕈߩ⸃᣿
ᧁ႐␞ሶ㧘ਛ᧛స᮸
ᕈࡎ࡞ࡕࡦ߇ᕈߩ⹺⍮ߦਈ߃ࠆᓇ㗀ࠍᬌ⸛ߔࠆߚ߼ߦ‫ࠍ࡞ࠩࡦࡎ࠾ޔ‬ኻ⽎ߣߒߡ‫ઁޔ‬୘૕ߩ㗻౮⌀ߥߤߩⷞⷡೝ
ỗߦኻߔࠆᑯ೎⢻ജ߿ㆬᅢᕈߣ޿ߞߚ⹺⍮ߣ‫ࡦࡕ࡞ࡎޔ‬േᘒߣߩ㑐ㅪᕈࠍ⺞ߴࠆ⸘↹ߢ޽ࠆ‫⺖ޕ‬㗴ㆀⴕ୘૕ߩዩߩ
ណขࠍ߅ߎߥ޿‫ࡦࡕ࡞ࡎޔ‬᷹ቯࠍ߅ߎߥ߁‫ޕ‬
F) ᄢ⣖⊹⾰␹⚻࿁〝ߩ↢ᓟ⊒㆐ߩ⎇ⓥ
ች࿾㊀ᒄ㧘ᄢ⍹㜞↢㧔⛔ว⣖ࠪࠬ࠹ࡓ㧕㧘㜞↰᣽ᒾ㧔⛔ว⣖ࠪࠬ࠹ࡓ㧕㧘᪍၂ዊ⊖ว㧔⛔ว⣖ࠪࠬ࠹ࡓ㧕㧘ችㇱ⾆ሶ
㧙㧙
㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕
ࡅ࠻ࠍ฽߻㔤㐳㘃ߩᄢ⣖ᣂ⊹⾰ߪ↢ᓟ߽⊒㆐ࠍ⛯ߌࠆߎߣ߇⍮ࠄࠇߡ޿ࠆ‫ⴕޕ‬േ೙ᓮߦߣߊߦ㊀ⷐߢ޽ࠆᄖ஥೨
㗡೨㊁ࠍ฽߻␹⚻࿁〝ߩ↢ᓟ⊒㆐ㆊ⒟ࠍ᣿ࠄ߆ߦߔࠆߚ߼‫ޔ‬ᐜ⧯ࠨ࡞ߩᄖ஥೨㗡೨㊁߳ߩ␹⚻౉ജ᭽ᑼࠍ⸃೬ቇ⊛
ߦ⸃ᨆߒߚ‫ޔߚ߹ޕ‬ኻᾖታ㛎ߣߒߡ‫ޔ‬ᚑᾫ୘૕ߦߟ޿ߡ߽ห᭽ߩታ㛎ࠍⴕߞߚ‫ޕ‬
G) ㆇേ㑐ㅪ⊹⾰ߩ↢ᓟ⊒㆐ߩ␹⚻⸃೬࡮␹⚻↢ℂቇ⊛⎇ⓥ
ች࿾㊀ᒄ㧘⑧භ㓷ผ
೨㗡⪲ߩߐ߹ߑ߹ߥㆇേ㑐ㅪ㗔㊁ߩᯏ⢻ߩ↢ᓟ⊒㆐ࠍ᣿ࠄ߆ߦߔࠆߚ߼‫࡞ࠨࠢࠞࡑޔ‬ᚑᾫ୘૕߅ࠃ߮ᐜ⧯୘૕ࠍ
ኻ⽎ߦ‫ࠆߥ⇣ߩ࠲࡯ࡔ࡜ࡄޔ‬㔚᳇ࡄ࡞ࠬߦࠃࠅ‫⾰⊹ߩࠇߙࠇߘޔ‬㗔ၞࠍೝỗߒ‫ޔ‬ೝỗߦࠃࠅ⺃⊒ߐࠇࠆߐ߹ߑ߹ߥ
ㆇേࠍⷰኤ‫⸥ޔ‬㍳ߒ‫᦬ޔ‬㦂‫ޔ‬ᐕ㦂ߏߣߦᲧセߔࠆ‫ᧄޕ‬ᐕᐲߪ‫↢ޔ‬ᓟ㧝ᐕߩᐜ⧯୘૕ࠍኻ⽎ߦ‫ޔ‬೨㗡⪲⊹⾰ࠍߐ߹ߑ
߹ߥೝỗࡄ࡜ࡔ࡯࠲ߢೝỗߒ‫ߚࠇߐ⊒⺃ޔ‬ㆇേࠍⷰኤߒߚ‫ޕ‬
H) ⦡ᑯ೎⺖㗴ㆀⴕਛߩࠨ࡞೨㗡ㅪว㊁ߦ߅ߌࠆ࠾ࡘ࡯ࡠࡦᵴേߩ⸃ᨆ
⍹Ꮉ⋥᮸㧘 ੗ ⡡㧘੗਄㓷ੳ㧘ች࿾㊀ᒄ㧘ਃ਄┨మ
⦡ᑯ೎ߣ⸥ᙘࠍ઻߁⌒⃿ㆇേ⺖㗴ㆀⴕਛߩࠨ࡞೨㗡ㅪว㊁߆ࠄ‫ࡦࡠ࡯ࡘ࠾ޔ‬ᵴേࠍ⸥㍳ߒ‫⊒࠻ࠬ࡯ࡃޔ‬Ἣߩ᦭ή
ߣࡄ࠲࡯ࡦߩ㆑޿ࠍᚻដ߆ࠅߣߒߡ࠲ࠗࡊಽ㘃ࠍⴕߞߚ‫ߩߘޕ‬ᓟ‫ޔ‬ฦ࠲ࠗࡊߩ⚦⢩ߣ⺖㗴ߣߩ㑐ㅪᕈࠍ⸃ᨆߒߚ‫ޕ‬
I) ࠨ࡞ߩ࡝࠭ࡓ೙ᓮߩ␹⚻ࡔࠞ࠾࠭ࡓߩ⸃᣿
㡨ᳰ⩿଻㧘ች࿾㊀ᒄ
࡝࠭ࡓ೙ᓮߩ␹⚻ᯏ᭴ࠍ᣿ࠄ߆ߦߔࠆߎߣࠍ⋡⊛ߣߒߡ‫ߦ࠻ࡅޔ‬ㄭ޿⊒㆐ߒߚ⣖ࠍᜬߜ‫ⶄޔ‬㔀ߥⴕേ⺖㗴ࠍቇ⠌
ߢ߈ࠆࡑࠞࠢࠨ࡞ࠍኻ⽎ߣߒߡ࡝࠭ࡒࠞ࡞ߦࡏ࠲ࡦࠍ᛼ߔ⺖㗴ࠍ⸠✵ߒߚ‫ᧄޕ‬ᐕᐲߪⴕേ࠺࡯࠲ߩ⸃ᨆࠍ߅ߎߥߞ
ߚ‫৻ޔߪߡ޿߅ߦ࡞ࠨࠢࠞࡑޕ‬ቯ㑆㓒ߩࡏ࠲ࡦ᛼ߒ⺖㗴ߢߪ࡜ࡦ࠳ࡓ㑆㓒ߩ႐วߦᲧߴߡ෻ᔕᤨ㑆߇⍴❗ߒ‫ޔ‬ᰴߩ
ࡏ࠲ࡦ᛼ߒߩ࠲ࠗࡒࡦࠣࠍ੍᷹ߒߡⴕേߒߡ޿ࠆߎߣ߇᣿ࠄ߆ߦߥߞߚ‫ޕ‬
J) 㐳ᦼ⸥ᙘ߅ࠃ߮⍴ᦼ⸥ᙘߦၮߠߊⴕേ᳿ቯߩ␹⚻ᯏ᭴ߩ⎇ⓥ
⑧භ㓷ผ㧘ች࿾㊀ᒄ㧘ୖጟᐽᴦ㧘ਛ᧛స᮸
ᧄ⎇ⓥߢߪ‫ޔ‬⍴ᦼ⸥ᙘߦၮߠߊⴕേ᳿ቯ߅ࠃ߮㐳ᦼ⸥ᙘߦၮߠߊⴕേ᳿ቯࠍࠨ࡞ߦⴕࠊߖ‫ߩߘޔ‬㓙ߩ␹⚻ᵴേࠍ
ᄖ஥೨㗡೨㊁ߦ߅޿ߡ⸥㍳‫⸃ޔ‬ᨆߒߚ‫ޔߦߢ߹ࠇߎޕ‬㐳ᦼ⸥ᙘߦၮߠߊⴕേ᳿ቯ‫߮ࠃ߅ޔ‬⍴ᦼ⸥ᙘߦၮߠߊⴕേ᳿
ቯߩߘࠇߙࠇߦ․⇣⊛ߥ␹⚻ᵴേࠍ⸥㍳ߢ߈ߚ‫ޕ‬
K) ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ߩ⊒ჿⴕേߩ⎇ⓥ
ᴰ᣿ብ㧘ጊญᥓᕺሶ㧘ਛ᧛స᮸
ࡑ࡯ࡕ࠮࠶࠻ߩ⊒ჿⴕേߩ୘૕ᕈߦߟ޿ߡᬌ⸛ߔࠆߚ߼ߦ‫ޔ‬หዬ㘺⢒ߐࠇߡ޿ࠆኅᣖ୘૕߆ࠄߩ㓒㔌߇⊒ჿⴕേ
ߦਈ߃ࠆᓇ㗀ߦߟ޿ߡᬌ⸛ߒߚ‫ޕ‬
L) ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ߩ⡬ⷡ♽೉ߩ⍮ⷡ᭽ᑼߩ⸃᣿
⣁↰⌀ᷡ
ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ࠍ↪޿ߡ⡬ⷡᑯ೎⸠✵ࠍⴕߞߚ‫౒ߪ⚛ⷐޕ‬ㅢߢ޽ࠆ߇㈩೉ߩ⇣ߥࠆੑߟߩ㖸♽೉ࠍ↪޿ߡ‫ޔ‬
⋧ኻᑯ೎⺖㗴ߣ⛘ኻᑯ೎⺖㗴ࠍⴕߞߚ‫⚿ޕ‬ᨐ‫⋧ޔ‬ኻᑯ೎᧦ઙߢߪᑯ೎߇ߢ߈ߡ߽⛘ኻᑯ೎᧦ઙߢߪᑯ೎߇ߢ߈ߥ߆
ߞߚ‫⚿ߩߎޕ‬ᨐߪࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻߇⡬ⷡ♽೉ߩⷙೣᕈࠍ᛽಴ߔࠆߎߣߪߢ߈ߡ߽‫ࠍࠄࠇߘޔ‬㐳ᦼ⸥ᙘߦ⾂⬿ߢ
߈ߥ޿ߎߣࠍ␜ߒߡ޿ࠆ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
2)
3)
4)
5)
6)
7)
8)
Kuraoka K, Nakamura K (2011) The use of nasal skin temperature measurements in studying emotion in macaque
monkeys. Physiology & Behavior 102:347-355.
Saito A, Izumi A, Nakamura K (2011) Development of infant common marmosets' (Callithrix jacchus) preference for
their parents over adults from another group. Primates 52:43-50.
Saito A, Izumi A, Nakamura K (2011) Fathers have higher motivation for parenting than mothers in common marmoset
(Callithrix jacchus). Behavior 148:1199-1214.
Saito A, Nakamura K (2011) Oxytocin changes primate paternal tolerance to offspring in food transfer. Journal of
Comparative Physiology 197:329-337.
Takemoto A, Izumi A, Miwa M, Nakamura K (2011) Development of a compact and general-purpose experimental
apparatus with a touch-sensitive screen for use in evaluating cognitive functions in common marmosets. Journal of
Neuroscience Methods 199:82-86.
Yumoto N, Lu X, Henry TR, Miyachi S, Nambu A, Fukai T, Takada M (2011) A neural correlate of the processing of
multi-second time intervals in primate prefrontal cortex. PLoS One.Apr 27 6(4):e19168.
㊀૑๟㧘ේ⧷ਯ㧘┻ᧄ◊ผ㧘࿯ᯅ↱ታ㧘ਛ᧛స᮸㧘᧻ᧄ㓉 (2011) ⣖ᵄ㧦SSVEP4 ࠢ࡜್ࠬ೎໧㗴ߩㆀᰴဳቇ
⠌㧦Sequential Monte Carlo ታⵝ. 㔚ሶᖱႎㅢାቇળ ାቇᛛႎ, Vol.110 No.460, MBE2010-125:125-130.
Naoi N, Minagawa-Kawai Y, Kobayashi A, Takeuchi K, Nakamura K, Yamamoto J, Kojima S (2012) Cerebral
responses to infant-directed speech and the effect of talker familiarity. NeuroImage 59:1735-1744.
㧙㧙
9)
Dobashi Y, Takemoto A, Shigezumi S, Shiraki T, Nakamura K, Matsumoto T (2012) Automatic determination of
stopping time of training phase in SSVEP-based brain-machine interface with Bayesian sequential learning.International
Conference on BioMedical Engineering, Feb. 2012, Vienna, Austria.
10) Wakita M, Hiraishi H (2011) Effects of handedness and viewing perspective on Broca's area activity. NeuroReport
22(2):331-336.
✚⺑
1)
2)
ਛ᧛స᮸ (2011) ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ࠍ↪޿ߚ㔤㐳㘃⎇ⓥߩേะ. ၮ␆ᔃℂቇ⎇ⓥ 30:79-85.
ਛ᧛స᮸ (2011) ♖␹∔ᖚߩ⸃᣿ߩߚ߼ߩ㔤㐳㘃ࡕ࠺࡞ ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ߩน⢻ᕈ. ࡈࠔࡑ࡞ࠪࠕ
47:799-803.
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
Saito A, Nakamura K (2011) Parent-infant relationship in marmosets. (Monkeys: Biology, Behavior and Disorders.) (ed.
Williams RM) p.77-95 Nova Science Publishers, Inc.
ߘߩઁၫ╩
1)
2)
ਛ᧛స᮸ (2011) ‫ޟ‬⣖ࠍ㎊߃ߚ޿ ⊝વ㧍ᣂ޽ߚ߹㆏႐‫ޠ‬໧㗴૞ᚑ Ფᣣᣂ⡞㧚
ਛ᧛స᮸ (2011) ࠗࠢࡔࡦߩࠨ࡞. p.79-84 ↢߈‛ߚߜߩߟߠࠇ❱ࠅ ╙ 5 Ꮞ㧔gCOE ᐢႎ⹹㧕㧚
ቇળ⊒⴫
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Konoike N, Kotozaki Y, Miyachi S, Miyachi CM, Yomogida Y, Akimoto Y, Kuraoka K, Sugiura M, Kawashima R,
Nakamura K (2011) Different contributions of frontal, parietal, and temporal cortices to working memory of rhythm.
41th Annual meeting, Society for Neuroscience (2011/11, Washington, DC).
Kuraoka K, Nakamura K (2011) Categorical representation of social information in the central nucleus of monkey
amygdale. 41th Annual meeting, Society for Neuroscience (2011/11, Washington, DC).
Nejime M, Inoue M, Saruwatari M, Nakamura K, Miyachi S (2011) Prefrontal neuron activities in short-term
memory-based and long-term memory-based decision processes. 5th International Symposium of the Biodiversity &
Evolution Global COE (2011/07, Kyoto)㧚
Nejime M, Inoue M, Saruwatari M, Nakamura K, Miyachi M (2011) Differential activities of monkey lateral prefrontal
neurons between decisions based on short-term and long-term memories. 41th Annual meeting, Society for
Neuroscience (2011/11, Washington, DC).
⩵ᳰℭℂ૫㧘ਛ᧛స᮸ (2011) ‫ߦ߾ߜ߽߅↪࠻ࡅߩ࠻࠶࠮ࡕ࡯ࡑࡦࡕࠦޟ‬ኻߔࠆㆬᅢᕈߩᕈᏅ‫ޠ‬. ╙ 27 ࿁ᣣᧄ
㔤㐳㘃ቇળ (2011/07/16-18, ›ጊᏒ)㧚
㟃↰ቁ๺㧘ᮮ੗ഞ㧘ጟỈ೰⿠㧘ㇹ↰⋥৻㧘ᐔ᧻ජ዆㧘ਃ਄┨మ㧘ࠞࡦ࠹ࠖ࡮ࠕ࡜ࡓ࡮࠙ࠖ࠳ࡗ࠹ࠖ㧘ች࿾㊀
ᒄ㧘ᚭᎹ᫪㓶㧘㜞ᧁᱜᶈ㧘ዊ᧻⧷ᒾ (2011) ੑ⦡ᕈࡑࠞࠢࠩ࡞ߩⴕേታ㛎ߦࠃࠆ⦡ⷡ࠹ࠬ࠻. ╙ 34 ࿁ᣣᧄ␹⚻
⑼ቇᄢળ (2011/09/14-17, ᮮᵿ)㧚
ୖጟᐽᴦ㧘ਛ᧛స᮸ (2011) ࠨ࡞ߩᚳ᩶૕ߣ⣻ᄖ஥೨㗡೨⊹⾰ߢ⇣ߥࠆ⴫ᖱᖱႎ⴫⃻ߩᤨ㑆⊛ᄌൻ. ╙ 34 ࿁ᣣ
ᧄ␹⚻⑼ቇᄢળ (2011/09/14-17, ᮮᵿ)㧚
ች࿾㊀ᒄ㧘ᐔ↰ᔟᵗ㧘ᯫ၂ዊ⊖ว㧘㤥↰๒ሶ㧘ችㇱ⾆ሶ㧘㜞↰᣽ᒾ㧘ᄢ⍹㜞↢ (2011) ᐜ⧯ࡑࠞࠢࠨ࡞ᄖ஥೨
㗡೨㊁߳ߩ⊹⾰⊹⾰౉ജ. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/17, ᮮᵿᏒ)㧚
ਛ᧛స᮸㧘┻ᧄ◊ผ㧘ᧁ႐␞ሶ㧘ਃベ⟤᮸㧘ጊญᥓᕺሶ (2011) ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ߦ߅ߌࠆⷞⷡ⊛ㆃᑧ⷗ᧄ
วߖ⺖㗴. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/14-17, ᮮᵿ)㧚
⣁↰⌀ᷡ (2011) ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻㧔Callithrix jacchus㧕ߦ߅ߌࠆಽ▵ߩ⡬ⷡᑯ೎. ᣣᧄേ‛ᔃℂቇળ㧔╙ 71
࿁㧕(2011/09/08-11, ᧲੩ㇺ)㧚
⣁↰⌀ᷡ (2011) ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ߦ߅ߌࠆ⡬ⷡ♽೉ᑯ೎. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/14-17, ᮮᵿ)㧚
⣁↰⌀ᷡ (2011) Ꮐਅ೨㗡㗔ၞߪㆇേߩᤨ㑆⊛೙ᓮߦ߽㑐ࠊࠆ. ᣣᧄᔃℂቇળ╙ 75 ࿁ᄢળ (2011/09/15-17, ᧲
੩ㇺ)㧚
Nejime M, Inoue M, Saruwatari M, Nakamura K, Miyachi S (2012) Neuronal activities of the monkey lateral prefrontal
cortex reflect difference of memory source in the memory-based behavioral choice. Kyoto University and The
University of Tokyo Global COE Joint Symposium (2012/01, Tokyo)㧚
Nakamura K, Takemoto A, Koba R, Miwa M (2012) Characteristics of serial order learning in Common Marmosets.
International Symposiums “Frontiers in Primate Neuroscience Research” (2012/02/22-24, Tokyo)㧚
⻠Ṷ
1) Nakamura K (2011/06/07) “Neural mechanisms of nonverbal communication.” The 15th annual meeting of the
Association for the Scientific Study of Consciousness (ASSC) Social Neuroscience Satellite, Joint Tamagawa Caltech,
Lecture Course 2011 Kyoto University, Kyoto㧚
2) ਛ᧛స᮸ (2012/02/01) ‫“ޟ‬᳇ߦߥࠆሶ”ߣ⊒㆐㓚ኂ㧔⥄㐽∝ࠍਛᔃߣߒߚ㧕ߩሶߤ߽ߚߜߩℂ⸃ߣᡰេ 㨪
KUMON ߣߩ౒ห⎇ⓥ߆ࠄ⷗߃ߡߊࠆሶ⢒ߡߣᢎ⢒㨪‫ ޠ‬ർᶏ㆏ࠛ࡝ࠕ㓚ኂఽᜰዉࡈࠜ࡯࡜ࡓ. ᧅᏻ㧚
3) ਛ᧛స᮸ (2012/03/28) ‫“ޟ‬᳇ߦߥࠆሶ”ߣ⊒㆐㓚ኂ㧔⥄㐽∝ࠍਛᔃߣߒߚ㧕ߩሶߤ߽ߚߜߩℂ⸃ߣᡰេ 㨪
KUMON ߣߩ౒ห⎇ⓥ߆ࠄ⷗߃ߡߊࠆሶ⢒ߡߣᢎ⢒㨪‫਻ ޠ‬Ꮊᴒ✽ࠛ࡝ࠕ㓚ኂఽᜰዉࡈࠜ࡯࡜ࡓ. ⑔ጟ㧚
㧙㧙
ಽሶ↢ℂ⎇ⓥㇱ㐷
⛔ว⣖ࠪࠬ࠹ࡓಽ㊁
㧨⎇ⓥ᭎ⷐ㧪
A) ␹⚻〝ㆬᛯ⊛ߥᵴേᛥ೙ߣ࠻࡟࡯ࠪࡦࠣߦࠃࠆᄢ⣖ࡀ࠶࠻ࡢ࡯ࠢߩ᭴▽ߣᯏ⢻ߩ⸃᣿
㜞↰᣽ᒾ㧘੗਄⻞৻
␹⚻〝ㆬᛯ⊛ᵴേᛥ೙ᴺߩ⏕┙ߣᔕ↪ߦߟ޿ߡߪ‫⹺ޔ‬⍮ㆊ⒟ࠍ฽߻⌒⃿ㆇേ⺖㗴߅ࠃ߮਄⢇ㆇേ⺖㗴ߩ 2 ⒳㘃ߩ
ⴕേ⺖㗴ࠍ⸠✵ߒߚࠨ࡞ߦ߅޿ߡ‫ࡦࠪࠠ࠻ࠬ࠿࠲࠹ޔ‬シ㎮ࡈ࡜ࠣࡔࡦ࠻ߩㆬᛯ⊛ዉ౉ߦࠃࠆ␹⚻ᵴേᛥ೙߇ⴕേߦ
෸߷ߔᓇ㗀ߩ⸃ᨆࠍㅴ߼ߡ޿ࠆ‫ౕޕ‬૕⊛ߦߪ‫⃿⌒ޔ‬ㆇേ⺖㗴ߢߪ೨㗡⪲⌒⃿ㆇേ㑐ㅪ㗔㊁㧔೨㗡⌒㊁߿⵬⿷⌒㊁㧕
߆ࠄߩ಴ജ♽ߦ‫ޔ‬਄⢇ㆇേ⺖㗴ߢߪ✢᧦૕߅ࠃ߮೨㗡೨㊁㗔ၞ㧔․ߦ⢛ᄖ஥ㇱߣ⢛ౝ஥ㇱ㧕߳ߩ౉ജ♽ߦ⌕⋡ߒߡ‫ޔ‬
ⴕേቇ⊛⸃ᨆ߿㔚᳇↢ℂቇ⊛⸥㍳ታ㛎ࠍⴕ߁‫ޔߚ߹ޕ‬ㄭᐕ⊒⴫ߐࠇߚ␹⚻ᵴേᛥ೙♽ߢ޽ࠆ GluCl-ivermectin ᛥ೙
♽߿ CNO-DREADDs ᛥ೙♽ߥߤߩᣂⷙᛥ೙♽ߦߟ޿ߡ߽Ⓧᭂ⊛ߦዉ౉ࠍᬌ⸛ߒ‫ޔ‬㔤㐳㘃ߦ߅޿ߡᦨ߽ታ↪⊛ߥ␹
⚻〝ㆬᛯ⊛ᵴേᛥ೙ࠪࠬ࠹ࡓࠍ⏕┙ߔࠆߚ߼ߩታ㛎⸘↹ࠍ╷ቯߒߚ‫〝⚻␹ޕ‬ㆬᛯ⊛ߥㅒⴕᕈ⿧ࠪ࠽ࡊࠬ⊛࠻࡟࡯ࠪ
ࡦࠣߩታ⃻ߣᔕ↪ߦߟ޿ߡ߽‫Ⱟޔ‬శ࠲ࡦࡄࠢ⾰ࠍ↪޿ߚ⿧ࠪ࠽ࡊࠬ⊛ᄙ㊀࠻࡟࡯ࠪࡦࠣᴺߦࠃࠅ‫ޔ‬೨㗡⪲⌒⃿ㆇേ
㑐ㅪ㗔ၞ㧔೨㗡⌒㊁߿⵬⿷⌒㊁㧕߅ࠃ߮೨㗡೨㊁㗔ၞ㧔․ߦ⢛ᄖ஥ㇱߣ⢛ౝ஥ㇱ㧕߳ߩᄙࠪ࠽ࡊࠬᕈ౉ജ᭽ᑼߩ⸃
ᨆࠍㅴ߼ࠆߣߣ߽ߦ‫ޔ‬᛫⁅›∛࠙ࠗ࡞ࠬ⚦⢩ౝ᛫૕ࠍ⊒⃻ߔࠆࠕ࠺ࡁ㓐઻࠙ࠗ࡞ࠬࡌࠢ࠲࡯ࠍ↪޿ߡ‫․ޔ‬ቯߩ␹⚻
〝ࠍㆬᛯ⊛ߦࡑࠬࠢߔࠆ OFF ೙ᓮဳㅒⴕᕈ⿧ࠪ࠽ࡊࠬ⊛࠻࡟࡯ࠪࡦࠣࠪࠬ࠹ࡓࠍ⏕┙ߔࠆߚ߼ߩታ㛎⸘↹ࠍ╷ቯ
ߒߚ‫ޔߚ߹ޕ‬Cre-loxP ޽ࠆ޿ߪ Tet-On/Off ㆮવሶ⊒⃻೙ᓮࠪࠬ࠹ࡓࠍᔕ↪ߒߡ‫․ޔ‬ቯߩ␹⚻〝ࠍㆬᛯ⊛ߦ࡜ࡌ࡞ߔ
ࠆ ON ೙ᓮဳㅒⴕᕈ⿧ࠪ࠽ࡊࠬ⊛࠻࡟࡯ࠪࡦࠣࠪࠬ࠹ࡓࠍࠨ࡞ߦ߅޿ߡ⏕┙ߔࠆߚ߼‫࡯࠲ࠢࡌޔ‬ᡷ⦟ߦ઻߁ᦨㆡൻ
ᬌ⸛ታ㛎ࠍታᣉߔࠆߚ߼ߩታ㛎⸘↹ࠍ╷ቯߒߚ‫ޕ‬
B) ㆮવሶᡷᄌ㔤㐳㘃ࡕ࠺࡞ߩ㐿⊒ߣ㜞ᰴ⣖ᯏ⢻ߩ⸃ᨆ
㜞↰᣽ᒾ㧘ᄢ⍹㜞↢㧘᧻ᧄᱜᐘ㧘੗਄⻞৻㧘ੑችᄥᐔ㧘㜞ේᄢテ㧘ᯫ၂ዊ⊖ว㧘ች࿾㊀ᒄ㧔㜞ᰴ⣖ᯏ⢻㧕
Ԙ ␹⚻〝ㆬᛯ⊛ㆮવሶᠲ૞ᴺࠍ↪޿ߚᄢ⣖ၮᐩᩭߩᖱႎಣℂᯏ᭴ߣᯏ⢻⊛ᓎഀߩ⸃ᨆ㧔㜞↰㧘੗਄㧘᧻ᧄ㧕
(1) ࠗࡓࡁ࠻ࠠࠪࡦ␹⚻〝ᮡ⊛ᴺࠍ↪޿ߚࡂࠗࡄ࡯⋥ធ〝ߩㆬᛯ⊛㒰෰
ࠗࡓࡁ࠻ࠠࠪࡦ␹⚻〝ᮡ⊛ᴺࠍ↪޿ߡ‫⋥࡯ࡄࠗࡂޔ‬ធ〝ࠍㆬᛯ⊛ߦ㒰෰ߒߚࡕ࠺࡞ࠩ࡞ࠍ૞⵾ߒ‫ޔ‬ㆇേ㊁ೝ
ỗߦኻߔࠆ᷆⫷⃿ౝ▵࠾ࡘ࡯ࡠࡦߩᣧ޿⥝ᅗᕈᔕ╵߇ࡂࠗࡄ࡯⋥ធ〝ࠍ⚻↱ߔࠆߎߣࠍ᣿ࠄ߆ߦߒߚ‫⎇ᧄޕ‬
ⓥᚑᨐߦ㑐ߔࠆේ⪺⺰ᢥࠍᛩⓂߒ‫⃻ޔ‬࿷‫ޔ‬ᩏ⺒⠪ߩࠦࡔࡦ࠻ߦᓥߞߡᡷ⸓ਛߢ޽ࠆ‫ޕ‬
(2) ࠗࡓࡁ࠻ࠠࠪࡦ␹⚻〝ᮡ⊛ᴺࠍ↪޿ߚ⊹⾰਄ਐ〝ߩㆬᛯ⊛㒰෰
ࠗࡓࡁ࠻ࠠࠪࡦ␹⚻〝ᮡ⊛ᴺࠍ↪޿ߡ‫⃿⌒ޔ‬ㆇേ೙ᓮߦ㊀ⷐߥᓎഀࠍᜂߞߡ޿ࠆߣ⠨߃ࠄࠇࠆ⊹⾰਄ਐ〝
ࠍㆬᛯ⊛ߦ㒰෰ߒߚࡕ࠺࡞ࠩ࡞ࠍ૞⵾ߔࠆߎߣߦᚑഞߒߚ‫⃻ޕ‬࿷‫ⶄޔ‬ᢙߩ⌒⃿ㆇേ⺖㗴ࠍࡕ࠺࡞ࠩ࡞ߦㆀ
ⴕߐߖߡ‫ޔ‬෻ᔕᤨ㑆߿ࠛ࡜࡯₸ߩᄌൻߥߤࠍ⸃ᨆߒߡ߅ࠅ‫⾰⊹ߦߢ߹ࠇߎޔ‬਄ਐ〝߇ᄖㇱᖱႎߦၮߠߊ
attention ߦ㑐ਈߔࠆߎߣࠍ␜ໂߔࠆ⚿ᨐࠍᓧߚ‫ޕ‬
(3) Cre-loxP ␹⚻〝ㆬᛯ⊛ㆮવሶ⊒⃻೙ᓮᴺࠍ↪޿ߚ㤥⾰✢᧦૕␹⚻〝ߩㆬᛯ⊛㒰෰
Cre-loxP ␹⚻〝ㆬᛯ⊛ㆮવሶ⊒⃻೙ᓮᴺࠍ↪޿ߡ‫᧦✢ޔ‬૕ߦᛩ኿ߔࠆ㤥⾰࠼࡯ࡄࡒࡦ࠾ࡘ࡯ࡠࡦߦߘߩᄌᕈ
ࠍ⺃ዉߔࠆࠕ࡞ࡈࠔࠪ࠿ࠢ࡟ࠗࡦࠍ⊒⃻ߐߖࠆߎߣߦࠃߞߡ‫ޔ‬㤥⾰✢᧦૕␹⚻〝ࠍㆬᛯ⊛ߦ㒰෰ߒߚࡄ࡯
ࠠࡦ࠰ࡦ∛ࡕ࠺࡞ࠩ࡞ࠍ૞⵾ߔࠆߎߣߦᚑഞߒߚ‫⃻ޕ‬࿷‫ⴕޔ‬േቇ⊛߅ࠃ߮⚵❱ቇ⊛ߥ⸃ᨆࠍ߅ߎߥߞߡ޿
ࠆ‫ޕ‬
ԙ ␹⚻〝ㆬᛯ⊛ᵴേ೙ᓮᴺࠍ↪޿ߚ㔤㐳㘃ࡕ࠺࡞ߩ૞⵾㧔㜞↰㧘੗਄㧕
(1) Tet-On ␹⚻〝ㆬᛯ⊛ㆮવሶ⊒⃻೙ᓮᴺࠍ↪޿ߚ㤥⾰✢᧦૕␹⚻〝ߩㆬᛯ⊛ᵴേ㒖ኂ
ዉ౉ㆮવሶߣߒߡ࠹࠲࠿ࠬ࠻ࠠࠪࡦシ㎮ࠍ↪޿ߚ Tet-On ␹⚻〝ㆬᛯ⊛ㆮવሶ⊒⃻೙ᓮᴺߦࠃࠅ‫ޔ‬㤥⾰✢᧦
૕␹⚻〝ߩᵴേࠍㆬᛯ⊛ߦ㒖ኂߒ‫⮎ޔ‬೷⺃ዉ⊛߆ߟนㅒ⊛ߦㆇേ㓚ኂࠍ⺃⊒ߔࠆࡕ࠺࡞ࠩ࡞ࠍ૞⵾ߔࠆߎ
ߣߦᚑഞߒߚ‫ޕ‬
(2) ㅒⴕᕈᗵᨴဳ࡟ࡦ࠴࠙ࠗ࡞ࠬࡌࠢ࠲࡯ߩᡷ⦟ߣࠨ࡞⣖߳ߩㆮવሶዉ౉
⑔ፉ⋵┙ක⑼ᄢቇߣߩ౒ห⎇ⓥߦࠃࠅ‫࡞ࠨޔ‬⣖ߦ߅޿ߡ㜞޿ㅒⴕᕈㆮવሶዉ౉ല₸ߣߣ߽ߦఝࠇߚ࠾ࡘ࡯
ࡠࡦ․⇣ᕈࠍ᦭ߔࠆㅒⴕᕈᗵᨴဳ࡟ࡦ࠴࠙ࠗ࡞ࠬࡌࠢ࠲࡯㧔FuG-C ဳ㧕ࠍ㐿⊒ߒ‫ߩߘޔ‬ᚑᨐࠍේ⪺⺰ᢥߣ
ߒߡ⊒⴫ߒߚ‫ޕ‬
Ԛ ታⴕᯏ⢻ߩ⣖ౝࡔࠞ࠾࠭ࡓߩ⎇ⓥ㧔ᄢ⍹㧘ች࿾㧘ᯫ၂㧕
⢛ᄖ஥೨㗡೨㊁߇㑐ਈߔࠆታⴕᯏ⢻ߦ⢛ᄖ஥೨㗡೨㊁߳ߩⶄᢙߩ౉ജ߇ߘࠇߙࠇߤߩࠃ߁ߦነਈߒߡ޿ࠆ߆
ࠍㅴ߼ߡ޿ࠆ‫⥄ޕ‬Ꮖⴕേߩࡕ࠾࠲࡯‫‛ޔ‬૕⹺⼂‫ޔ‬ⓨ㑆⹺⼂ߩ߁ߜ‫․ޔ‬ቯߩ߽ߩߩࡢ࡯ࠠࡦࠣࡔࡕ࡝࡯߇ᔅⷐ
ߥ 3 ߟߩⴕേ⺖㗴ࠍోߡ⸠✵ߒߚ୘૕ࠍ૞ᚑߒߚ‫✵⸠ޕ‬ቢੌ୘૕ߩ߁ߜ 1 㗡ߪ࠼ࠠࠪࠨࠗࠢ࡝ࡦᛩਈଐሽ⊛
ߦ․ቯߩ␹⚻〝ߩࠪ࠽ࡊࠬવ㆐ࠍᛥ೙ߔࠆߚ߼ߩࡌࠢ࠲࡯ᵈ౉ࠍⴕߞߚ‫ޕ‬
ԛ 㔤㐳㘃⣖ߩㆮવሶ⊒⃻ࡄ࠲࡯ࡦߣߘߩട㦂ߦ㑐ߔࠆ⎇ⓥ㧔ᄢ⍹㧘ᯫ၂㧕
ࡑࠞࠢ⣖ౝߩㆮવሶ⊒⃻ߩട㦂ᄌൻࠍ✂⟜⊛ㆮવሶ⊒⃻⸃ᨆߢᬌ⸛ߒߚ‫ޕ‬ᯏ⢻ㆮવሶ⟲ߩ⊒⃻ߩჇᷫࠍᲧセ
ߒߚߣߎࠈ‫ޔ‬㓽㓶࡮ㇱ૏ࠍ໧ࠊߕ఺∉‫ޔ‬Ἳ∝෻ᔕ㑐ㅪㆮવሶ⊒⃻߇Ⴧടߒ‫ࠕ࡝࠼ࡦࠦ࠻ࡒޔ‬㑐ㅪㆮવሶߩ⊒
⃻ߪᷫዋߒߡ޿ߚ‫ࠬࡊ࠽ࠪޕ‬વ㆐ߦ㑐ࠊࠆㆮવሶ⟲ߩ⊒⃻ߪ㓶ࠃࠅ߽㓽ߢ‫ޔ‬೨㗡೨㊁ࠃࠅᶏ㚍ߢૐਅ߇⪺ߒ
㧙㧙
߆ߞߚ‫ޕ‬
C) ࠨ࡞ࡕ࠺࡞ߦࠃࠆ⊹⾰⣄㜑〝ߩน႟ᕈ೙ᓮᯏ᭴ߩᬌ⸛
㜞↰᣽ᒾ㧘ੑችᄥᐔ
ࠨ࡞ࠍ↪޿ߡ ஥⣄㜑៊்ࡕ࠺࡞ࠍ૞⵾ߒ‫ⴕޔ‬േቇ⊛߅ࠃ߮ᒻᘒቇ⊛⸃ᨆࠍታᣉߒߚ‫ౕޕ‬૕⊛ߦߪ‫ޔ‬⣄㜑៊்ߩ
೨ᓟߦ♖ኒᛠᜬ⺖㗴ࠍㆀⴕߐߖߚ⚿ᨐ‫்៊ޔ‬ᓟᢙᣣߢᯏ⢻࿁ᓳ߇ᆎ߹ࠅ‫ޔ‬1㨪3 ࡩ᦬ᓟߦߪ៊்೨ߣ߶߷ᄌࠊࠄߥ
޿⒟ᐲߦ߹ߢ࿁ᓳߔࠆߎߣࠍ⏕⹺ߒߚ‫⾰⊹ߢ࡞ࠩ࡞࠺ࡕߥ߁ࠃߩߎޕ‬⣄㜑〝ߩ㗅ⴕᕈ␹⚻〝࠻࡟࡯ࠪࡦࠣࠍⴕ޿‫ޔ‬
៊்ߒߚ⣄㜑ߩᡰ㈩஥ߣ෻ኻ஥ߩᄢ⣖ඨ⃿ߦ߅޿ߡ‫৻ޔ‬ᰴㆇേ㊁߆ࠄ⣄㜑 motor neurons ߦะ߆ߞߡ␹⚻ゲ⚝߇ᣂ
ߚߦિ㐳ߒߡ޿ࠆߎߣࠍ⷗಴ߒߚ‫ޔߦࠄߐޕ‬෻⊒ᕈゲ⚝⺃ዉ࿃ሶߢ޽ࠆ ephrin B3 ߇⣄㜑ᱜਛઃㄭߦᒝߊ⊒⃻ߒߡ
޿ࠆߎߣࠍ㔤㐳㘃ߢೋ߼ߡ᣿ࠄ߆ߦߒߚ‫᧪ޕ‬ᐕᐲߪ‫ߚ޿↪ࠍࠬ࡞ࠗ࠙∛›⁅ޔ‬ㅒⴕᕈ⿧ࠪ࠽ࡊࠬ⊛␹⚻〝࠻࡟࡯ࠪ
ࡦࠣߦࠃࠅ‫ޔ‬⣄㜑៊்ࡕ࠺࡞ߦ߅޿ߡ‫৻ޔ‬ᰴㆇേ㊁ࠍ฽߻೨㗡⪲ㆇേ㑐ㅪ㗔㊁߆ࠄ⣄㜑㧔propriospinal neurons,
segmental interneurons, motor neurons㧕‫╭ߦࠄߐޔ‬⡺ߦ⥋ࠆᄙࠪ࠽ࡊࠬᕈ␹⚻〝ߩᒻᘒቇ⊛⸃ᨆࠍⴕ߁ߣߣ߽ߦ‫␹ޔ‬
⚻࿁〝ౣ✬ᚑߦ㑐ࠊࠆᯏ⢻ಽሶߩ⊒⃻ߣߘߩᓎഀߦߟ޿ߡ⹦⚦ߦᬌ⸛ߔࠆ‫ޕ‬
D) 㔤㐳㘃ߩᄢ⣖̆ዊ⣖̆ၮᐩᩭࡀ࠶࠻ࡢ࡯ࠢߦ߅ߌࠆㆇേᖱႎಣℂߩಽᢔߣ⛔ว
㜞↰᣽ᒾ㧘ᄢ⍹㜞↢㧘᧻ᧄᱜᐘ㧘੗਄⻞৻㧘㜞ේᄢテ㧘ᧁ᧛ᵴ↢
(1) 㗅ⴕᕈ࠻࡟࡯ࠪࡦࠣᴺߦࠃࠆ᭴ㅧ⸃ᨆ
㗅ⴕᕈ࠻࡟࡯ࠨ࡯ߢ޽ࠆ BDA ࠍࡑࠞࠢࠩ࡞ߩㆇേ೨㊁⢛஥ㇱߩ਄⢇㗔ၞߦᵈ౉ߒ‫৻ޔ‬ᰴㆇേ㊁ߦ߅޿ߡ
࡜ࡌ࡞ߐࠇߚ✢⛽⚳ᧃߩጀ․⇣⊛ಽᏓ᭽ᑼࠍ⸃ᨆߒߚ‫੹ޕ‬ᓟߪ‫ޔ‬2 ⒳㘃ߩ㗅ⴕᕈ࠻࡟࡯ࠨ࡯㧔BDA ߣ
WGA-HRP㧕ࠍㆇേ೨㊁⢛஥ㇱߣ⵬⿷ㆇേ㊁ߩ਄⢇㗔ၞߦᵈ౉ߒ‫৻ߩᧃ⚳⛽✢ߚࠇߐ࡞ࡌ࡜ޔ‬ᰴㆇേ㊁ߦ
߅ߌࠆጀ․⇣ᕈߦߟ޿ߡᬌ⸛ߔࠆ੍ቯߢ޽ࠆ‫ޕ‬
(2) ␹⚻〝ㆬᛯ⊛ᠲ૞ࡕ࠺࡞ࠨ࡞ߩ૞⵾
ዊᨋࠣ࡞࡯ࡊߣ౒ห㐿⊒ߒߚ㜞㗫ᐲㅒⴕᕈㆮવሶዉ౉㧔HiRet㧕ࡌࠢ࠲࡯ࠍ↪޿ߚࠗࡓࡁ࠻ࠠࠪࡦ␹⚻〝ᮡ
⊛ᴺ߿ࠗࡓࡁ࠹࠲࠿ࠬ␹⚻〝ᮡ⊛ᴺߦࠃࠅ‫ޔ‬ᄢ⣖‫ޔ‬ዊ⣖‫ޔ‬ၮᐩᩭࠍ❬ߋࡀ࠶࠻ࡢ࡯ࠢࠍ᭴ᚑߔࠆ୘೎ߩ␹
⚻〝ࠍㆬᛯ⊛ߦ㒰෰޽ࠆ޿ߪᵴേᛥ೙ߒߚㆮવሶᡷᄌࠨ࡞ࡕ࠺࡞ߩ૞⵾ࠍ⸘↹ਛߢ޽ࠆ‫ޕ‬
E) ㆇേ㓚ኂߣ⹺⍮㓚ኂࠍಾࠅಽߌࠆࡄ࡯ࠠࡦ࠰ࡦ∛ߩࠨ࡯ࠠ࠶࠻ࡄ࠰ࡠࠫ࡯
㜞↰᣽ᒾ㧘੗਄⻞৻
(1) -synuclein ⊒⃻࠙ࠗ࡞ࠬࡌࠢ࠲࡯ߩ૞⵾
⊒⃻೙ᓮࡌࠢ࠲࡯ߣߒߡ‫ޔ‬AAV-floxed/stop-GFP ࡌࠢ࠲࡯ߩ GFP ㈩೉ࠍ‫ޔ‬wild-type human -synuclein ߦട
߃ߡ‫ޔ‬A53P ᄌ⇣ဳ -synuclein ߩ㈩೉ߦ⟎឵ߒߚࡌࠢ࠲࡯ࠍ૞⵾ߒߚ‫ޕ‬
(2) -synuclein ⺃ዉᕈࡄ࡯ࠠࡦ࠰ࡦ∛ࡕ࠺࡞ࠩ࡞ߩ૞಴
Cre ࡝ࠦࡦࡆ࠽࡯࠯ߣ GFP ࠍ⊒⃻ߔࠆㅒⴕᕈ࡟ࡦ࠴࠙ࠗ࡞ࠬࡌࠢ࠲࡯ࠍࠨ࡞ߩ✢᧦૕ో૕ߦᵈ౉ߒ‫ߩߘޔ‬
ᢙㅳ㑆ᓟߦ‫ޔ‬wild-type -synuclein ߿ᄌ⇣ဳ -synuclein ࠍ⊒⃻ߔࠆ AAV-floxed/stop--synuclein ࠍ㤥⾰ߦᵈ
౉ߒߚࠨ࡞ࠍ૞⵾ߒߚ‫ߦ࡞ࠨߩࠄࠇߎޕ‬೨߽ߞߡ࠻࡟࡯࠾ࡦࠣߒߡ߅޿ߚណ㙄ㆇേ⺖㗴ࠍታⴕߐߖߡ‫ޔ‬ㆇ
േ㓚ኂߩ⒟ᐲࠍ⸃ᨆߔࠆߣߣ߽ߦ‫ࡦࠪࡠ࠴ޔ‬᳓㉄ൻ㉂⚛㧔TH㧕߅ࠃ߮ -synuclein ߩ఺∉ᨴ⦡ߦࠃࠅ㤥⾰
࠼࡯ࡄࡒࡦ࠾ࡘ࡯ࡠࡦߩᄌᕈ࡮⣕⪭ߩ⒟ᐲࠍ⸃ᨆߒߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬ᄌ⇣ဳ -synuclein ߩᣇ߇ wild-type
-synuclein ࠃࠅ߽ലᨐ⊛ߦ⚦⢩ᱫࠍ⺃ዉߢ߈ࠆߎߣࠍ⏕⹺ߒߚ‫ޕ‬
(3) ࡄ࡯ࠠࡦ࠰ࡦ∛㑐ㅪⴕേ⺖㗴ߩ࠻࡟࡯࠾ࡦࠣ
ࡄ࡯ࠠࡦ࠰ࡦ∛ߦ㑐ㅪߔࠆⴕേ⺖㗴ߣߒߡ‫ޔ‬ណ㙄ㆇേ⺖㗴ߦട߃ߡ೔㆐ㆇേ⺖㗴‫ޔߦࠄߐޔ‬Wisconsin Card
Sorting Test㧔WCST㧕߿ࠡࡖࡦࡉ࡞⺖㗴ߥߤߩ⹺⍮⺖㗴ࠍࠨ࡞ߦ࠻࡟࡯࠾ࡦࠣߔࠆߎߣࠍ㐿ᆎߒߚ‫ޕ‬
F) ࠕࠗࠞ࡞࠺ࠖ࡮ࠧ࡯࠹ࠖࠛ∝୥⟲╬ߩࡆࠝࡊ࠹࡝ࡦઍ⻢⇣Ᏹࠍ઻߁∔ᖚߩ⸻ᢿᣇᴺ⏕┙߅ࠃ߮ᴦ≮ᴺ㐿⊒ߩߚ
߼ߩᮮᢿ⊛⎇ⓥ
㜞↰᣽ᒾ㧘৻ἑብ㧔᧲੩Ꮏᬺᄢቇ㧕
ᧄ⎇ⓥߩ⋡⊛ߪ‫∝⊒ߩ∛ࡦ࠰ࡦࠠ࡯ࡄޔ‬ㆊ⒟ߦ߅ߌࠆࡆࠝࡊ࠹࡝ࡦ߅ࠃ߮ߘߩઍ⻢↥‛ߩ㊂⊛ᄌൻࠍ⸃ᨆߔࠆߎ
ߣߦࠃࠅ‫ޔ‬㤥⾰࠼࡯ࡄࡒࡦ⚦⢩ᱫߣࡆࠝࡊ࠹࡝ࡦઍ⻢ߩ⋧੕㑐ଥࠍ᣿ࠄ߆ߦߔࠆߎߣߢ޽ࠅ‫৻ޔ‬ἑࠣ࡞࡯ࡊߣㅪ៤
ߒߡ‫ޔ‬MPTP ᛩਈߦࠃߞߡ૞⵾ߒߚࡄ࡯ࠠࡦ࠰ࡦ∛ࡕ࠺࡞ࠩ࡞߆ࠄ‫ޔ‬ㆇേ㓚ኂߩ⊒⃻ࠍࡕ࠾࠲࡯ߔࠆߣߣ߽ߦ‫⚻ޔ‬
ᤨ⊛ߦ⣖⣄㜑ᶧࠍណขߒ‫ߩ╬ࡦ࡝࠹ࡊࠝࡆޔ‬฽᦭㊂ߣ࠼࡯ࡄࡒࡦ⚦⢩ߩᄌᕈ㨯⣕⪭ߩ⒟ᐲࠍ⺞ߴߚ‫ޕ‬
G) 㔤㐳㘃⣖ߩォ౮࿃ሶㆮવሶ⊒⃻ߣߘߩ⊒㆐ߦ㑐ߔࠆ⎇ⓥ
ᄢ⍹㜞↢㧘ᯫ၂ዊ⊖ว
ᄢ⣖ᣂ⊹⾰ߩ 7 㗔㊁‫ޔ‬ᶏ㚍‫ⵍޔ‬Ზ߅ࠃ߮ዊ⣖ߩ⊒㆐ߦ߅ߌࠆォ౮࿃ሶㆮવሶ⊒⃻⸃ᨆࠍⴕߞߡ޿ࠆ‫ߩߤޕ‬ㇱ૏ߢ
߽⊒⃻߇㕖Ᏹߦૐ޿߽ߩ‫⃻⊒ޔ‬ᄌേ߇߶ߣࠎߤߥ޿߽ߩ‫ߩߤޔ‬ㇱ૏ߢ߽⊒㆐ㆊ⒟ߢห᭽ߥ⊒⃻ᄌൻࠍ␜ߔ߽ߩ‫ޔ‬ㇱ
૏㑆ߢ⊒⃻ᄌൻࡄ࠲࡯ࡦ߇⇣ߥࠆ߽ߩ‫ޔ‬ห৻ㇱ૏ߩⶄᢙࠨࡦࡊ࡞ߢ⊒⃻߇቟ቯߒߥ޿߽ߩ߇޽ߞߚ‫ޕ‬
H) ᗧ᰼ࠍ↢ߺ಴ߔ␹⚻ࡔࠞ࠾࠭ࡓߩ⸃᣿㧦೨㗡೨㊁߳ߩਛ⣖࠼࡯ࡄࡒࡦ౉ജߩᓎഀ
᧻ᧄᱜᐘ㧘㜞↰᣽ᒾ
⋡ᮡࠍ㆐ᚑߒߡႎ㈽ࠍᓧࠃ߁‫ࠍ⟏ߪ޿ࠆ޽ޔ‬ㆱߌࠃ߁ߣ޿߁‫ޟ‬ᗧ᰼‫ߪޠ‬೨㗡೨㊁ߩ௛߈ߩ৻ߟߢ޽ࠆ‫ᦨޕ‬ㄭߩ⎇
㧙㧙
ⓥߪ‫ޔ‬೨㗡೨㊁ߦႎ㈽߿⟏ߦኻߒߡᔕ╵ߔࠆ␹⚻⚦⢩㧔࠾ࡘ࡯ࡠࡦ㧕߇ሽ࿷ߒ‫ࡦࡠ࡯ࡘ࠾ߩࠄࠇߎޔ‬ᔕ╵߇ᗧ᰼ߩ
ࠦࡦ࠻ࡠ࡯࡞ߦ㑐ਈߔࠆߎߣࠍ␜ໂߒߡ޿ࠆ‫ޔߒ߆ߒޕ‬ᗧ᰼ߦ㑐ㅪߒߚ೨㗡೨㊁ߩ␹⚻ᵴേ߇ߤߩࠃ߁ߥࡔࠞ࠾࠭
ࡓߦࠃߞߡ↢ߓࠆߩ߆ߣ޿߁ᩮḮ⊛ߥ໧㗴ߪᧂ⸃᣿ߩ߹߹ߢ޽ࠆ‫ޕ‬
ᧄ⎇ⓥߢߪ‫ࠍࡓ࠭࠾ࠞࡔߩߘޔ‬᣿ࠄ߆ߦߔࠆߚ߼‫ޔ‬ਛ⣖࠼࡯ࡄࡒࡦ࠾ࡘ࡯ࡠࡦ߆ࠄ೨㗡೨㊁ߦવ㆐ߐࠇࠆ␹⚻ࠪ
ࠣ࠽࡞ߦᵈ⋡ߔࠆ‫ߪࡦࡠ࡯ࡘ࠾ࡦࡒࡄ࡯࠼ޕ‬ႎ㈽߿⟏ߦ㑐ㅪߒߚᖱႎࠍࠦ࡯࠼ߒߡ߅ࠅ‫ࠍ࡞࠽ࠣࠪߩߘޔ‬೨㗡೨㊁
ߦવ㆐ߔࠆߎߣߦࠃߞߡ‫ޔ‬ᗧ᰼ߦ㑐ㅪߒߚ೨㗡೨㊁ߩᵴേࠍᒻᚑߔࠆၮ⋚ߣߥߞߡ޿ࠆน⢻ᕈ߇޽ࠆ‫⎇ᧄߢߎߘޕ‬
ⓥߢߪ‫ࠄ߆ࡦࡠ࡯ࡘ࠾ࡦࡒࡄ࡯࠼ޔ‬೨㗡೨㊁ߦߤߩࠃ߁ߥࠪࠣ࠽࡞߇વ㆐ߐࠇߡ޿ࠆߩ߆⺞ߴࠆߎߣࠍ⋡⊛ߣߒ‫ޔ‬
೨㗡೨㊁߇⊒㆐ߒߚࡑࠞࠢዻߩࠨ࡞ࠍታ㛎േ‛ߣߒߡ↪޿ߚ㔚᳇↢ℂታ㛎ࠍ⸘↹ߒߡ޿ࠆ‫ౕޕ‬૕⊛ߦߪ‫⹺ޔ‬⍮⺖㗴
ࠍࠨ࡞ߦⴕࠊߖ‫ߣࡦࡠ࡯ࡘ࠾ࡦࡒࡄ࡯࠼ޔ‬೨㗡೨㊁࠾ࡘ࡯ࡠࡦߩᵴേࠍ⸥㍳ߔࠆ‫⹺ޕ‬⍮⺖㗴ߢߪ‫⺖ޔ‬㗴ߩ㔍ᤃᐲ߿
ႎ㈽㊂ࠍࡄ࡜ࡔ࡯࠲ߣߒߡᠲ૞ߒ‫ߩ࡞ࠨޔ‬ᗧ᰼ࠍࠦࡦ࠻ࡠ࡯࡞ߔࠆ‫ߣࡦࡠ࡯ࡘ࠾ࡦࡒࡄ࡯࠼ޔߡߒߘޕ‬೨㗡೨㊁ߩ
ᵴേࠍᲧセߔࠆߎߣߦࠃࠅ‫ޔ‬ᗧ᰼ߦ㑐ࠊࠆ㔍ᤃᐲ߿ႎ㈽㊂ߩାภ߇࠼࡯ࡄࡒࡦ࠾ࡘ࡯ࡠࡦ߆ࠄ೨㗡೨㊁ߦવ㆐ߐࠇ
ߡ޿ࠆߩ߆ᬌ⸽ߔࠆ‫ޕ‬
ᐔᚑ 23 ᐕᐲߪ‫߇࡞ࠨޔ‬਄ㅀߩ⹺⍮⺖㗴ࠍ߅ߎߥ߃ࠆࠃ߁ߦ⸠✵ߒ‫⚻␹ࠄ߆ࡦࡠ࡯ࡘ࠾ࡦࡒࡄ࡯࠼ޔ‬ᵴേࠍ⸥㍳
ߒߚ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
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✚⺑
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ᦠ⹏
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Shinomiya K, Matsuda K, Oishi T, Otsuna H, Ito K (2011) Flybrain neuron database: a comprehensive database system
of the Drosophila brain neurons. Journal of Comparative Neurology 519(5):807-833.
Tohno Y, Tohno S, Sato H, Hayashi M, Oishi T, Minami T, Mahakkanukrauh P (2011) Gender differences in the
phosphorus content of the sino-atrial nodes and other cardiac regions of monkeys. Biol Trace Elem Res. 143(2):871-881.
Higaki S, Takumi K, Itoh M, Watanabe G, Taya K, Shimizu K, Hayashi M, Oishi T (2012) Response of ER and
aromatase expression in the monkey hippocampal formation to ovariectomy and menopause. Neurosci Res
72(2):148-54.
Hiraoka M, Inoue K, Kawano H, Takada M (2012) Localization of papillofoveal bundles in primates. Anat Rec
295:347-354.
Hiraoka M, Inoue K, Ninomiya T, Takada M (2012) Ischaemia in the Zinn–Haller circle and glaucomatous optic
neuropathy in macaque monkeys. Br J Ophthalmol 96:597-603.
Ninomiya T, Sanada TM, Ohzawa I (2012) Contributions of excitation and suppression in shaping spatial frequency
selectivity of V1 neurons as revealed by binocular measurements. J Neurophysiol 107:2220-2231.
ᄢ⍹㜞↢ (2011) ⣖ߣߎߎࠈ㧙⣖ߩㅴൻ߆ࠄ⠨߃ࠆ㧙ᬉጊੱ㑆ቇ⎇ⓥ 7:82-92.
ᧁ᧛ᵴ↢㧘ጊᧄశ೑ (2012) ᛫ Parkinson ∛⮎ߦࠃࠆ restless legs syndrome ߩᴦ≮㧔⸃⺑/․㓸㧕␹⚻ౝ⑼
(0386-9709) 76(1):67-74.
㜞↰᣽ᒾ (2011) ̌ࠕ࡚ࠢࠪࡦ̍ߩ૞േේℂࠍ⸃߈᣿߆ߔ೨㗡⪲࡟ࡆࡘ࡯ߩவ૞. ㅳೀකቇળᣂ⡞╬.
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
Kato S, Kuramochi M, Kobayashi K, Inoue K, Takada M, Kobayashi K (2011) Viral Gene Therapy (ed. Ke Xu) p.450
Intech, Croatia.
㧙㧙
2)
3)
ᄢ⍹㜞↢ (2011) ‫ߩࠢࠞࡑޟ‬⣖‫ߟ߼߇ߔߟ߼ߚޔ‬, ↢߈‛ߚߜߩߟߠࠇ❱ࠅ, ╙྾Ꮞ‫ ޠ‬p.29-34 ੩ㇺᄢቇࠣࡠ
࡯ࡃ࡞ COE ࡊࡠࠣ࡜ࡓ㧚
㜞↰᣽ᒾ (2011) ‫ޔ੹ޟ‬ㆮવሶᡷᄌࠨ࡞߇ࡎ࠶࠻㧍 ↢߈‛ߚߜߩߟߠࠇ❱ࠅ, ╙੖Ꮞ‫ ޠ‬p.18-24 ੩ㇺᄢቇࠣࡠ
࡯ࡃ࡞ COE ࡊࡠࠣ࡜ࡓ㧚
ቇળ⊒⴫
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Bromberg-Martin ES, Matsumoto M, Hikosaka O (2011) A pallidus-habenula-dopamine pathway transmits both
motivational and visuospatial signals. 41th Annual Meeting of Society for Neuroscience (2011/11/12-16, Washington
DC, USA).
Inoue K, Kato S, Kobayashi K, Takada M (2011) Development in pathway-selective gene delivery and neuronal ablation
with enhanced retrograde transfer of a modified lentiviral vector in primate brain. The 8th IBRO World Congress Of
Neuroscience (2011/07/14-18, Florence, Italy)㧚
Kimura K, Yokochi F, Okiyama R, Aoki Y, Kawasaki T, Taniguchi M (2011) A case of Myoclonus-Dystonia (DYT-11)
improved by bilateral pallidal stimulation. 5th International dystonia symposium (2011/10/20-22, Barcelona, Spain).
Kimura K, Yokochi F, Okiyama R, Aoki Y, Kawasaki T, Taniguchi M (2011) First report of a patient with
Myoclonus-Dystonia (DYT-11) improved by bilateral pallidal stimulation in Japan. 15th International Congress of
Parkinson’s Disease and Movement Disorders (2011/06/05-09, Toronto ON, Canada)㧚
Matsumoto M, Takada M (2011) Midbrain dopamine neurons represent behavioral relevance in a working memory task.
41th Annual Meeting of Society for Neuroscience (2011/11/12-16, Washington DC, USA).
Ninomiya T, Sawamura H, Inoue K, Takada M (2011) Organization of multisynaptic top-down pathways from frontal
cortex to visual areas MT and V4 in macaques. 41th Annual Meeting of Society for Neuroscience (2011/11/12-16,
Washington DC, USA).
Uesaka M, Nishimura O, Uno K, Ueda HR, Oishi T, Imai H, Agata K, Imamura T (2011) Species-specific pseudogene
insertions generate cis-acting RNA for promoter demethylation in the macaque. ╙ 34 ࿁ᣣᧄಽሶ↢‛ቇળᐕળ
(2011/12/16, ᮮᵿ)㧚
⇌ਛિᒾ㧘㊄ሶ዁਽㧘㜞⦟ᴕᐘ㧘㜞↰᣽ᒾ㧘ධㇱ◊ (2011) ㆇേ⺖㗴ㆀⴕਛߩࠨ࡞ߦ߅ߌࠆ᷆⫷⃿࠾ࡘ࡯ࡠࡦ
ᵴേߩࠣ࡞࠲ࡒࡦ㉄߅ࠃ߮ GABA ૞േᕈ⺞▵. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/15, ᮮᵿ)㧚
ᐔ↰ᔟᵗ㧘ች࿾㊀ᐢ㧘ᄢㄼବੑ㧘੹⷏⟤⍮ሶ㧘㤥↰๒ሶ㧘㜞↰᣽ᒾ (2011) ࡜࠶࠻⊹⾰✢᧦૕〝ߩ㔚᳇ೝỗߦ
ࠃߞߡ⋥ធ〝߅ࠃ߮㑆ធ〝࠾ࡘ࡯ࡠࡦߦ c-fos ⊒⃻߇⺃ዉߐࠇࠆ. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/15, ᮮ
ᵿ)㧚
੹᧛ᜏ਽㧘਄ဈ዁ᒄ㧘⷏᧛ℂ㧘ᄢ⍹㜞↢㧘੹੗໪㓶㧘㒙ᒻᷡ๺ (2011) ࡑࠞࠢࠩ࡞ߦ߅ߌࠆனㆮવሶ↱᧪
promoter-associated noncoding RNA (pancRNA) ߦࠃࠆ⒳․⇣⊛ォ౮ᵴᕈൻ. ╙ 152 ࿁ᣣᧄ₞කቇળᄢળ
(2011/09/20, ႓)㧚
੗਄⻞৻㧘➚➓ᄢテ㧘ട⮮ᚑ᮸㧘ዊᨋ๺ੱ㧘ධㇱ◊㧘㜞↰᣽ᒾ (2011) ࠗࡓࡁ࠻ࠠࠪࡦ␹⚻〝ᮡ⊛ᴺߦࠃࠆࠨ
࡞ᄢ⣖ၮᐩᩭࡂࠗࡄ࡯⋥ធ〝ߩㆬᛯ⊛㒰෰. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/15, ᮮᵿ)㧚
દ૒ᱜ㧘ዊፉବ↵㧘⢈ᓟ▸ⴕ㧘ᄢ⍹㜞↢㧘የ਄ᶈ㓉 (2011) ࡑࠞࠢࠩ࡞ㆇേ㑐ㅪ⊹⾰㗔㊁ߦ߅ߌࠆㆮવሶ⊒⃻
ߩࡑࠗࠢࡠࠕ࡟ࠗᴺߦࠃࠆ✂⟜⊛⸃ᨆ. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/17, ᮮᵿ)㧚
દ⮮ືผ㧘㜞↰᣽ᒾ (2011) ࠾ࡎࡦࠩ࡞⡬ⷡ␹⚻ᩭߦ߅ߌࠆࠣ࡞࠲ࡒࡦ㉄‫ޔ‬GABA‫ࡦࠪ࡝ࠣޔ‬૞േᕈ࠾ࡘ࡯ࡠ
ࡦߩಽᏓ. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/15, ᮮᵿ)㧚
ᧁ᧛ᵴ↢㧘ᮮ࿾ᚱሶ㧘ᴒጊ੫৻㧘㕍ᧁካሶ㧘⼱ญ⌀㧘Ꮉፒ㓉㧘⏷የ✍ሶ㧘⍹੗৻ᒾ㧘ᾢ↰⡡ሶ (2011) ࡒࠝࠢ
ࡠ࡯࠿ࠬߦኻߔࠆ DBS ߩലᨐ. ╙ 5 ࿁ᣣᧄࡄ࡯ࠠࡦ࠰ࡦ∛࡮ㆇേ㓚ኂ∔ᖚࠦࡦࠣ࡟ࠬ (2011/10, ᧲੩)㧚
ች࿾㊀ᒄ㧘ᐔ↰ᔟᵗ㧘ᯫ၂ዊ⊖ว㧘㤥↰๒ሶ㧘ችㇱ⾆ሶ㧘㜞↰᣽ᒾ㧘ᄢ⍹㜞↢ (2011) ᐜ⧯ࡑࠞࠢࠨ࡞ᄖ஥೨
㗡೨㊁߳ߩ⊹⾰⊹⾰౉ജ. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/17, ᮮᵿ)㧚
᧛↰ᑿ㧘⢈ᓟ▸ⴕ㧘⷏᧛ᐘ↵㧘ᨋᜏ਽㧘᧖ጊኈሶ㧘ᄢ⍹㜞↢㧘Ⴆ↰⑲ᄦ㧘દ૒ᱜ㧘የ਄ᶈ㓉 (2011) ╙৻ᰴㆇ
േ㊁៊்ᓟߩᛠីേ૞࿁ᓳߦኻߔࠆㆇേ೨㊁⣻஥ㇱߩ㑐ࠊࠅ. ൮᜝⣖ࡀ࠶࠻ࡢ࡯ࠢᄐߩࡢ࡯࡚ࠢࠪ࠶ࡊ
(2011/08/23, ␹ᚭ)㧚
㜞 ↰ ᣽ ᒾ (2011) ࠨ ࡞ ⣖ ߳ ߩ ㆮ વ ሶ ዉ ౉ ᚻ ᴺ ߩ 㐿 ⊒ 㧦 ㆮ વ ሶ ᴦ ≮ ߳ ߩ ᔕ ↪ . ╙ 28 ࿁ ᣣ ᧄ ක ቇ ળ ✚ ળ
(2011/04/10, ᧲੩)㧚
㜞↰᣽ᒾ (2011) ࠼࡯ࡄࡒࡦ⚦⢩ᱫߣᛥ೙ߔࠆᯏ⢻ಽሶߩㆮવሶዉ౉ߦࠃࠆࡄ࡯ࠠࡦ࠰ࡦ∛స᦯ߩߚ߼ߩࠕ
ࡊࡠ࡯࠴. ╙ 26 ࿁ᣣᧄᄢ⣖ၮᐩᩭ⎇ⓥળ (2011/07/02, ▫ᩮ)㧚
Ⴆర⪲ሶ㧘␉᧛ቱ๺㧘੹⷏⟤⍮ሶ㧘Ⴆ↰⒤㧘㜞↰᣽ᒾ (2011) ࡜࠶࠻ᶏ㚍㧛ᄢ⣖⊹⾰ߦ߅ߌࠆ prototypic
afterdischarge ⊒⃻ߩ㗔ၞ․⇣ᕈ. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/15, ᮮᵿ)㧚
ᧁ᧛ᵴ↢ (2012) ⹺⍮∝ߩ⮎‛ᴦ≮㧦ࠛࡆ࠺ࡦࠬ߆ࠄ⠨߃ࠆ. ╙ 11 ࿁㜞᧻ࡄ࡯ࠠࡦ࠰ࡦ∛ࠪࡦࡐࠫ࠙ࡓ
(2012/02/13, 㜞᧻)㧚
㜞↰᣽ᒾ (2012) ೨ㇱᏪ⁁⊹⾰߳ߩ਄ⴕᕈ∩ⷡવዉ〝ߩ᭴▽. ╙ 117 ࿁ᣣᧄ⸃೬ቇળ✚ળ࡮ో࿖ቇⴚ㓸ળ
(2012/03/27, ↲ᐭ)㧚
᧛↰ᑿ㧘⢈ᓟ▸ⴕ㧘ᨋᜏ਽㧘⷏᧛ᐘ↵㧘᧖ጊኈሶ㧘ᄢ⍹㜞↢㧘Ⴆ↰⑲ᄦ㧘દ૒ᱜ㧘የ਄ᶈ㓉 (2011) Plastic changes
in the ventral premotor area after primary motor cortex lesion in macaque monkeys: possible involvement in functional
㧙㧙
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24)
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compensation of manual dexterity. 41st annual meeting of Society for Neuroscience (2011/11/12-16, Washington DC,
USA)㧚
ੑችᄥᐔ㧘Ỉ᧛⵨ᱜ㧘੗਄⻞৻㧘㜞↰᣽ᒾ (2011) ࡑࠞࠢࠩ࡞೨㗡⪲߆ࠄ MT ߅ࠃ߮ V4 ߳ߩᄙࠪ࠽ࡊࠬᕈ౉
ജ᭽ᑼ. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/17, ᮮᵿ)㧚
᧻ᧄᱜᐘ㧘㜞↰᣽ᒾ (2011) ૞ᬺ⸥ᙘ⺖㗴ߦ߅ߌࠆ࠼࡯ࡄࡒࡦ࠾ࡘ࡯ࡠࡦߩ⺖㗴㑐ㅪᕈᔕ╵. 34 ࿁ᣣᧄ␹⚻⑼
ቇᄢળ (2011/09/17, ᮮᵿ)㧚
ዊᨋ๺ੱ㧘ട⮮ᚑ᮸㧘ୖᜬ⌀ੱ㧘ዊᨋᙗᄥ㧘੗਄⻞৻㧘㜞↰᣽ᒾ (2011) 㜞㗫ᐲㅒⴕᕈㆮવሶዉ౉ࡌࠢ࠲࡯ߩ
㔤㐳㘃⣖⑼ቇ߳ߩᔕ↪. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/17, ᮮᵿ)㧚
ୖᜬ⌀ੱ㧘ട⮮ᚑ᮸㧘ዊᨋᙗᄥ㧘㜞૑⾫ม㧘㜞ේᄢテ㧘੗਄⻞৻㧘ፉ↰㓉㧘㜞↰᣽ᒾ㧘ዊᨋ๺ੱ (2011) ⣖ᯏ
⢻⎇ⓥߩߚ߼ߩ␹⚻․⇣⊛ߥ㜞㗫ᐲㅒⴕᕈㆮવሶዉ౉ࡌࠢ࠲࡯ߩ㐿⊒. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/17,
ᮮᵿ)㧚
㜞ේᄢテ㧘ᐔ↰ᔟᵗ㧘ੑችᄥᐔ㧘㜞↰᣽ᒾ (2011) ࠕ࠶ࠨࡓࡕࡦࠠ࡯㧔Macaca assamensis㧕ߦ߅ߌࠆㆇേ㊁ߩ
り૕ㇱ૏ዪ࿷. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/16, ᮮᵿ)㧚
਄ဈ዁ᒄ㧘⷏᧛ℂ㧘ᄢ⍹㜞↢㧘੹੗໪㓶㧘㒙ᒻᷡ๺㧘੹᧛ᜏ਽ (2011) ࡑࠞࠢࠩ࡞ߦ߅ߌࠆனㆮવሶ↱᧪
promoter-associated noncoding RNA (pancRNA) ߦࠃࠆ⒳․⇣⊛ォ౮ᵴᕈൻ. ╙ 104 ࿁ᣣᧄ❥ᱺ↢‛ቇળᄢળ
(2011/09/16, ⋓ጟ)㧚
⻠Ṷ
ᧁ᧛ᵴ↢ (2011/07/01) ࡄ࡯ࠠࡦ࠰ࡦ∛ߩේ࿃ߣᄖ⑼ᴦ≮ߦߟ޿ߡ. 㚅Ꮉ⋵┙ਛᄩ∛㒮␹⚻ౝ⑼. 㜞᧻㧚
᧻ᧄᱜᐘ (2011/10/14) ⣖ߩႎ㈽♽㧦ᄖ஥ᚻ✁ᩭߣ࠼࡯ࡄࡒࡦ࠾ࡘ࡯ࡠࡦߩᓎഀ. ᐔᚑ 23 ᐕᐲࠕ࡞ࠦ࡯࡞࡮⮎
‛ଐሽ㑐ㅪቇળวหቇⴚ✚ળ. ฬฎደ㧚
3) ᧻ᧄᱜᐘ (2011/12/01) ⣖ߩႎ㈽♽㧦ᄖ஥ᚻ✁ᩭߣ࠼࡯ࡄࡒࡦ࠾ࡘ࡯ࡠࡦߩᓎഀ. ✚ว⎇ⓥᄢቇ㒮ᄢቇో㗵ᢎ⢒
੐ᬺ‫↢ޟ‬๮⑼ቇ࡝࠻࡝࡯࠻‫ޠ‬. ដᎹ㧚
4) 㜞↰᣽ᒾ (2011/07/08) ㆮવሶዉ౉ᛛⴚߦࠃࠆࡄ࡯ࠠࡦ࠰ࡦ∛ߩ㔤㐳㘃ࡕ࠺࡞ߩ㐿⊒ߣㆮવሶᴦ≮ߦ㑐ߔࠆၮ
␆⊛ࠕࡊࡠ࡯࠴. ർᶏ㆏ᄢቇᄢቇ㒮⮎ቇ⎇ⓥ㒮. ᧅᏻ㧚
5) 㜞↰᣽ᒾ (2011/10/08) Novel strategies for primate brain research with neurotropic viruses and viral vectors. ╙ 59 ࿁
࿖㓙ᱤ⑼⎇ⓥቇળᣣᧄᡰㇱ㧔JADR㧕✚ળ࡮ቇⴚ㓸ળ. ᐢፉ㧚
6) ᄢ⍹㜞↢ (2011/06/04) ࠨ࡞ࠍቇ߱‫ߢ࡞ࠨޔ‬ቇ߱. ἥ㜞╬ቇᩞ. ␹ᚭ㧚
7) ᄢ⍹㜞↢ (2011/07/12) ࠨ࡞ࡕ࠺࡞ࠍ↪޿ߚ⣖⣄㜑៊்߆ࠄߩㆇേᯏ⢻࿁ᓳߩ⎇ⓥ. 㕍ਤቇ࿦㑐⷏ቇ⎇ක≮⑔
␩ቇ㒮⋴⼔ቇ⑼. ᄹ⦟㧚
8) ᄢ⍹㜞↢ (2011/10/4) ⣖ߣߎߎࠈ 㧙⣖ߩㅴൻ߆ࠄ⠨߃ࠆ㧙. ᬉጊᅚቇ࿦ᄢቇᬉጊੱ㑆ቇ⎇ⓥ࠮ࡦ࠲࡯. ฬฎደ㧚
9) Takada M (2012/02/22)̌Frontiers in Primate Neuroscience Researches̍, Novel approaches to pathway-selective
neuronal manipulation in the primate brain. 㔤㐳㘃⣖⑼ቇࠪࡦࡐࠫ࠙ࡓ. ᧲੩㧚
10) ੗਄⻞৻㧘㜞↰᣽ᒾ (2012/01/14) 㔤㐳㘃ࡄ࡯ࠠࡦ࠰ࡦ∛ࡕ࠺࡞ߩ૞⵾ߣหࡕ࠺࡞ࠍ↪޿ߚㆮવሶᴦ≮⎇ⓥ. ␹
⚻∔ᖚߩࡕ࠺࡞േ‛⎇ⓥળ. ᄢ㒋㧚
11) ᧁ᧛ᵴ↢ (2012/01/31) PD ߣ Movement Disorder ߦ߅ߌࠆᆫ൓⇣Ᏹ㧦ၮ␆කቇߣ⥃ᐥࠍߟߥߋᦨㄭߩ⍮⷗. ᮮᵿ
ࡄ࡯ࠠࡦ࠰ࡦ∛ᴦ≮⎇ⓥળ㧔YPD㧕. ᮮᵿ㧚
12) ੑችᄥᐔ (2012/01/06) ࡑࠞࠢࠩ࡞ߩ MT ߅ࠃ߮ V4 ߳ߩᄙࠪ࠽ࡊࠬᕈ౉ജ᭽ᑼ. ⥄ὼ⑼ቇ⎇ⓥᯏ᭴↢ℂቇ⎇ⓥ
ᚲ. ጟፒ. ᐔᚑ 23 ᐕᐲ↢ℂቇ⎇ⓥᚲ⎇ⓥળ㧚
1)
2)
ㆮવሶᖱႎಽ㊁
㧨⎇ⓥ᭎ⷐ㧪
A) ࠥࡁࡓਇᲫㆃṛ㧔RCRO㧕ߩㅴൻߣᗧ⟵
ᐔ੗໪ਭ㧘ฎ⾐┨ᒾ㧔ࠥࡁࡓᄙ᭽ᕈಽ㊁㧕㧘ᐔ੗⊖วሶ㧔ᛛ⢻⵬㧕㧘㡻Ლବผ㧔ᾢᧄࠨࡦࠢ࠴ࡘࠕ࡝㧕㧘᧻ᨋᷡ᣿㧔ฬ
⹷ᢎ᝼㧕
RCRO ߩ᭴ᚑⷐ⚛ߩ߭ߣߟߢ޽ࠆࠨࡉ࠲࡯ࡒ࠽࡞ࠨ࠹࡜ࠗ࠻㧔StSat㧕ࠍࠢࡠ࡯ࡦൻߒ‫ ߥ⚦⹦ޔ‬FISH ⸃ᨆࠍ߅ߎ
ߥߞߚ‫࡯ࠫࡦࡄࡦ࠴ޕ‬43 ୘૕ߩᨴ⦡૕ࠍ⸃ᨆߒ‫ޔ‬ᄌ⇣ߩ⁁ᴫࠍ᣿ࠄ߆ߦߒߚ‫ޕ‬ᷫᢙಽⵚ♖Უ⚦⢩ߩᨴ⦡૕ࡉ࡯ࠤ
ߣߩ㑐ㅪ߆ࠄ RCRO ߩሽ࿷ᗧ⟵ࠍ⼏⺰ߒ‫⺰ޔ‬ᢥߣߒߡ߹ߣ߼ߚ‫ޕ‬
B) ࠹࠽ࠟࠩ࡞㘃ߩᄙ᭽ᕈߣ♽⛔↢‛࿾ℂቇ
ᐔ੗໪ਭ㧘ࠬ࠳࡜ࠬ࡮ࡃࠗ࠴ࡖࡠࠛࡦ㧔࠲ࠗേ‛࿦දળ㧘ࠞ࠮ࠨ࡯࠻ᄢቇ㧕㧘ࠗࠬ࡜࠻࡮ࠫࡖࡂࡦ㧔ᄢቇ㒮↢㧕㧘ฎ
⾐┨ᒾ㧔ࠥࡁࡓᄙ᭽ᕈ㧕㧘ᐔ੗⊖วሶ㧔ᛛ⢻⵬㧕㧘᧻੗ᷕ㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯⎇ⓥຬ㧕
ࡒ࠻ࠦࡦ࠼࡝ࠕࠥࡁࡓోႮၮߩ⸃ᨆ߆ࠄ‫ޔ‬Hoolock hoolock ࠍട߃ߡ࠹࠽ࠟࠩ࡞ 4 ዻߩಽሶ♽⛔ಽጘࠍ᣿ࠄ߆ߦ
ߒߚ‫ߩࡦࡑࠕࠪޕ‬ᨴ⦡૕ᧃ┵ߦሽ࿷ߔࠆࡋ࠹ࡠࠢࡠࡑ࠴ࡦߩ DNA ࠍ⸃ᨆߒ‫ޔ‬4 ዻߦ߅޿ߡߘࠇߙࠇ․⇣⊛ߥᨴ⦡
૕਄ಽᏓࡄ࠲࡯ࡦࠍ๒ߔࠆߎߣࠍ᣿ࠄ߆ߦߒߚ‫ࠕࡔࡠ࠹ߣࠕࡔࡠ࠻ࡦ࠮ޕ‬๟ㄝߩࡋ࠹ࡠࠢࡠࡑ࠴ࡦߩᒻᚑߦ㑐ࠊࠆ
DNA ߩ㑐ㅪᕈߦߟ޿ߡ⸃ᨆߒߚ‫ޕ‬
㧙㧙
C) ࡑࡦ࠰ࡦ૑ⴊๆ⯻ߩᕈᨴ⦡૕㧔Z‫ޔ‬W㧕ߩㅴൻ
ᐔ੗໪ਭ㧘ᐔ੗⊖วሶ㧔ᛛ⢻⵬㧕㧘ࡈࠖ࡝࠶ࡊ࡮ࡠࡌ࡞࠺㧔࠹ࠠࠨࠬᄢቇ㧕
BAC mapping ߦࠃߞߡ᣿ࠄ߆ߦߥߞߚࠢࡠ࡯ࡦߩሽ࿷᭽ᑼ߆ࠄ‫ޔ‬Z ߅ࠃ߮ W ᕈᨴ⦡૕ߩㅴൻ߇ 4 ࿁ߩㅒ૏ߦࠃ
ߞߡ↢ߓߚߎߣࠍផቯߒߚ‫ోޔߚ߹ޕ‬ᨴ⦡૕ߩᓀ⦡ࡊࡠ࡯ࡉࠍ૞ᚑߒ‫ޔ‬Z ߣ W ᨴ⦡૕ߩ⋧ห߅ࠃ߮㕖⋧หㇱ૏ࠍ
᣿ࠄ߆ߦߒߚ‫⺰ޕ‬ᢥߣߒߡ߹ߣ߼ߚ‫ޕ‬
D) ࠴ࡦࡄࡦࠫ࡯⧰๧ฃኈ૕ߩᄙဳ⸃ᨆ
ᣧᎹථᔒ㧘⩲ේ੧㧔⃻ ᚑ⢒ක≮࠮ࡦ࠲࡯㧕
㧘㡻Ლବผ㧘᫪᧛ᚑ᮸㧔એ਄㧘ᾢᧄࠨࡦࠢ࠴ࡘࠕ࡝㧕
㧘෹᳗㓷Ꮖ㧔ᕁ⠨⸒⺆㧕
㧘
ᄢ᧲⡸㧔⑔੗⋵┙ᄢቇ㧕
㧘ㇹᐽᐢ㧘ᐔ੗໪ਭ㧘੹੗໪㓶
࿖ౝᣉ⸳㘺⢒࠴ࡦࡄࡦࠫ࡯ࠍኻ⽎ߦ TAS2R ㆮવሶ⟲ߩ⒳ౝᄙဳࠍ⸃ᨆߒߚ‫ ⚂ޕ‬50 ୘૕ࠍኻ⽎ߦߒߚ⷏࠴ࡦࡄࡦ
ࠫ࡯ߢߪᐔⴧㆬᛯ⊛ߥ௑ะ߇ߺࠄࠇߚ㧔Sugawara et al.,2011㧕
‫৻ޕ‬ᣇ‫ޔ‬േ‛࿦╬߆ࠄ㓸߼ߡ⸃ᨆߒߚ᧲࡮ਛᄩ࠴ࡦࡄ
ࡦࠫ࡯ߦߟ޿ߡߪ‫ߚߞߥ⇣ߣ࡯ࠫࡦࡄࡦ࠴⷏ޔ‬௑ะࠍ␜ߔߎߣ߇᣿ࠄ߆ߦߥߞߚ‫ޕ‬ㅴൻ⊛࡮↢ᘒ⊛ߥᗧ⟵ߦߟ޿ߡ‫ޔ‬
⎇ⓥࠍㅴ߼ߡ޿ࠆ‫ޕ‬
E) ࡑࠞࠢ㘃ߩ⧰๧ฃኈ૕ߩᄙဳ⸃ᨆ
㋈ᧁධ⟤㧘⩲ේ੧㧔⃻ ᚑ⢒ක≮࠮ࡦ࠲࡯㧕
㧘᧻੗ᷕ㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯㧕㧘᧻Ꮉື਽㧔ㄭ⇰ᄢቇ㧕㧘ㇹᐽ
ᐢ㧘ᐔ੗໪ਭ㧘੹੗໪㓶
ฦ࿾ߩ࠾ࡎࡦࠩ࡞ߦߟ޿ߡ⧰๧ฃኈ૕ TAS2R ߩㆮવሶᄙဳ⸃ᨆࠍⴕߞߚ‫ ߦ․ޕ‬TAS2R38 ߦߟ޿ߡᄙߊߩㆮવሶ
ᄙဳ߇⊒⷗ߐࠇ‫♿ޔ‬દඨፉ಴りߩ⟲ߢ․⇣⊛ߦᯏ⢻ࠍᰳ៊ߒߡ޿ࠆߎߣ߇ಽ߆ߞߚ‫ߩߎޕ‬ᄌ⇣߇↢ߓߚᐕઍផቯߣ
ߘࠇߦ㑐ㅪߒߚ߆ࠎ߈ߟ㘃ߩᬀ↢╬ߦߟ޿ߡ⸃ᨆࠍㅴ߼ߡ޿ࠆ‫ޕ‬
F) ࡑࠞࠢ㘃ߩ⧰๧ฃኈ૕ߩ⊒⃻⸃ᨆ
੹੗໪㓶㧘㋈ᧁධ⟤㧘ષᎹ⟤Ⓞ㧔ࠣࡠ࡯ࡃ࡞ COE㧕
㧘᪉੗ᢘዷ㧘⍹ਣ༑ᦶ㧘ਃဈᏁ㧘㒙ㇱ໪ሶ㧔એ਄᧲ᄢ㒮ㄘ↢⑼㧕
⚦⢩࡟ࡌ࡞ߢࠞ࡞ࠪ࠙ࡓࠗࡔ࡯ࠫࡦࠣᴺߦࠃࠅࡅ࠻ߣࡑࠞࠢߩ⧰๧ฃኈ૕ߩᲧセᯏ⢻⸃ᨆࠍⴕߞߚ‫ޔߚ߹ޕ‬ኻᔕ
ߔࠆⴕേታ㛎ࠍⴕߞߚ‫⚿ߩߘޕ‬ᨐ‫ߩࠡ࠽ࡗߪߢࠢࠞࡑޔ‬᮸⊹ߦ฽߹ࠇࠆࠨ࡝ࠪࡦߦኻߔࠆ⧰๧ᗵฃᕈ߇ࡅ࠻ࠃࠅ߽
10 ୚એ਄ૐ޿ߎߣ߇᣿ࠄ߆ߦߥߞߚ㧔Imai et al.,2012㧕
‫⚿ߩߎޕ‬ᨐߪ‫߇࡞ࠩࡦࡎ࠾ޔ‬౻ቄߦࡗ࠽ࠡ╬ߩ᮸⊹ࠍ㘩ߴࠆ
ߎߣߣ㑐ଥ߇޽ࠆน⢻ᕈࠍ␜ໂߒߡ޿ࠆ‫ޕ‬
G) ࠦࡠࡉࠬ㘃ߩ⧰๧ฃኈ૕ߣណ㘩ߩ㑐ଥ
㋈ᧁධ⟤㧘ዊᴰᢘᒾ㧔ℂቇㇱቇㇱ↢㧕㧘᧛਄ᄩᒎ㧔ㄘቇㇱቇㇱ↢㧕㧘Yin Lijie㧘Pan Wenshi㧔એ਄ർ੩ᄢቇ㧕㧘ષᎹ⟤
Ⓞ㧔ࠣࡠ࡯ࡃ࡞ COE㧕
㧘੹੗໪㓶
ਛ࿖ᐢ⷏࠴ࡢࡦᣖ⥄ᴦ඙ፏᏀᏒߢⷰኤߐࠇߡ޿ࠆ white-headed langur ߦߟ޿ߡ‫ޔ‬ណ㘩ᵴേߣ๧ⷡߩ㑐ଥࠍᬌ⸛ߔ
ࠆߚ߼ߦ౒ห⎇ⓥࠍㅴ߼ߡ޿ࠆ‫ޕ‬ᤓᐕᐲߦᒁ߈⛯߈‫ޔ‬ណ㘩ᬀ‛ߩ⺞ᩏߣࡈࡦ߆ࠄߩ DNA ಽᨆࠍⴕߞߚ‫ޕ‬
H) ᶖൻེߦ⊒⃻ߔࠆ๧ⷡᖱႎવ㆐♽ߩត⚝
ᮭ↰ᓀ㧘᧻᧛⑲৻㧔એ਄ጘ㒂ᄢቇ㧕
㧘ㇹᐽᐢ㧘੹੗໪㓶
㔤㐳㘃ߩᶖൻེߦ⊒⃻ߒߡ޿ࠆ๧ⷡᖱႎવ㆐㑐ㅪ࠲ࡦࡄࠢ⾰ߦߟ޿ߡ‫ޔ‬ඨቯ㊂⊛ RT-PCR ᴺߦࠃࠅᬌ⸛ߒߚ‫ߘޕ‬
ߩ⚿ᨐ‫ޔ‬⣺▤♽ߢߩ⊒⃻ࡄ࠲࡯ࡦߦߟ޿ߡ⒳Ꮕ߇ⷰኤߐࠇߚ‫ޕ‬
I) ⣖ᯏ⢻ߦ㑐ࠊࠆㆮવሶߩᄙဳ⸃ᨆ
ષᎹ⟤Ⓞ㧔ࠣࡠ࡯ࡃ࡞ COE㧕
㧘ᯅᧄ੫ᄥ㧔ᄢ㒋ᄢቇ㧕
㧘੹੗໪㓶
⣖ᯏ⢻ߣ⥄ὼ⊒↢⊛ㆮવሶᄌ⇣ߣߩ㑐ㅪࠍᬌ⸛ߔࠆߚ߼‫ޔ‬᭽‫ߥޘ‬⣖ᯏ⢻ߦ㑐ࠊࠆㆮવሶߩᄙဳ⸃ᨆߦ⌕ᚻߒߚ‫ޕ‬
੹ᐕᐲߪࠞ࠹ࠦ࡯࡞ࡔ࠴࡞ၮォ⒖㉂⚛ㆮવሶ㧔COMT㧕ߩᄙဳ⸃ᨆࠍਥߦⴕߞߚ‫ޕ‬
J) ࠴ࡦࡄࡦࠫ࡯ߩᲧセࠥࡁࡓ࡮Ყセ࠻࡜ࡦࠬࠢ࡝ࡊ࠻࡯ࡓ⸃ᨆ
ㇹᐽᐢ㧘⼾↰ᢕ㧔ㆮવᚲ㧕㧘ㄖᧄ዁ม㧔ㆮવ⎇㧕㧘⮮ጊ⑺૒ᄦ㧔ㆮવ⎇㧕㧘㤥ᧁ㓁ሶ㧔ℂ⎇㧕㧘ᐔ੗໪ਭ㧘෹᳗㓷Ꮖ㧔ᕁ
⠨⸒⺆㧕
㧘᧻ᴛື㇢㧔ᕁ⠨⸒⺆㧕㧘⷏᧛ℂ㧔੩ᄢℂ࡮ࠣࡠ࡯ࡃ࡞ COE㧕
㧘㒙ᒻᷡ๺㧔੩ᄢℂ࡮↢‛‛ℂ㧕
ࡅ࠻ߩㅴൻࠍ⠨߃ࠆ਄ߢ‫߽ᦨޔ‬ㄭ✼⒳ߢ޽ࠆ࠴ࡦࡄࡦࠫ࡯ߩࠥࡁࡓ⸃ᨆ߅ࠃ߮࠻࡜ࡦࠬࠢ࡝ࡊ࠻࡯ࡓ⸃ᨆߪᔅ㗇
ߢ޽ࠆ‫ޕ‬㔤㐳㘃⎇ⓥᚲߩ࠴ࡦࡄࡦࠫ࡯ⷫሶ࠻࡝ࠝߩ⊕ⴊ⃿⚦⢩߅ࠃ߮ࡅ࠻ߩ࠮࡞࡜ࠗࡦࠍ↪޿ߡᰴ਎ઍࠪ࡯ࠤࡦࠨ
࡯ߦࠃࠆ⊒⃻ቯ㊂ൻࠍⴕߥߞߚ‫ߚ߹ޕ‬࿖┙ㆮવቇ⎇ⓥᚲߣߩ౒ห⎇ⓥߦࠃࠅⷫሶ࠻࡝ࠝߩోࠥࡁࡓ⸃ᨆࠍߔߔ߼ߡ
߅ࠅ‫ࠆߥ⇣ޔ‬ᰴ਎ઍࠪ࡯ࠤࡦࠨ࡯㧔SOLiD4 ߣ HiSeq2000㧕ߦࠃࠆⷫሶ 3 ୘૕ߩోࠥࡁࡓ⸃ᨆࠍⴕߞߡ޿ࠆ‫ޕ‬
K) 㔤㐳㘃ߦ߅ߌࠆ⣖Ყセࠝࡒ࠶ࠢࠬ⎇ⓥ
ㇹᐽᐢ㧘੗਄⻞৻㧔⛔ว⣖㧕
㧘ᄢ⍹㜞↢㧔⛔ว⣖㧕
㧘ᷰᚒㇱᤘ຦㧔ၮ↢⎇㧕
㧘㊀ା⑲ᴦ㧔ၮ↢⎇㧕
㧘ጊ᫪ື㓶㧔ၮ↢⎇㧕
㧘
㇊ᵄብਯ㧔ᣂẟᄢ⣖⎇㧕
㧘ᩑ↰᣿⟤㧔ᣂẟᄢ⣖⎇㧕
㧘㜞↰᣽ᒾ㧔⛔ว⣖㧕
㧘ᐔ੗໪ਭ
ࡅ࠻ࠄߒߐࠍᡰ߃ࠆ⣖ࠥࡁࡓၮ⋚ߩ⸃᣿ࠍ⋡ᜰߒߡ‫ߩ࡞ࠩࠢࠞࡑޔ࡞ࠩࠟ࠽࠹ޔ࡜࡝ࠧޔ࡯ࠫࡦࡄࡦ࠴ޔ࠻ࡅޔ‬
ᱫᓟ⣖ࠃࠅᄢ⣖ᣂ⊹⾰ࠍਛᔃߦⶄᢙ㗔㊁߆ࠄ DNA ߅ࠃ߮ RNA ࠍᯏ⢻㗔㊁޽ࠆ޿ߪᯏ⢻࠾ࡘ࡯ࡠࡦන૏ߢขᓧߒ‫ޔ‬
ᰴ਎ઍࠪ࡯ࠤࡦࠨ࡯ߦࠃࠆ✂⟜⊛࠻࡜ࡦࠬࠢ࡝ࡊ࠻࡯ࡓ⸃ᨆ߅ࠃ߮ࡔ࠴ࡠ࡯ࡓ⸃ᨆ㧔ࠥࡁࡓࡢࠗ࠼ࡔ࠴࡞ൻ⸃ᨆ㧕
ࠍⴕߞߡ޿ࠆ‫ޕ‬
㧙㧙
L) ࡑࠞࠢࠩ࡞ߦ߅ߌࠆࠛࠢ࠰࡯ࡓ⸃ᨆ
ㇹᐽᐢ㧘⼾↰ᢕ㧔ㆮવᚲ㧕
㧘੹੗໪㓶㧘ጊ᫪ື㓶㧔ၮ↢⎇㧕
㧘દ૒ᱜ㧔↢ℂ⎇㧕
㧘ᐔ੗໪ਭ
ࡑࠞࠢࠩ࡞ߩታ㛎േ‛ൻߦะߌߚᦨೋߩ⹜ߺߣߒߡ‫ޔ‬ታ㛎ߦଏਈߐࠇࠆ୘૕⟲ߩㆮવ⊛ࡃ࠶ࠢࠣ࡜࠙ࡦ࠼ࠍᛠី
ߔࠆᔅⷐ߇޽ࠆ‫ ࡞ࠩࠥࠞࠕ߮ࠃ߅࡞ࠩࡦࡎ࠾ޕ‬48 ୘૕ࠍ↪޿ߡ‫↪ࠍ࠻࠶ࠠࡓ࡯࠰ࠢࠛߚࠇߐࡦࠗࠩ࠺ߦ↪࠻ࡅޔ‬
޿ߚࠛࠢ࠰࡯ࡓ⸃ᨆࠍⴕߞߡ޿ࠆ‫ޕ‬
M) ․ᱶߥⅣႺߦㆡᔕߒߚ߶੃㘃ߩ༦ⷡฃኈ૕ㆮવሶ⟲ߩㆡᔕㅴൻ
ㇹᐽᐢ㧘ᣂ᧛⧐ੱ㧔᧲੩ක⑼ᱤ⑼ᄢ㧕㧘㘖↰⪲ሶ㧔✚⎇ᄢ㧕㧘ਭ㊁㚅㧔✚⎇ᄢ㧕㧘㜞⇌ዏਯ㧔✚⎇ᄢ㧕
ㅴൻߩㆊ⒟ߢ․ᱶߥⅣႺߦㆡᔕߒߚ↢‛ߦߪ‫ߩߘޔ‬ⅣႺߦᔕߓߚ⴫⃻ဳߩ․ᱶൻ߇ߒ߫ߒ߫ⷰኤߐࠇࠆ‫․ߩߎޕ‬
ᱶൻߦ㓙ߒߡ⿠߈ࠆಽሶ࡟ࡌ࡞ߩᄌൻࠍតࠆߚ߼ߦ‫ޔ‬ᶏ᫮ㆡᔕ߽ߒߊߪ㘧⠍⢻ജࠍ₪ᓧߒߚ߶੃㘃ߦ߅ߌࠆ༦ⷡฃ
ኈ૕ㆮવሶߩㆡᔕㅴൻߩㆊ⒟ࠍ⺞ߴߚ‫ޕ‬
N) ࠗ࡞ࠞߩ⧰๧ฃኈ૕ㆮવሶߩࠥࡁࡓ⸃ᨆ
ㇹᐽᐢ㧘ᵻᎹୃ৻㧔᧲ᄢ㧕㧘ᷡ᳓ෘᔒ㧔ᘮᔕᄢ㧕㧘૒‫⾆ᧁޘ‬ผ㧔ᘮᔕᄢ㧕㧘ᷡ᳓ା⟵㧔ᘮᔕᄢ㧕
ᶏ᫮ㆡᔕߒߚࠗ࡞ࠞ㘃ߦ߅ߌࠆ๧ⷡฃኈ૕ㆮવሶߩㆮવሶㅴൻࠍ⺞ߴࠆߚ߼ߦ‫ ࠞ࡞ࠗޔ‬BAC ࡜ࠗࡉ࡜࡝࡯ࠃࠅ
T2R ㆮવሶ⟲߇ሽ࿷ߔࠆ BAC ࠢࡠ࡯ࡦࠍหቯߒ‫ޔ‬㈩೉⸃ᨆࠍⴕߥߞߚ‫ޔߚ߹ޕ‬หᤨߦ⸃ᨆ߇ㅴⴕߒߡ޿ࠆోࠥࡁ
ࡓ㈩೉ࠍ೑↪ߒߚ in silico ⸃ᨆ߽ⴕ޿‫ޔ‬ታ㛎ߢᓧߚ࠺࡯࠲ߣᲧセࠍⴕߥߞߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬หቯߒߚ㈩೉ߔߴߡ߇ᯏ
⢻ࠍ༚ᄬ㧔னㆮવሶൻ㧕ߒߡ޿ࠆߎߣ߇ಽ߆ߞߚ‫ޕ‬
O) ࡚࡚ࠪ࠙ࠫ࠙ࡃࠛߦ߅ߌࠆᲧセ࠻࡜ࡦࠬࠢ࡝ࡊ࠻࡯ࡓ⸃ᨆ
ㇹᐽᐢ㧘P Fontanillas (Broad Institute)㧘D Hartl (Harvard ᄢ)
⒳߿ᕈߦ߅ߌࠆ⴫⃻ဳߩ㆑޿ࠍ↢ߺ಴ߔ RNA ࡟ࡌ࡞ߢߩᯏ᭴ࠍ⺞ߴࠆߚ߼ߦ‫࡟ࠕࡠࠢࠗࡑߥ⊛⇣․ࡦ࠰ࠢࠛޔ‬
ࠗࠍ૞ᚑߒ‫ࠛࡃ࡚࡚࠙ࠫ࠙ࠪࡠࠗࠠޔ‬㧔Drosophila melanogaster㧕
ߣߘߩㄭ✼ 2 ⒳ߦ߅ߌࠆㆮવሶ⊒⃻ᄌൻࠍ⺞ߴߚ‫ޕ‬
ߘߩ⚿ᨐ‫ޔ‬ᕈ․⇣⊛ߥㆮવሶ⊒⃻ࡄ࠲࡯ࡦࠍ␜ߔㆮવሶࠍᄙᢙᬌ಴ߒߚ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
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7)
Katoh I, Míróva A, Kurata S, Murakami Y, Horikawa K, Nakakuki N, Sakai T, Hashimoto K, Maruyama A, Yonaga T,
Fukunishi N, Moriishi K, Hirai H (2011) Activation of the long terminal repeat of human endogenous retrovirus K by
melanoma-specific transcription factor MITF-M. Neoplasia 13(11):1081-1092.
Koga A, Notohara M, Hirai H (2011) Evolution of subterminal satellite (StSat) repeats in hominids. Genetica
139:167-175.
Lawton SP, Hirai H, Ironside JE, Johnstone DA, Rollinson D (2011) Genomes and geography: genomic insights into the
evolution and phylogeography of the genus Schistosoma. Parasites & Vectors 10:131 doi:10.1186/1756-3305-4-131.
Nagai H, Terai Y, Sugawara Y, Imai H, Nishihara H, Hori M, Okada N (2011) Reverse evolution in RH1 for adaptation
of cichlids to water depth in Lake Tanganyika. Mol. Biol. Evol. 28:1769-1776.
Takaki A, Yamazaki A, Maekawa T, Shibata H, Hirayama K, Kimura A, Hirai H, Yasunami M (2011) Positive selection
of Toll-like receptor 2 polymorphisms in two closely related old world monkey species, rhesus and Japanese macaques.
Immunogenetics DOI:10.1007/S00251-011-0556-2.
Yoshida K, Terai Y, Mizoiri S, Aibara M, Nishihara H, Watanabe M, Kuroiwa A, Hirai H, Hirai Y, Matsuda Y, Okada
N (2011) B chromosomes have a functional effect on femal sex determination in lake Victoria cichlid fishes. PLoS
Genetics 7(8):e1002203.
Katayama K, Furutani Y, Imai H, Kandori H (2012) Protein-Bound Water Molecules in Primate Red- and
Green-Sensitive Visual Pigments. Biochemistry 51:1126-1133.
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
2)
Hirai H (2012) Evolution and biological meaning genomic wastelands (RCRO): Proposal of hypothesis. (Post-Genome
Biology of Primates.) (ed. Hirai H, Imai H, Go Y) p.227-240 Springer.
Sugawara T, Imai H (2012) Post-Genome Biology of Primates Focusing on Taste Perception. (Post-Genome Biology of
Primates) (ed. Hirai H, Imai H, Y. Go) p.79-92 Springer.
✬㓸
1)
Hirai H, Imai H, Go Y (2012) Post-Genome Biology of Primates, Primatology Monographs. p.286 Tokyo, Dordrecht,
Heidelberg, London, New York: Springer.
ቇળ⊒⴫
1)
2)
3)
Go Y (2011) Comparative transcriptome and genome analysis in a chimpanzee trio. International Symposium of
Developmental Systems Biology on Gene Regulation and Aging (2011/10/13, Shanghai, CHINA㧔᜗ᓙ⻠Ṷ㧕)㧚
Go Y (2011) Comparative transcriptome and genome analysis in a chimpanzee trio. Young Researchers Conference on
Evolutionary Genomics (2011/08/02, Tokyo).
Imai H (2011) Functional evolution of primate TAS2Rs, ࠪࡦࡐࠫ࠙ࡓ‫ޟ‬ⅣႺㆡᔕߩᦨ೨✢㧦ᗵⷡฃኈ૕ߩᯏ⢻ㅴ
ൻ‫ޠ‬Adaptive molecules: functional evolution of sensory receptors. ᣣᧄㅴൻቇળ╙ 13 ࿁ᄢળ (2011/07/30, ੩ㇺ)㧚
㧙㧙
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⻠Ṷ
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2)
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Suzuki N, Matsui A, Go Y, Ishimaru Y, Misaka T, Abe K, Hirai H, Imai H (2011) Region specific dysfunction of bitter
taste receptor TAS2R38 in Japanese macaques. The 9th International Symposium on Molecular and Neural Mechanisms
of Taste and Olfactory Perception (2011/11/06, Fukuoka).
ㇹᐽᐢ (2011) ࠥࡁࡓࠍㅢߒߡᚒ߇りࠍ⍮ࠆ㨪ࡅ࠻ߣ࠴ࡦࡄࡦࠫ࡯ߩ㑆ߦ޽ࠆ߽ߩ㨪. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇ
ળᄢળ (2011/07/18, ›ጊ)㧚
ㇹᐽᐢ㧘⼾↰ᢕ㧘ળᵤᥓᐘ㧘੹੗໪㓶㧘⮮ጊ⑺૒ᄦ㧘ᐔ੗໪ਭ (2011) ࠾ࡎࡦࠩ࡞ࠛࠢ࠰࡯ࡓ⸃ᨆ㨪ታ㛎േ‛
ൻߦ߻ߌߚㆮવ⊛ࡃ࠶ࠢࠣ࡜࠙ࡦ࠼ߩ⸃᣿㨪. ╙ 13 ࿁ᣣᧄㅴൻቇળᄢળ (2011/07/30, ੩ㇺ)㧚
ㇹᐽᐢ㧘⼾↰ᢕ㧘ળᵤᥓᐘ㧘੹੗໪㓶㧘⮮ጊ⑺૒ᄦ㧘ᐔ੗໪ਭ (2011) ࠾ࡎࡦࠩ࡞ࠛࠢ࠰࡯ࡓ⸃ᨆ㨪ታ㛎േ‛
ൻߦ߻ߌߚㆮવ⊛ࡃ࠶ࠢࠣ࡜࠙ࡦ࠼ߩ⸃᣿㨪. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/18, ›ጊᏒ)㧚
ᣧᎹථᔒ㧘⩲ේ੧㧘ㇹᐽᐢ㧘㡻Ლବผ㧘ᐔ੗໪ਭ㧘੹੗໪㓶 (2011) ࠴ࡦࡄࡦࠫ࡯3 ੝⒳ߦ߅ߌࠆ⧰๧ฃኈ૕
ㆮવሶࡈࠔࡒ࡝࡯ߩಽሶㅴൻ. ᣣᧄㅴൻቇળ╙ 13 ࿁ᄢળ (2011/07/30, ੩ㇺ)㧚
ᣧᎹථᔒ㧘⩲ේ੧㧘ㇹᐽᐢ㧘㡻Ლବผ㧘ᐔ੗໪ਭ㧘੹੗໪㓶 (2011) ࠴ࡦࡄࡦࠫ࡯ߩ๧ⷡߦ࿾ၞᏅߪ޽ࠆ߆㧫
㨪ಽሶㆮવቇ߆ࠄߩ⠨ኤ㨪. SAGA14 (2011/11/12, ᾢᧄ)㧚
ᣧᎹථᔒ㧘⩲ේ੧㧘ㇹᐽᐢ㧘㡻Ლବผ㧘ᐔ੗໪ਭ㧘੹੗໪㓶 (2011) ࠴ࡦࡄࡦࠫ࡯3 ੝⒳ߦ߅ߌࠆ⧰๧ฃኈ૕
ㆮવሶࡈࠔࡒ࡝࡯ߩಽሶㅴൻ. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/17, ›ጊ)㧚
ᐔ੗໪ਭ㧘ᐔ੗⊖วሶ㧘ฎ⾐┨ᒾ㧘㡻Ლବผ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ޽ߞߡࡅ࠻ߦߥ޿ᨴ⦡૕┵ㇱࠥࡁࡓਇᲫ
࿾ Ꮺ :(1) ሽ ࿷ ᭽ ᑼ ᄌ ⇣ ߆ ࠄ ផ ᷹ ߐ ࠇ ࠆ 㕖 ⋧ ห ᨴ ⦡ ૕ 㑆 ᧃ ┵ ⚵ ឵ ߃ . ╙ 27 ࿁ ᣣ ᧄ 㔤 㐳 㘃 ቇ ળ ᄢ ળ
(2011/07/16-18, ›ጊᏒ)㧚
ᐔ੗໪ਭ㧘ᐔ੗⊖วሶ㧘ฎ⾐┨ᒾ㧘㡻Ლବผ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ޽ߞߡࡅ࠻ߦߥ޿ᨴ⦡૕┵ㇱࠥࡁࡓਇᲫ
࿾Ꮺ㧦ሽ࿷᭽ᑼᄌ⇣. ╙ 62 ࿁ᨴ⦡૕ቇળ (2011/11/11-13, ᐔႦ)㧚
੹੗໪㓶㧘ㇹᐽᐢ㧘ᐔ੗໪ਭ (2011) 㔤㐳㘃ࠥࡁࡓࠬࠢ࡝࡯࠾ࡦࠣߦࠃࠆ⥄ὼ⊒↢⊛ㆮવሶᄌ⇣ࡕ࠺࡞ߩត⚝.
╙ 34 ࿁ᣣᧄ␹⚻⑼ቇળᄢળࠪࡦࡐࠫ࠙ࡓ (2011/09/18, ᮮᵿ)㧚
੹੗໪㓶㧘㋈ᧁධ⟤㧘᧻੗ᷕ㧘ㇹᐽᐢ㧘⍹ਣ༑ᦶ㧘ਃဈᏁ㧘㒙ㇱ໪ሶ㧘ᐔ੗໪ਭ (2011) ⧰๧ฃኈ૕ TAS2R16
ᗵฃᕈߩ⒳㑆Ꮕߣಽሶᯏ᭴. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/17, ›ጊ)㧚
ฎ⾐┨ᒾ㧘ᐔ੗⊖วሶ㧘ᐔ੗໪ਭ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ޽ߞߡࡅ࠻ߦߥ޿ᨴ⦡૕┵ㇱࠥࡁࡓਇᲫ࿾Ꮺ:(2) ၭ
㙃⚦⢩ࠍ↪޿ߚ⚵឵߃ߩᬌ಴. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/16-18, ›ጊᏒ)㧚
᧻੗ᷕ㧘Jahn I㧘Islam MA㧘Rahman ZMM㧘ᐔ੗໪ਭ (2011) ࡒ࠻ࠦࡦ࠼࡝ࠕࠥࡁࡓߦࠃࠆ࠹࠽ࠟࠩ࡞ߩಽሶ
♽⛔ㅴൻ. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/16-18, ›ጊᏒ)㧚
㋈ᧁධ⟤㧘᧻੗ᷕ㧘ㇹᐽᐢ㧘⍹ਣ༑ᦶ㧘ਃဈᏁ㧘㒙ㇱ໪ሶ㧘ᐔ੗໪ਭ㧘੹੗໪㓶 (2011) ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ
࿾ၞ․⇣⊛ߥ⧰๧ᗵฃᕈᄌ⇣. ᣣᧄㅴൻቇળ╙ 13 ࿁ᄢળ (2011/07/30, ੩ㇺ)㧚
㋈ᧁධ⟤㧘᧻੗ᷕ㧘ㇹᐽᐢ㧘⍹ਣ༑ᦶ㧘ਃဈᏁ㧘㒙ㇱ໪ሶ㧘ᐔ੗໪ਭ㧘੹੗໪㓶 (2011) ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ
⧰๧ฃኈ૕ TAS2R38 ߩ࿾ၞ․⇣⊛ߥᗵฃᕈᄌ⇣. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/17, ›ጊ)㧚
੹੗໪㓶㧘㋈ᧁධ⟤㧘ᣧᎹථᔒ㧘⩲ේ੧㧘᧻੗ᷕ㧘ㇹᐽᐢ㧘ᰞ੗ᢘዷ㧘⍹ਣ༑ᦶ㧘Lijie Yin㧘Wenshi Pan, 㒙
ㇱ໪ሶ㧘ਃဈᏁ㧘ᐔ੗໪ਭ (2012) 㔤㐳㘃๧ⷡฃኈ૕ߩㅴൻ. ᣣᧄ↢ℂቇળᄢળ╙ 89 ࿁ᄢળࠪࡦࡐࠫ࠙ࡓ
(2012/03/31, ᧻ᧄ)㧚
੹੗໪㓶 (2011/10/12) ‫ޟ‬㔤㐳㘃ࡃࠗࠝ࡝࠰࡯ࠬߩ⃻⁁ߣዷᦸ‫ޠ‬. ᾢᧄᄢቇ╙ 16 ࿁ㆮવሶታ㛎ᣉ⸳࠮ࡒ࠽࡯‫ࡃޟ‬
ࠗࠝ࡝࠰࡯ࠬᦨ೨✢‫ޠ‬. ᾢᧄ㧚
੹੗໪㓶 (2011/11/19) ‫ࡓࡁࠥޟ‬ᄙဳߩᯏ⢻⸃ᨆ㧦㔤㐳㘃‫ޠ‬. ╙ੑ࿁⣖⴫⃻ဳߩಽሶࡔࠞ࠾࠭ࡓ⎇ⓥળ 㧔᜗ᓙ
⻠Ṷ㧕㧚
ᐔ੗໪ਭ (2012/03/19) Chimpanzee chromosomes and gibbon oversea project. Linconl University, UK㧚
ነ㒝⎇ⓥㇱ㐷
Ყセ⹺⍮⊒㆐㧔ࡌࡀ࠶࠮ࠦ࡯ࡐ࡟࡯࡚ࠪࡦ㧕⎇ⓥㇱ㐷
㧨⎇ⓥ᭎ⷐ㧪
A) ࠴ࡦࡄࡦࠫ࡯ߩ⍮ⷡ⹺⍮⢻ജߩᲧセ⹺⍮⑼ቇ⊛⎇ⓥ
દ᧛⍮ሶ
࠴ࡦࡄࡦࠫ࡯ߣࡅ࠻ࠍኻ⽎ߦ‫‛ޔ‬૕ߩ⾰ᗵ⍮ⷡߦ㑐ߔࠆ⢻ജ‫ޔ‬ᒻᘒᖱႎߣㆇേᖱႎߩ⛔ว⢻ജࠍ⋥ធᲧセߔࠆ⎇
ⓥࠍ߅ߎߥߞߚ‫ޕ‬
B) ࡅ࠻ߣ࠾ࡎࡦࠩ࡞੃ఽߩ⍮ⷡ⊒㆐ߩᲧセ
દ᧛⍮ሶ
ࡅ࠻ߩ੃ఽߣ࠾ࡎࡦࠩ࡞੃ఽࠍኻ⽎ߦ‫ޔ‬ਇน⢻࿑ᒻߥߤࠍ↪޿ߚᅏⴕ߈⍮ⷡߩ⢻ജߩ⊒㆐߿‫ޔ‬ᒻᘒᖱႎߣㆇേᖱ
ႎߩ⛔ว⢻ജߩ⊒㆐ߦߟ޿ߡ‫ޔ‬ᵈⷞᤨ㑆ࠍᜰᮡߦ⺞ߴߚ‫ޕ‬
㧙㧙
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
Goto K, Imura T, Tomonaga M (2012) Perception of emergent configurations in humans (Homo sapiens) and
chimpanzees (Pan troglodytes). Journal of Experimental Psychology: Animal Behavior Processes, 38:125-138. doi:
10.1037/a0026899.
ቇળ⊒⴫
1)
2)
3)
Imura T (2011) Visual temporal integration on object recognition in chimpanzees and humans. 34th European
Conference on Visual Perception (2011/09, Toulouse, France).
દ᧛⍮ሶ (2011) ࠴ࡦࡄࡦࠫ࡯ߣࡅ࠻ߦ߅ߌࠆࠬ࡝࠶࠻ⷞ᧦ઙਅߩ‛૕⹺⼂. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળ
(2011/07, ᗲ⍮)㧚
દ᧛⍮ሶ㧘⊕੗ㅀ (2011) ࠬ࡝࠶࠻ⷞ᧦ઙߦ߅ߌࠆᒻᘒߣㆇേߩ⛔ว⢻ജߩೋᦼ⊒㆐. ᣣᧄၮ␆ᔃℂቇળ╙ 30
࿁ᄢળ (2011/12, ᧲੩)㧚
㒝ዻᣉ⸳
ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯
ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯ߪᚲౝߩฦ⒳⎇ⓥߩᡰេ߿࠽࡚ࠪ࠽࡞ࡃࠗࠝ࡝࠰࡯ࠬࡊࡠࠫࠚࠢ࠻㧔NBRP㧕߳ߩ
ࠨ࡞ߩଏ⛎ߩߚ߼ߦ‫ޔ‬ᣉ⸳ᢛ஻‫ޔ‬ฦ⒳Უ⟲ߩ⛽ᜬ‫ޔ‬㘺⢒࡮❥ᱺ‫ޔ‬ஜᐽ▤ℂࠍ߅ߎߥ߁ߣߣ߽ߦ‫ߟߦ࡞ࠨߩࠄࠇߎޔ‬
޿ߡߩ⒳‫⎇ߩޘ‬ⓥࠍផㅴߒߡ޿ࠆ‫ޕ‬
2011 ᐕᐲߪ‫ޔ‬WISH ੐ᬺߩ৻Ⅳߣߒߡ‫࡯ࠫࡦࡄࡦ࠴ޔ‬㘺⢒ࠛ࡝ࠕౝߩ᧲⷏ߩࠨࡦ࡞࡯ࡓߩᡷୃࠍⴕ߁ߣߣ߽ߦ╙
3 ᡼㘺႐ߦ࠴ࡦࡄࡦࠫ࡯㘺⢒ࠤ࡯ࠫࠍᣂ⸳ߒߚ‫᧲ޔߜ߁ߩߘޕ‬஥ߩࠨࡦ࡞࡯ࡓߦߟ޿ߡߪ‫ޔ‬ᐕᐲᧃࠃࠅ౒↪ࠍ㐿ᆎ
ߒߚ‫ޔߚ߹ޕ‬㘺⢒ࠛ࡝ࠕߩ᜛ᄢߦ઻ߞߡ‫ޔ‬㕖Ᏹൕ⡯ຬ 1 ฬࠍᣂⷙߢ㈩⟎ߒߚ‫ޕ‬
ㄭᐕ‫ޔ‬㔤㐳㘃⎇ⓥᚲߩ࠾ࡎࡦࠩ࡞ߦ߅޿ߡ‫ోޔ‬り߆ࠄߩ಴ⴊࠍ઻ߞߡᱫ੢ߔࠆ∛᳇㧔࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝㧕
߇⊒↢ߒߡ޿ߚ߇‫ޔ‬2010 ᐕᐲߦᄖㇱᯏ㑐ߩදജߩ߽ߣ♖ജ⊛ߦ⎇ⓥࠍㅴ߼ߚ⚿ᨐ‫ߩ∝ᧄޔ‬ේ࿃ߪ‫࠙ࡠ࠻࡟࡞ࠨޔ‬
ࠗ࡞ࠬ 4 ဳ㧔SRV-4㧕ߢ޽ࠆߎߣ߇᣿ࠄ߆ߣߥߞߚ‫ޕ‬2011 ᐕᐲߪ‫ޔ‬೨ᐕᐲᧃߦ⸳⟎ߒߚᬌᩏ▤ℂㇱ߇ਛᔃߣߥߞߡ
ᚲౝߩࡑࠞࠢዻࠨ࡞ో㗡ߩ SRV ᬌᩏࠍታᣉߒ‫ޔ‬ᗵᨴߩ⇼޿ߩ޽ࠆ୘૕ߪ‫ߡߴߔޔ‬㓒㔌᫟‫ޔ‬ᬌ∉᫟ߦ㓒㔌ߒߚ‫ߘޕ‬
ߩ⚿ᨐ‫ޔ‬2011 ᐕᐲએ㒠‫ޔ‬ᣂⷙߩ⊒∝ߪ 1 ଀߽⷗ࠄࠇߡ޿ߥ޿‫ޔߚ߹ޕ‬8 ᦬ࠃࠅ NBRP ⚻⾌ߦߡ․ቯഥᢎࠍ㓹↪ߒ‫ޔ‬
ห∝ߩ⊒∝ᯏᐨ⸃᣿ߦขࠅ⚵߻ߣߣ߽ߦ‫ޔ‬ᬌᩏㇱ߇ਛᔃߣߥߞߡ‫ޔ‬SRV ߩᬌᩏᴺߩᡷ⦟ࠍⴕߞߚ‫ޕ‬
NBRP ߩ࠾ࡎࡦࠩ࡞ߦ㑐ߒߡߪ‫ⴊޟޔ‬ዊ᧼ᷫዋ∝‫޿઻ߦ↢⊒ߩޠ‬Უ⟲ߪ 228 㗡ߦ߹ߢᷫዋߒߚ߇‫ޔ‬⢒ᚑ⟲ߩ❥ᱺ
ߪ㗅⺞ߦㅴࠎߢ߅ࠅ‫⃻ޔ‬࿷ว⸘ 350 㗡ߩ NBR ࡊࡠࠫࠚࠢ࠻↪࠾ࡎࡦࠩ࡞ࠍ㘺㙃ߒߡ޿ࠆ‫ޕ‬
ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯ߩᛛⴚ⡯ຬߩዬቶߪ࠮ࡦ࠲࡯᫟ߦ⸳⟎ߐࠇߡ޿ߚ߇‫ᧄޔ‬㙚 1 㓏ߩᣥ․೎ળ⼏ቶߦ⒖
⸳ߐࠇߚ‫ޔࠅࠃߦࠇߎޕ‬ᢎຬ‫ޔ‬ᛛⴚ⡯ຬ‫࠻࡯ࡄޔ‬⡯ຬ‫⎇ޔ‬ⓥຬߩዬቶߪోߡᧄ᫟ౝߦ㈩⟎ߐࠇࠆߎߣߣߥߞߚ‫ޕ‬
ੱ੐㕙ߢߪ 2011 ᐕ 8 ᦬ 1 ᣣߦ․ቯഥᢎߩ૒⮮⧷ᰴ߇ዞછߒߚ‫․ޕ‬ቯ⎇ⓥຬߣߒߡ㥱⮮ᥙ‫᧲ޔ‬Ớ◊ᓼࠍ 6 ᦬߆ࠄ
ណ↪‫ޕ‬㕖Ᏹൕ⡯ຬߦߪએਅߩ⇣േ߇޽ߞߚ‫ޕ‬2011 ᐕ 4 ᦬ࠃࠅᢎോ⵬૒ຬߦ↢㚤ᥓሶ‫ᧄ᧖ޔ‬ᄥ㇢ࠍណ↪‫࡞ࠨޕ‬㘺⢒
ᜂᒰߣߒߡ 5 ᦬Ꮉᷝᥓ㚅‫⩆ޔ‬㊁ᄹ⟤‫ޔ‬6 ᦬೨↰Ꮣ⟤ሶ‫ޔ‬㐳⼱Ꮉᄕ⟤ሶ‫ޔ‬7 ᦬ᳯญ⡛ሶ‫ޔ‬2012 ᐕ 2 ᦬㜞ᧁ᦮ሶ‫ޔ‬ᱞ⮮
ਭ⟤‫ޔ‬ဈ੗ዏ⟤‫ޔ‬3 ᦬ᄐ⋡ዅᅢ‫⮮ޔ‬᫪໑ࠍᛛ⢻⵬૒ຬߦណ↪‫ޕ‬4 ᦬ታ㛎⵬ഥᜂᒰߣߒߡ቟ᳯ⟤㔐㧔ᛛⴚ⵬૒ຬ㧕ࠍ
ណ↪‫ޕ‬4 ᦬ᄢၳ⟤૫㧔⎇ⓥᡰេផㅴຬ㧕⎇ⓥഥᚑ߆ࠄ㈩⟎឵‫ޕ‬5 ᦬ศ↰⟤ජሶ‫ޔ‬6 ᦬ᳯญ⡛ሶ‫ޔ‬2012 ᐕ 3 ᦬દ⮮๺
ሶ‫ޔ‬⍹↰ᕺᵤሶ‫ޔ‬ᄢ┻౏ሶ㧔ᛛ⢻⵬૒ຬ㧕‫ޔ‬ฎᯅ଻ᔒ㧔⎇ⓥᡰេផㅴຬ㧕߇ㅌ⡯ߒߚ‫ޕ‬
㧨⎇ⓥ᭎ⷐ㧪
A) ࠥࡁࡓਇᲫ࿾Ꮺ㧔RCRO㧕ߩㅴൻߣᗧ⟵
ᐔ੗໪ਭ㧘ฎ⾐┨ᒾ㧔ࠥࡁࡓᄙ᭽ᕈಽ㊁㧕㧘ᐔ੗⊖วሶ㧔ᛛ⢻⵬㧕㧘㡻Ლବผ㧔ᾢᧄࠨࡦࠢ࠴ࡘࠕ࡝㧕㧘᧻ᨋᷡ᣿㧔ฬ
⹷ᢎ᝼㧕
RCRO ߩ᭴ᚑⷐ⚛ߩ߭ߣߟߢ޽ࠆࠨࡉ࠲࡯ࡒ࠽࡞ࠨ࠹࡜ࠗ࠻㧔StSat㧕ࠍࠢࡠ࡯ࡦൻߒ‫ ߥ⚦⹦ޔ‬FISH ⸃ᨆࠍ߅ߎ
ߥߞߚ‫࡯ࠫࡦࡄࡦ࠴ޕ‬43 ୘૕ߩᨴ⦡૕ࠍ⸃ᨆߒ‫ޔ‬ᄌ⇣ߩ⁁ᴫࠍ᣿ࠄ߆ߦߒߚ‫ޕ‬ᷫᢙಽⵚ♖Უ⚦⢩ߩᨴ⦡૕ࡉ࡯ࠤ
ߣߩ㑐ㅪ߆ࠄ RCRO ߩሽ࿷ᗧ⟵ࠍ⼏⺰ߒ‫⺰ޔ‬ᢥߣߒߡ߹ߣ߼ߚ‫ޕ‬
B) ࠹࠽ࠟࠩ࡞㘃ߩᄙ᭽ᕈߣ↢‛࿾ℂቇ
ᐔ੗໪ਭ㧘ࠬ࠳࡜ࠬ࡮ࡃࠗ࠴ࡖࡠࠛࡦ㧔࠲ࠗേ‛࿦දળ㧘ࠞ࠮ࠨ࡯࠻ᄢቇ㧕㧘ࠗࠬ࡜࠻࡮ࠫࡖࡂࡦ㧔ᄢቇ㒮↢㧕㧘ฎ
⾐┨ᒾ㧔ࠥࡁࡓᄙ᭽ᕈ㧕㧘ᐔ੗⊖วሶ㧔ᛛ⢻⵬㧕㧘᧻੗ᷕ㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯⎇ⓥຬ㧕
ࡒ࠻ࠦࡦ࠼࡝ࠕࠥࡁࡓోႮၮߩ⸃ᨆ߆ࠄ‫ޔ‬Hoolock hoolock ࠍട߃ߡ࠹࠽ࠟࠩ࡞ 4 ዻߩಽሶ♽⛔ಽጘࠍ᣿ࠄ߆ߦ
ߒߚ‫ߩࡦࡑࠕࠪޕ‬ᨴ⦡૕ᧃ┵ߦሽ࿷ߔࠆࡋ࠹ࡠࠢࡠࡑ࠴ࡦߩ DNA ࠍ⸃ᨆߒ‫ޔ‬4 ዻߦ߅޿ߡߘࠇߙࠇ․⇣⊛ߥᨴ⦡
૕਄ಽᏓࡄ࠲ࡦࠍ๒ߔࠆߎߣࠍ᣿ࠄ߆ߦߒߚ‫ࠕࡔࡠ࠹ߣࠕࡔࡠ࠻ࡦ࠮ޕ‬๟ㄝߩࡋ࠹ࡠࠢࡠࡑ࠴ࡦߩᒻᚑߦ㑐ࠊࠆ
DNA ߩ㑐ㅪᕈߦߟ޿ߡ⸃ᨆߒߚ‫ޕ‬
C) ࡑࡦ࠰ࡦ૑ⴊๆ⯻ߩᕈᨴ⦡૕㧔Z‫ޔ‬W㧕ߩㅴൻ
ᐔ੗໪ਭ㧘ᐔ੗⊖วሶ㧔ᛛ⢻⵬㧕㧘ࡈࠖ࡝࠶ࡊ࡮ࡠࡌ࡞࠺㧔࠹ࠠࠨࠬᄢቇ㧕
㧙㧙
BAC mapping ߦࠃߞߡ᣿ࠄ߆ߦߥߞߚࠢࡠ࡯ࡦߩሽ࿷᭽ᑼ߆ࠄ‫ޔ‬Z ߅ࠃ߮ W ᕈᨴ⦡૕ߩㅴൻ߇ 4 ࿁ߩㅒ૏ߦࠃ
ߞߡ↢ߓߚߎߣࠍផቯߒߚ‫ోޔߚ߹ޕ‬ᨴ⦡૕ߩᓀ⦡ࡊࡠ࡯ࡉࠍ૞ᚑߒ‫ޔ‬Z ߣ W ᨴ⦡૕ߩ⋧ห߅ࠃ߮㕖⋧หㇱ૏ࠍ
᣿ࠄ߆ߦߒߚ‫⺰ޕ‬ᢥߣߒߡ߹ߣ߼ߚ‫ޕ‬
D) ࠹࠾ࠕ⑼᧦⯻ᐜ⯻ᗵᨴኅ⇓୘૕ߩ⼂೎ߦ᦭↪ߥᣂߒ޿ᛛⴚ㐿⊒ߣ࡝ࠬࠢ⹏ଔ߳ߩᔕ↪
ጟᧄቬ⵨
ࡅ࠻ࠍ⚳ኋਥߣߔࠆ࠹࠾ࠕ⑼᧦⯻ߦߪ‫᦭ޔ‬㋭᧦⯻‫ޔ‬ή㋭᧦⯻‫ ߩࠕ࠾࠹ࡦࡢࠗ࠲ޔ‬3 ⒳߇⍮ࠄࠇߡ޿ࠆ‫ޕ‬ᐔᚑ 23
ᐕᐲߪ‫࡝ࡃ࡮ࠕࠪࡀ࠼ࡦࠗޔ‬ፉߩᵹⴕ࿾ߢ⺞ᩏࠍታᣉߒߚߣߎࠈ‫࡝ࡃޔ‬ፉߩർㇱߩዊ᧛ߪ᦭㋭᧦⯻ߩỚෘᳪᨴ࿾ၞ
ߢ޽ࠆߎߣ߇⏕⹺ߢ߈ߚ‫⃻ޕ‬࿾ߢታᣉߒߚ ELISA ߦࠃࠅ‫᦭ޔ‬㋭ྙ⯻ߦኻߔࠆ᛫૕ࠍ଻᦭ߒߡ޿ࠆࡅ࠻߅ࠃ߮ࡉ࠲
ࠍ⏕⹺ߒߚ‫ࠍ࠲ࡉߩߎޕ‬೬ᬌߒߚߣߎࠈ‫ޔ‬ᄙᢙߩ᦭㋭ྙ⯻ߩነ↢߇⏕⹺ߢ߈ߚߎߣ߆ࠄ‫࡝ࡃޔ‬ፉߦ߅޿ߡ᦭㋭᧦⯻
ߩ↢ᵴⅣ߇ᚑ┙ߒߡ޿ࠆߎߣ߇᣿ࠄ߆ߣߥߞߚ‫ޔߚ߹ޕ‬ᚒ‫ߩޘ‬㐿⊒ߒߚ ELISA ߪ․೎ߥᯏེߩߥ޿ᵹⴕ࿾ߢ߽ච
ಽ᦭ലߢ޽ࠆߎߣ߇᣿ࠄ߆ߣߥߞߚ‫ޕ‬
E) 㔍ᴦᕈነ↢⯻∛ߦ㑐ߔࠆㆮવሶ⸻ᢿᴺߩ㐿⊒
ጟᧄቬ⵨
ㄭᐕᚒ‫⎇ߩޘ‬ⓥࠣ࡞࡯ࡊߪ‫ޔ‬ή㋭᧦⯻ߣࠕࠫࠕ᧦⯻੤㔀૕߇ሽ࿷ߔࠆߎߣࠍ⊒⷗ߒߚ‫ߩࠄࠇߎޔ‬੤㔀૕ߪᓥ᧪ߩ
ㆮવሶ⸻ᢿᴺߢߪ⸻ᢿߢ߈ߥ޿ߚ߼‫ߩࠄࠇߘޔ‬ㆮવሶࠍ⸃ᨆߒ‫ޔ‬ᣂߒ޿⸻ᢿᴺࠍ㐿⊒ߔࠆᔅⷐ߇޽ࠆ‫ޕ‬ᐔᚑ 23 ᐕ
ᐲߪ‫ߩࠄࠇߎޔ‬࿾ၞߩ߁ߜ࠲ࠗߣਛ࿖྾Ꮉ⋭ߢ∉ቇ⺞ᩏࠍታᣉߒ‫ߦ࠻ࡅޔ‬ነ↢ߒߡ޿ࠆᚑ⯻߅ࠃ߮ࡅ࠻߳ߩᗵᨴḮ
ߣߥࠆኅ⇓ߦነ↢ߒߡ޿ࠆᐜ⯻ࠍណขߒߚ‫⯻ߩࠄࠇߎޕ‬૕ߦߟ޿ߡ‫ߣࠕ࡝࠼ࡦࠦ࠻ࡒޔ‬ᩭߩㆮવሶࠍߢ߈ࠆߛߌ✂
⟜⊛ߦ⺞ߴࠆߣߣ߽ߦ‫ޔ‬ᣂߚߦ㐿⊒ߒߚ LAMP ᴺ߇ࡈࠖ࡯࡞࠼ߢߩ⸻ᢿߦ೑↪ߢ߈ࠆߎߣࠍ⏕⹺ߒߚ‫ޔߦࠄߐޕ‬
ᖚ⠪ߩዬ૑࿾ߣᗵᨴኅ⇓ߩ㘺⢒႐ᚲߩ㑐ଥࠍ‫ޔ‬GIS㧔࿾ℂᖱႎࠪࠬ࠹ࡓ㧕ࠍ೑↪ߒߡ⸃ᨆߒߚ‫ޕ‬
F) ᄙ൮⯻∝ߦ㑐ߔࠆ㕖㐿⣻⊛ᴦ≮ᴺ߳ߩ᜸ᚢߣ⹏ଔᴺߩ㐿⊒
ጟᧄቬ⵨
࡜࠶࠻ߩ⢄⤳ߩᚲቯߩ૏⟎ߦᄙ൮⯻∝∛Ꮍࠍ૞⵾ߔࠆᣇᴺߦߟ޿ߡᬌ⸛ߒߚ‫ోޕ‬り㤗㈮ਅߢ‫ޔ‬⣻ㇱᱜਛࠍ㐿⣻ᓟ‫ޔ‬
⢄⤳ߩ৻ㇱࠍ૕ᄖߦ㔺಴ߒߚ‫ߢࠬࡔޕ‬⢄⤳ߦ⚂ 5mm ߩಾ㐿ഃࠍ૞⵾ߒ‫߼ߓ߆ࠄ޽ޔ‬Ḱ஻ߒߡ߅޿ߚᄙ൮⯻ߩࠪࠬ
࠻ࠍಾ㐿ഃߦ⒖ᬀߒߚ‫ޕ‬ᱛⴊߣࠪࠬ࠻ߩ⣻⣧ౝᵹ಴㒖ᱛߩߚ߼ߦ‫ޔ‬ᱛⴊࠪ࡯࠻ߢಾ㐿ഃࠍⵍⷒߒߚ‫ߦ଀ోޕ‬ᣈ޿ߡ‫ޔ‬
⒖ᬀㇱ૏ߢߩࠪࠬ࠻ߩ↢㐳ࠍ MRI ߢ⚻ᤨ⊛ߦㅊ〔ߢ߈ߚ‫ᧄޕ‬ᴺߪᚲቯߩ૏⟎ߦᄙ൮⯻∝∛Ꮍࠍ૞⵾ߔࠆ਄ߢᭂ߼
ߡ᦭↪ߥᣇᴺߢ޽ࠆߎߣࠍ⏕⹺ߒߚ‫∛ޔߡߞ૶ࠍ࠻࠶࡜࡞࠺ࡕߩߎޔߢߎߘޕ‬ᐥ㒰෰ᓟߩ Em18 ᛫ේߦኻߔࠆ᛫૕
ଔߩផ⒖ࠍ⺞ߴߚ‫ޕ‬
G) ࠞ࠾ࠢࠗࠩ࡞/human immunodeficiency virus type-1 (HIV-1) ᗵᨴࡕ࠺࡞ߩ㐿⊒ߦ㑐ߔࠆ⎇ⓥ
㥱⮮ᥙ㧘᣿㉿ብᢥ
ᧄ⎇ⓥߢߪㄭᐕ⏕┙ߐࠇߚ‫⚦࡞ࠨޔ‬⢩ߢჇᱺน⢻ߥࠨ࡞ᜰะᕈ HIV-1 ࠢࡠ࡯ࡦ㧔macaque-tropic HIV-1: HIV-1mt㧕
ࠍ↪޿ߡߎࠇ߹ߢਇน⢻ߣߐࠇߡ߈ߚࡕ࠺࡞േ‛ߢ޽ࠆታ㛎↪ࠨ࡞㘃㧛HIV-1 ᗵᨴ࡮⊒∝ࠪࠬ࠹ࡓࠍ⏕┙ߔࠆߎߣ
߇⋡⊛ߢ޽ࠆ‫ޕ‬㜞㚔ൻဳ╙ 3 ਎ઍ HIV-1mt ߢ޽ࠆ MN4Rh-3 ߪ‫ ࡞ࠩࠗࠢ࠾ࠞޔ‬PBMC ߅ࠃ߮୘૕ߣ߽ߦ╙ 2 ਎ઍ
HIV-1 ߣᲧセߒߡ࠙ࠗ࡞ࠬჇᱺല₸߇ 10 ୚ะ਄ߒߚ㧔105 viral RNA copies/ml㧕
‫ࠬ࡞ࠗ࠙ߚ߹ޕ‬Ⴧᱺߦ઻޿‫৻ޔ‬ㆊᕈ
ߢߪ޽ࠆ߇㗼⪺ߥ CD4 㓁ᕈ T ⚦⢩ߩᷫዋ߇⏕⹺ߐࠇߚ‫ࠬ࡞ࠗ࠙ߪ∝ⴊࠬ࡞ࠗ࠙ޕ‬ធ⒳ᓟ 6 ㅳߢᬌ಴㒢⇇એਅߣߥ
ߞߚ߇‫ࠬ࡞ࠗ࠙ޔ‬ធ⒳ᓟ 17 ㅳߩᤨὐߢ CD8 ․⇣᛫૕ࠍᛩਈߒ CD8 㓁ᕈ T ⚦⢩㒰෰ࠍⴕߥߞߚߣߎࠈ HIV-1mt ౣ
ᵴᕈൻ߇↢ߓ࠙ࠗ࡞ࠬ RNA ߇ᬌ಴ߐࠇߚ‫ޔࠄ߆ߣߎߩߎޕ‬MN4Rh-3 ߪࠞ࠾ࠢࠗࠩ࡞ߦ߅޿ߡ㐳ᦼẜફᗵᨴ߇น⢻
ߢ޽ࠆߎߣ߇␜ߐࠇߚ‫ޕ‬
H) ࠞ࠾ࠢࠗࠩ࡞ߦ߅ߌࠆ macaque-tropic human immunodeficiency virus type-1 (HIV-1mt) Ⴧᱺߩ୘૕Ꮕࠍⷙቯߔ
ࠆኋਥౝ࿃ᕈ࿃ሶ TRIM5 ߦ㑐ߔࠆಽሶㆮવቇ⎇ⓥ
㥱⮮ᥙ㧘᣿㉿ብᢥ
ᚒ‫ޔߦߢ߹ࠇߎߪޘ‬HIV-1mt Ⴧᱺേᘒ߇ធ⒳ࠞ࠾ࠢࠗࠩ࡞୘૕ߦࠃߞߡ㗼⪺ߦ⇣ߥࠆߎߣ‫ޔ‬TRIM5 ㆮવሶࠕ࡝࡞
߇ߘߩਥߥⷙቯⷐ࿃ߢ޽ࠆߎߣࠍ␜ໂߔࠆ⍮⷗ࠍᓧߚ‫ޔߢߎߘޕ‬TRIM5 ߩ HIV-1mt ߳ߩᯏ⢻⊛ᗧ⟵ࠍᬌ⸛ߒߚߣ
ߎࠈ‫ޔ‬TRIMCyp㧔TRIM5 ߩ৻ㇱ㗔ၞߦ cyclophilin A ߩ৻ㇱ߇ᝌ౉ߐࠇߚᄌ⇣ဳࠕ࡝࡞㧕ࠍ᦭ߔࠆࠞ࠾ࠢࠗࠩ࡞୘
૕ߢߪ㊁↢ဳߢ޽ࠆ TRIM5 ࠕ࡟࡞଻᦭୘૕ߣᲧセߒߡ‫ޔ‬HIV-1mt ᗵᨴߦ߅ߌࠆⴊਛ࠙ࠗ࡞ࠬ㊂߇⚂ 50 ୚㜞޿ߎ
ߣࠍ⷗಴ߒߚ‫⥝ޕ‬๧ᷓ޿ߎߣߦ‫ޔ‬TRIM5/TRIMCyp 㗫ᐲߪࠞ࠾ࠢࠗࠩ࡞ߩේ↥࿾ߦࠃߞߡᄢ߈ߊ⇣ߥߞߡ߅ࠅ‫ࡈޔ‬
ࠖ࡝ࡇࡦ↱᧪୘૕⟲ߦ߅ߌࠆ TRIMCyp ࠕ࡝࡞㗫ᐲ㧔87.0%㧕ߪࠗࡦ࠼ࡀࠪࠕ෸߮ࡑ࡟࡯ࠪࠕ↱᧪୘૕⟲ߩࠕ࡝࡞
㗫ᐲ㧔34.8%‫ޔ‬48.9%㧕ߣᲧセߒߡ᦭ᗧߦ㜞߆ߞߚ‫ޕ‬HIV-1 ᗵฃᕈߦ㑐ߔࠆࠨ࡞୘૕Ꮕࠍ TRIM5 ㆮવሶဳ߇ⷙቯߒ
ߡ޿ࠆߎߣࠍ in vitro ߩߺߥࠄߕ in vivo ߢ߽ታ⸽ߒ‫ޔ‬HIV-1 ࠨ࡞ࡕ࠺࡞⏕┙ߦะߌߡࡈࠖ࡝ࡇࡦ↥ࠞ࠾ࠢࠗࠩ࡞୘
૕ߩ᦭↪ᕈ߇ㆮવሶ࡟ࡌ࡞ߢታ⸽ߐࠇߚߎߣߪ․╩ߔߴ߈ᚑᨐߢ޽ࠆ‫ ߩ࡞ࠩࠗࠢ࠾ࠞߦࠄߐޕ‬TRIMCyp ࠕ࡝࡞㗫
ᐲߦ߅ߌࠆ࿾ℂ⊛ᄙ᭽ᕈ߇ᒝߊ␜ໂߐࠇ‫੹ޔ‬ᓟߎߩᄙ᭽ᕈࠍ↢ߺ಴ߒߚᄖ⊛ⷐ࿃ࠍ᣿ࠄ߆ߦߒߡ޿߈ߚ޿‫ޕ‬
I) C ဳ⢄Ἳ࠙ࠗ࡞ࠬᴦ≮࡮੍㒐ࡢࠢ࠴ࡦߩ቟ోᕈ࡮᦭ലᕈ⹏ଔߦ㑐ߔࠆ⎇ⓥ
᧲Ớ◊ᓼ㧘᣿㉿ብᢥ
C ဳ⢄Ἳ࠙ࠗ࡞ࠬ㧔HCV㧕ߪ਎⇇⊛ߦ⬧ᑧߒߡ߅ࠅ‫ޔ‬1 ం 7 ජਁ㧔਎⇇ੱญߩ⚂ 3%㧕߇ࠠࡖ࡝ࠕߣ⷗ࠄࠇߡ޿
㧙㧙
ࠆ‫ޕ‬ᣣᧄߢߩᜬ⛯ᗵᨴ⠪ߪ 190 ਁੱ㨪230 ਁੱሽ࿷ߔࠆߣផቯߐࠇߡ޿ࠆ߇‫߇ߣߎ޿ߥ߇⁁∝ⷡ⥄ޔ‬ᄙߊ‫ޔ‬⢄⎬ᄌ
߿⢄≸߳⒖ⴕߔࠆᗵᨴ⠪߇ᄙߊሽ࿷ߔࠆߎߣ߇໧㗴ߣߥߞߡ޿ࠆߚ߼‫ޔ‬᛫࠙ࠗ࡞ࠬ೷߿ᴦ≮↪ࡢࠢ࠴ࡦ㐿⊒߇ᕆോ
ߢ޽ࠆ‫⎇ᧄޕ‬ⓥߢߪ‫ޔ‬⣁↰ࠄ߇⊒⷗ߒߚၭ㙃น⢻ߥ HCV ߢ޽ࠆ JFH-1 ᩣࠍᄢ㊂♖⵾‫ޔ‬ਇᵴൻߔࠆߎߣߢ C ဳ⢄Ἳ
ߩ੍㒐࡮ᴦ≮ࡢࠢ࠴ࡦߣߒߡߩታ↪ൻࠍ⋡ᜰߒߡ޿ࠆ‫⎇ߩߢ߹ࠇߎޕ‬ⓥߦࠃࠅࡑ࠙ࠬ߳ߩធ⒳ߦࠃࠅ఺∉ේᕈ߇␜
ໂߐࠇߚᣂⷙਇᵴൻ HCV ☸ሶࡢࠢ࠴ࡦߦߟ޿ߡ‫੹ޔ‬ᐕᐲߪࠕࠞࠥࠩ࡞ߦ߅ߌࠆหࡢࠢ࠴ࡦߩ቟ోᕈ࡮᦭ലᕈࠍ⹏
ଔߒߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬HCV ߦኻߔࠆ․⇣⊛᛫૕⺃ዉ⢻߅ࠃ߮ធ⒳୘૕ߦ߅ߌࠆ቟ోᕈ߇⏕⹺ߐࠇߚ‫⺃ߚ߹ޕ‬ዉߐࠇ
ߚ᛫૕ߪ HCV ߦኻߔࠆᗵᨴ㒖ኂᵴᕈ߇ᓧࠄࠇߚ‫ޕ‬એ਄ߩᚑᨐߪ‫ޔ‬ਇᵴൻ HCV ☸ሶߦࠃࠆ੍㒐࡮ᴦ≮ࡢࠢ࠴ࡦ㐿
⊒ߦะߌߡᣂߚߥㅴᱠߢ޽ࠆߣ⠨߃ࠄࠇߚ‫ޕ‬
J) 㔤㐳㘃఺∉ਇో࠙ࠗ࡞ࠬߦࠃࠆ఺∉ᛥ೙ᯏ᭴ߦ㑐ߔࠆಽሶ᭴ㅧ↢‛ቇ⊛⎇ⓥ
㘵ፉᴕᐘ㧘᣿㉿ብᢥ
㔤㐳㘃఺∉ਇో࠙ࠗ࡞ࠬߪኋਥ఺∉ᔕ╵ࠍ․⇣⊛ߦ೙ᓮߔࠆߎߣߢ‫↢ޔ‬૕ౝߦ߅޿ߡ㐳ᦼᜬ⛯ᗵᨴߔࠆߎߣ߇⍮
ࠄࠇߡ޿ࠆ‫ߩߎޕ‬ᯏᐨߣߒߡ‫ޔ‬ห࠙ࠗ࡞ࠬߩ⺞▵Ⱞ⊕ߩ৻ߟߢ޽ࠆ Nef Ⱞ⊕߇᛫ේឭ␜ಽሶߢ޽ࠆ MHC ࠢ࡜ࠬ I
㧔MHC-I㧕ߩ⊒⃻ࠍᛥ೙ߔࠆߎߣ߇ႎ๔ߐࠇߡ޿ߚ߇‫ߥ⚦⹦ߩߘޔ‬ಽሶᯏ᭴ߪߎࠇ߹ߢᧂ⸃᣿ߢ޽ߞߚ‫⎇ᧄޕ‬ⓥߢ
ߪ‫ޔ‬MHC-I ⊒⃻೙ᓮߦᔅ㗇ߢ޽ࠆ Nef Ⱞ⊕ߣ‫ޔ‬ャㅍⰮ⊕ߩ৻ߟߢ޽ࠆ AP-1 ⶄว૕ mu-1A ࠨࡉ࡙࠾࠶࠻ߦࠃࠆ․⇣
⊛⚿วߦ㑐ߔࠆಽሶ᭴ㅧቇ⊛ߥࡔࠞ࠾࠭ࡓࠍ᣿ࠄ߆ߦߔࠆߎߣߦᚑഞߒߚ‫ߩߎޕ‬ᚑᨐߪ‫ޔ‬Nef Ⱞ⊕ࠍᜬߟ㔤㐳㘃఺
∉ਇో࠙ࠗ࡞ࠬߩኋਥ఺∉೙ᓮᯏ᭴ࠍ⸃᣿ߒߚ㊀ⷐߥ⊒⷗ߢ޽ࠆ‫ޕ‬
K) ࠨ࡞㘃ߩࠬ࠻࡟ࠬቯ㊂߅ࠃ߮േ‛⑔␩ߩߚ߼ߩၮ␆⎇ⓥ
㋈ᧁ᮸ℂ
㘺⢒ⅣႺߢߩࠬ࠻࡟ࠬ෻ᔕࠍቯ㊂ߔࠆߎߣߣߘߩシᷫ╷ߩᬌ⸛ߩߚ߼ߦ‫♮ߩ࡯ࠫࡦࡄࡦ࠴߮ࠃ߅ࠢࠞࡑޔ‬ਛࠦ࡯
࠴࠱࡞᷹ቯࠍⴕߞߚ‫ޕ‬㐳ᦼࠬ࠻࡟ࠬቯ㊂ߦ᦭ലߥᲫ㜬ਛࠦ࡯࠴࠱࡞᷹ቯ♽ߩ⏕┙ࠍᒁ߈⛯߈ⴕߞߡ޿ࠆ‫ޕ‬
L) ࠾ࡎࡦࠩ࡞ߩ∩ߺߩ⴫ᖱߦ㑐ߔࠆ੍஻⊛⎇ⓥ
ችㇱ⾆ሶ㧘㊍ਛᘮᦶ㧘G Tzimiropoulos㧘H Zulch㧘D Mills (Universityof Lincoln, UK)㧘ᐔ੗໪ਭ
࠾ࡎࡦࠩ࡞ߩ∩ߺߩ⴫ᖱࠍតࠆߎߣࠍ⋡⊛ߦ‫⴫ޔ‬ᖱߦ㑐ߔࠆ੍஻⊛ߥ⎇ⓥࠍ߅ߎߥߞߚ‫ߣ޿ߥ߇ߺ∩ޕ‬ᕁࠊࠇࠆ
࠾ࡎࡦࠩ࡞ߩ㗻౮⌀ࠍ 2397 ᨎߣ‫ ࠍࡊ࠶࡝ࠢࠝ࠺ࡆޔ‬6 ߟ᠟ᓇߒߚ‫ߩࠄࠇߎޕ‬౮⌀߆ࠄ‫⸃ޔ‬ᨆߦㆡߒߚ߽ߩࠍㆬᛯ
ߒ‫⸃࠲࡯ࡘࡇࡦࠦޔ‬ᨆࡕ࠺࡞ࠍ૞ᚑߒߡ޿ࠆߣߎࠈߢ޽ࠆ‫੹ޕ‬ᓟ‫ޔ‬᭽‫⺞ߥޘ‬ᢛࠍ߅ߎߥ޿‫ޔ‬ᣇᴺࠍᵞ✵ߐߖߡ޿ߊ
੍ቯߢ޽ࠆ‫ޕ‬㧔㗡⣖ᓴⅣࡊࡠࠣ࡜ࡓߢታᣉ㧕
M) ࡅ࠻♽⛔ߦ߅ߌࠆࠪࠕ࡞㉄ฃኈ૕ Siglec-11‫ޔ‬Siglec-16 ߩㅴൻ
ᣧᎹᢅਯ㧘X Wang㧔ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇࠨࡦ࠺ࠖࠛࠧᩞ㧕㧘N Mitra㧔ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇࠨࡦ࠺ࠖࠛࠧᩞ㧕㧘N Varki
㧔ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇࠨࡦ࠺ࠖࠛࠧᩞ㧕㧘A Varki㧔ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇࠨࡦ࠺ࠖࠛࠧᩞ㧕㧘቟ᒻ㜞ᔒ㧔ℂൻቇ⎇ⓥᚲ㧕
ࠪࠕ࡞㉄ߪ‫⚦ޔ‬⢩⤑⴫㕙ߩ♧㎮ߩᧃ┵ߦ޽ࠆ㉄ᕈන♧ߢ޽ࠅ‫⚦ޔ‬⢩㑆⹺⼂ᯏ᭴߿ኋਥ—∛ේ૕⋧੕૞↪ߦ߅޿ߡ
࡝ࠟࡦ࠼ߣߒߡ௛߈‫ߢߤߥ∉఺ޔ‬㊀ⷐߥᓎഀࠍᨐߚߒߡ޿ࠆ‫ޕ‬Siglec-11 ߣ Siglec-16 ߪ‫⚦ߒ⼂⹺ࠍ㉄࡞ࠕࠪޔ‬⢩ౝ
ࠪࠣ࠽࡞વ㆐ࠍ߅ߎߥ߁ฃኈ૕ߢ޽ࠆ‫ޕ‬Siglec-11 ㆮવሶߪࡅ࠻♽⛔ߢ Siglec-16 ㆮવሶߦࠃࠆㆮવሶᄌ឵ࠍฃߌ‫ޔ‬
ࡅ࠻․⇣⊛ߦ⣖ߢߩ⊒⃻ࠍ₪ᓧߒ‫⼂⹺㉄࡞ࠕࠪޔ‬⢻ࠍᄌൻߐߖߡ޿ࠆ‫ ߩߎޕ‬Siglec-11 ߩࡅ࠻♽⛔ߢߩㅴൻߩߐࠄ
ߥࠆ⍮⷗ࠍᓧࠆߚ߼‫࠻ࡅޔ‬⣖ߢߩ⊒⃻⚦⢩ߩ․ቯ‫࠻ࡅޔ‬⣖ౝߢߩ࡝ࠟࡦ࠼ߩᬌ಴‫߮ࠃ߅ޔ‬ㆮવሶᄌ឵ߩ⹦⚦ߥ⸃ᨆ
ࠍ߅ߎߥߞߚ‫⚿ߩߘޕ‬ᨐ‫ ࠻ࡅޔ‬Siglec-11 ߪ⣖ߩࡒࠢࡠࠣ࡝ࠕߢ⊒⃻ߒߡ߅ࠅ‫ޔ‬⣖ౝߦ࡝ࠟࡦ࠼߇⏕⹺ߐࠇ‫ޔ‬⣖ౝ
఺∉ߦ㑐ࠊߞߡ޿ࠆߎߣ߇ࠊ߆ߞߚ‫ޔߚ߹ޕ‬Siglec-11 ㆮવሶߪ Siglec-16 ㆮવሶߩਇᵴᕈൻࠕ࡝࡯࡞ߦࠃࠅㆮવሶ
ᄌ឵ߐࠇߡ߅ࠅ‫ߩߘޔ‬ㆮવሶᄌ឵ߪ Siglec-11 ㆮવሶߩனㆮવሶൻࠍ࿁ㆱߔࠆⶄ㔀ߥ߽ߩߢ޽ࠆߎߣࠍ⷗޿ߛߒߚ‫ޕ‬
ߎߩߎߣߪ‫ޔ‬ㆮવሶᄌ឵ߦࠃࠆ Siglec-11 ߩ⊒⃻ߣᯏ⢻ߩᄌൻߦ‫ޔ‬ㅴൻ⊛ㆬᛯ߇௛޿ߡ޿ࠆߎߣࠍ␜ໂߒߡ޿ࠆ‫ޕ‬
ߐࠄߦㆮવሶᄌ឵ߩᐕઍផቯ߆ࠄ‫ ࠻ࡅޔ‬Siglec-11 ߩ⊒⃻ߣᯏ⢻ߩᄌൻߪ⚂ 100 ਁᐕ೨ߦ߅ߎߞߚߣ⠨߃ࠄࠇߚ‫ޕ‬
એ਄ߩࠃ߁ߦ‫ޔ‬Siglec-11 ߪࡎࡕዻ಴⃻ᓟߦ⣖ౝ఺∉ߦ㑐ࠊࠆࠃ߁ߦߥࠅ‫ߩ࠻ࡅ߇ࠇߘޔ‬ㅴൻߦ㊀ⷐߢ޽ߞߚߎߣࠍ
␜ߔ⚿ᨐࠍᓧߡ޿ࠆ‫ޕ‬
N) 㔤㐳㘃ߦ߅ߌࠆࠪࠕ࡞㉄ฃኈ૕ Siglec-11‫ޔ‬Siglec-16 ߩㅴൻ
ᣧᎹᢅਯ㧘቟ᒻ㜞ᔒ㧔ℂൻቇ⎇ⓥᚲ㧕
Siglec-11 ㆮવሶߪࡅ࠻♽⛔․⇣⊛ߦ Siglec-16 ㆮવሶߦࠃࠆㆮવሶᄌ឵ࠍฃߌ‫ߦ⊛⇣․࠻ࡅޔ‬⣖ߢߩ⊒⃻ࠍ₪ᓧ
ߒ‫⼂⹺㉄࡞ࠕࠪޔ‬⢻ࠍᄌൻߐߖߡ޿ࠆ‫ ߩߎޕ‬Siglec-11 ߩࡅ࠻․⇣⊛ߥᄌൻߩ․ᱶᕈࠍ⍮ࠆߚ߼‫࠻ࡅޔ‬એᄖߩ㔤㐳
㘃ߩ Siglec-11 ߣ Siglec-16 ߩ⸃ᨆࠍ߅ߎߥߞߡ޿ࠆ‫ޕ‬
O) 㔤㐳㘃ߦ߅ߌࠆࠪࠕ࡞㉄ฃኈ૕ Siglec-13 ߩㅴൻ
ᣧᎹᢅਯ㧘X Wang㧔ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇࠨࡦ࠺ࠖࠛࠧᩞ㧕㧘N Mitra㧔ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇࠨࡦ࠺ࠖࠛࠧᩞ㧕㧘A Varki
㧔ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇࠨࡦ࠺ࠖࠛࠧᩞ㧕
Siglec-13 ߪ‫⚦ߒ⼂⹺ࠍ㉄࡞ࠕࠪޔ‬⢩ౝࠪࠣ࠽࡞વ㆐ࠍ߅ߎߥ߁ฃኈ૕ߢ޽ࠅ‫ߩߘޔ‬ㆮવሶᐳߪࡅ࠻․⇣⊛ߦᰳᄬ
ߒߡ޿ࠆ‫ߩߎޕ‬ᰳᄬߩේ࿃ߣᰳᄬߩㅴൻ⊛ߥᗧ๧ࠍ⍮ࠆߚ߼‫ޔ‬㔤㐳㘃ߩ Siglec-13 ߩࠥࡁࡓ㈩೉‫㉄࡞ࠕࠪޔ⃻⊒ޔ‬
⹺⼂⢻ߩ⸃ᨆࠍ߅ߎߥߞߡ޿ࠆ‫ޕ‬
㧙㧙
P) 㔤㐳㘃ࡑ࡜࡝ࠕේ⯻ࠍߪߓ߼ߣߒߚ⃻↢ࡑ࡜࡝ࠕේ⯻ߩ⿠Ḯ
ᣧᎹᢅਯ㧘ᯌ⌀৻㇢㧔ᄢ㒋ᄢቇ㧕㧘ᒾဈஜఽ㧔ᄢ㒋ᄢቇ㧕㧘᦭ᧃિሶ㧔ᄢ㒋ᄢቇ㧕㧘᧻੗ᷕ㧘ၳ੗ବብ㧔ᄢ㒋ᄢቇ㧕㧘
↰ㆺ๺ⵜ㧔ᄢ㒋ᄢቇ㧕
ࡑ࡜࡝ࠕේ⯻ߪ‫ޔ‬㔤㐳㘃߿ߍߞᱤ㘃ߣ޿ߞߚູ੃㘃߿㠽㘃‫ޔ‬Ῥ⯻㘃ࠍኋਥߣߒߡᗵᨴߒ‫ࠍࠕ࡝࡜ࡑޔ‬ᒁ߈⿠ߎߔ‫ޕ‬
ࡒ࠻ࠦࡦ࠼࡝ࠕࠥࡁࡓߩ⸃ᨆ߆ࠄ‫ࠕ࡝࡜ࡑ↢⃻ޔ‬ේ⯻ߩ⿠Ḯߦ߅޿ߡ‫ޔ‬ኋਥߩᄙ᭽ൻߦߣ߽ߥߞߚኋਥォ឵ߦࠃࠆ
ᕆㅦߥᄙ᭽ൻࠍ⷗޿ߛߒ‫ࡦࡃࠣ࠶ࡆࠕ࡝࡜ࡑ“ޔ‬઒⺑”ࠍឭ໒ߒߡ޿ࠆ‫ࡦࡃࠣ࠶ࡆࠕ࡝࡜ࡑޕ‬઒⺑ߩᬌ⸽ߩߚ߼‫ޔ‬ᩭ
ࠥࡁࡓߦࠦ࡯࠼ߐࠇߚㆮવሶ㧔ࠛࡀ࡞ࠡ࡯ઍ⻢ߦ㑐ࠊࠆㆮવሶ⟲㧕ߩㅴൻ⸃ᨆࠍ߅ߎߥ޿‫ࠕ࡝࡜ࡑ↢⃻ޔ‬ේ⯻ߩ౒
ㅢ␲వߪẋᣂ਎ߩᤨᦼߦ಴⃻ߒߚߎߣࠍ⷗޿ߛߒߡ޿ࠆ‫౒ߩߎޕ‬ㅢ␲వߩ಴⃻ᐕઍߪ‫ࡦࡃࠣ࠶ࡆࠕ࡝࡜ࡑޔ‬઒⺑ࠍ
ᡰᜬߔࠆ‫ޕ‬
Q) ࠕࡈ࡝ࠞ㊁↢ᄢဳ㘃ੱ₎ߦ߅ߌࠆ IgA ᛫૕ࠬࠢ࡝࡯࠾ࡦࠣߦࠃࠆੱ₞౒ㅢᗵᨴ∝ߩታᘒ⺞ᩏ
ศ↰෹ᢎ㧘┻రඳᐘ㧘૒⮮⧷ᰴ㧘ဈᏎື਽㧘ችㇱ⾆ሶ㧘↢㚤ᥓሶ㧘ᷰㆺᦶ㊁㧘౗ሶ᣿ਭ㧘ᷰㆺ␽ᐔ㧘೨↰ౖᒾ㧘ᣧ
Ꮉᢅਯ㧘㋈ᧁ᮸ℂ㧘ጟᧄቬ⵨㧘᧻ᴛື㇢㧘ฎᏒ೰ผ㧘᣿㉿ብᢥ
ㄭᐕ‫⹺ޔ‬⍮⎇ⓥ޽ࠆ޿ߪᔃߩ⎇ⓥ╬ߐ߹ߑ߹ߥ⎇ⓥߦߡᵈ⋡ߐࠇࠆ㊁↢ᄢဳ㘃ੱ₎߇‫ߡߞߚࠊߦ᧪ᧂޔ‬ሽ⛯น⢻
ߥ⁁ᘒߢ↢ᕷߔࠆߎߣ߇㕖Ᏹߦ࿎㔍ߥ⁁ᘒߦ޽ࠆ‫ߩߘޕ‬ਥߥℂ↱ߩ৻ߟߪ‫ߚૃߦ࠻ࡅߦ․ޔ‬ㆮવሶ᭴ᚑࠍᜬߟ㘃ੱ
₎ߢߪ‫ߩઁޔ‬േ‛⒳ߣ⇣ߥࠅ‫ੱޟ‬₞౒ㅢᗵᨴ∝‫߇ࠢ࡯࡟ࡉ࠻࠙ࠕߩޠ‬࿾ၞ୘૕⟲ߩሽ⛯ߦᄢ߈ߥ⢿ᆭߣߥߞߡ޿ࠆ
߆ࠄߢ޽ࠆ‫ᦨޕ‬ㄭᚒ‫ੱ᧪↱࠻ࡅޔߪޘ‬₞౒ㅢᗵᨴ∝ߩታᘒᛠីࠍⴕߥ߁ߚ߼‫ޔ‬㊁↢㘃ੱ₎♮ଢਛ߆ࠄේ࿃∛ේ૕․
⇣⊛ IgA ᛫૕ࠍᬌ಴ߢ߈ࠆ♽ࠍ⏕┙ߒߚ‫੹ߢߎߘޕ‬ᐕᐲߪ‫ᧄޔ‬ᴺࠍ↪޿ߡ‫ޔ‬㊁↢ᄢဳ㘃ੱ₎ߦ߅ߌࠆੱ₞౒ㅢᗵᨴ
∝ߩේ࿃∛ේ૕ߩታᘒ⸃᣿ࠍⴕߥ߁ߚ߼‫ޔ‬㊁↢ࡏࡁࡏߦ߅޿ߡ๭ๆེᗵᨴ∝ࠍᒁ߈⿠ߎߔ∛ේ૕ߦኻߔࠆ᛫૕ߩࠬ
ࠢ࡝࡯࠾ࡦࠣࠍⴕߥߞߚ‫⚿ߩߘޕ‬ᨐ‫↢ޔ‬ᕷၞߩᐢ▸࿐ߦᷰࠅⶄᢙߩ୘૕⟲ߢ‫ ࠩࡦࠛ࡞ࡈࡦࠗޔ‬A‫ޔ‬B‫ ߣޔ‬respiratory
syncytial virus (RSV) A‫ޔ‬B‫ޔ‬Parainfluenza type1~3‫ޔ‬Mumps ޽ࠆ޿ߪ Rhinovirus ߦኻߔࠆ᛫૕ࠍ଻᦭ߒߡ޿ࠆߎߣ߇
᣿ࠄ߆ߦߥߞߚ‫ߩࠄࠇߎޕ‬ᖱႎߪ‫∛ޔ‬ේ૕ߦኻߔࠆ᛫૕ࠍ଻᦭ߒߡ޿ࠆ୘૕߇ᄙᢙ޿ࠆߎߣ‫ߚߒ߁ߎߡߒ߁ߘޔ‬ᗵ
ᨴᱧࠍᜬߟߣផ᷹ߐࠇࠆ୘૕߇ᔅߕߒ߽㊀◊ߥ∔∛ࠍ๒ߔࠆߣߪ㒢ࠄߥ޿ߎߣ߇␜ໂߐࠇߚ‫ߚߒ߁ߎޕ‬ᣂ⍮⷗ߪ‫ޔ‬
㊁↢㘃ੱ₎ߦ߅ߌࠆੱ₞౒ㅢᗵᨴ∝ߩࠕ࠙࠻ࡉ࡟࡯ࠢࠍ੍᷹ߔࠆ਄ߢ‫⾆ޔ‬㊀ߥᖱႎḮߣߥࠆߣᦼᓙߢ߈ࠆ‫ޕ‬
R) ࠾ࡎࡦࠩ࡞ߦ߅޿ߡⴊዊ᧼ᷫዋ∝ࠍᒁ߈⿠ߎߔࠨ࡞࡟࠻ࡠ࠙ࠗ࡞ࠬ 4 ဳ㧔SRV-4㧕ߩቯ㊂ᴺߩ㐿⊒
૒⮮⧷ᰴ㧘ศᎹ⑍ഥ㧔࠙ࠗ࡞ࠬ⎇ⓥᚲ㧕㧘ችᴛቁᐘ㧔࠙ࠗ࡞ࠬ⎇ⓥᚲ㧕㧘ศ↰෹ᢎ㧘ጟᧄቬ⵨
㔤㐳㘃⎇ⓥᚲౝߢ⊒↢ߒߚ‫ⴊ࡞ࠩࡦࡎ࠾ޟ‬ዊ᧼ᷫዋ∝‫∛ߩޠ‬ේ૕ߪࠨ࡞࡟࠻ࡠ࠙ࠗ࡞ࠬ 4 ဳ㧔SRV-4㧕ߢ޽ࠆߎ
ߣ߇್᣿ߒߚ‫ⷐ∝⊒ߩߎޕ‬࿃ߩ⸃᣿ࠍ⋡⊛ߣߒߡ∛ේᕈߣ࠙ࠗ࡞ࠬ㊂ߩ⋧㑐ᕈࠍ⺞ߴࠆߚ߼‫ޔ‬ᚒ‫ ߪޘ‬SRV-4 ࠥࡁࡓ
ߢ޽ࠆ RNA ࠍࠬ࠲ࡦ࠳࡯࠼ߣߒߡ↪޿ߚ real time RT-PCR ᴺࠍ㐿⊒ߒߚ‫ޕ‬SRV-4 ᗵᨴ࠾ࡎࡦࠩ࡞ߩⴊẏߦ߅޿ߡ⊒
∝୘૕ߣ㕖⊒∝୘૕ߩ࠙ࠗ࡞ࠬ㊂ࠍᲧセߒ‫∝⊒ߚ߹ޔ‬୘૕ߦ߅޿ߡⴊዊ᧼ᢙߣ࠙ࠗ࡞ࠬ㊂ߩ⚻ᤨ⊛ᄌൻࠍ⺞ߴߚߣ
ߎࠈ‫ⴊޔ‬ዊ᧼ᢙ߇ዋߥ޿ߣ࠙ࠗ࡞ࠬ㊂߇ᄙ޿௑ะߦ޽ࠆߣ޿߁੍஻⊛⍮⷗߇ᓧࠄࠇߚ‫੹ޕ‬ᓟߪᬌ૕ᢙߩჇട߅ࠃ߮
ᬌ಴ᣇᴺߩᡷༀࠍⴕ޿‫ࠅࠃޔ‬ᱜ⏕ߥ⸃ᨆࠍㅴ߼ࠆ੍ቯߢ޽ࠆ‫ޕ‬
S) ࡒ࠻ࠦࡦ࠼࡝ࠕࠥࡁࡓߦࠃࠆ࠹࠽ࠟࠩ࡞ߩಽሶ♽⛔ㅴൻ
᧻੗ᷕ㧘I Jahan㧔ㆮવሶᖱႎ㧕㧘Md A Islam (University of Dhaka and Wildlife Trust of Bangladesh)㧘ZM Rahman
(Bangabandhu Sheikh Mujib Safari Park)㧘ᐔ੗໪ਭ㧔ㆮવሶᖱႎ㧕
ోࡒ࠻ࠦࡦ࠼࡝ࠕࠥࡁࡓ㈩೉ࠍ↪޿ߡ‫ ࡞ࠩࠟ࠽࠹ޔ‬4 ዻ㑆ߩ㑐ଥࠍಽሶ♽⛔⸃ᨆߒߚ‫ޕ‬ಽጘᐕઍߩផቯ߆ࠄ‫࠹ޔ‬
࠽ࠟࠩ࡞ߪᭂ߼ߡ⍴޿ᦼ㑆ߦዻ࡟ࡌ࡞ߩ᡼኿⊛ߥ⒳ಽൻ߇߅ߎࠅ‫߇ߣߎߩߎޔ‬ዻ㑆ߩ♽⛔㑐ଥࠍਇ᣿⏕ߦߒߡ޿ࠆ
ߣ⠨߃ࠄࠇࠆ‫↢ޕ‬ᕷ࿾ၞߢ޽ࠆ᧲ධࠕࠫࠕߩඨፉ࡮ፉ‫ޔߢޘ‬ฎઍߩ࠹࠽ࠟࠩ࡞㘃߇ಽᏓࠍᐢߍߥ߇ࠄ⒳ಽൻߒߡ޿
ߞߚ᭽⋧ࠍ↢‛࿾ℂቇ⊛ߦ⠨ኤߒߚ‫ޕ‬
T) 㔤㐳㘃ߩ༦ⷡฃኈ૕ㆮવሶ࡟ࡄ࡯࠻࡝࡯ߩㅴൻ
᧻੗ᷕ㧘ᣂ᧛⧐ੱ㧔᧲੩ක⑼ᱤ⑼ᄢቇ㧕
༦ⷡฃኈߪ‫ޔ‬༦ⷡฃኈ૕㧔OR㧕߇ⅣႺਛߩߦ߅޿‛⾰ࠍಽሶ⹺⼂ߔࠆߎߣߦࠃࠅ㐿ᆎߐࠇࠆ‫ߣ‛↢ߥ⊛ⷡⷞޕ‬
ߐࠇࠆ㔤㐳⋡ߢߪ‫ޔ‬ㅴൻߩㆊ⒟ߢ༦ⷡߩ⋧ኻ⊛ߥ㊀ⷐᕈ߇ૐਅߒ‫ޔ‬༦ⷡᯏ⢻߇ᄢ߈ߊᄬࠊࠇߚߣ޿ࠊࠇࠆ‫ޕ‬㔤㐳⋡
5 ⒳ߣ‫੃ູߩઁޔ‬㘃 8 ⒳ߩ OR ᯏ⢻ㆮવሶߩ⋧หㆮવሶࠍหቯߒ‫ޔ‬OR ㆮવሶߩ࡟ࡄ࡯࠻࡝࡯ࠍᲧセ⸃ᨆߒߚ‫ޕ‬㔤
㐳⋡ 5 ⒳ߩ౒ㅢ␲వᲑ㓏ߢ‫੃ູ↢⃻ߩઁޔ‬㘃 8 ⒳ߣห⒟ᐲߦ߹ߢ OR ㆮવሶߩ࡟ࡄ࡯࠻࡝࡯ࠍᷫዋߐߖߡ߅ࠅ‫⃻ޔ‬
↢㔤㐳⋡ߢߪߘࠇߙࠇߩ♽⛔ߢߐࠄߦ‫ޔ‬ㆮવሶߩ࡟ࡄ࡯࠻࡝࡯ࠍᄬߞߡ޿ࠆߎߣ߇ࠊ߆ߞߚ‫ޔߚ߹ޕ‬㔤㐳⋡ߦ߅ߌ
ࠆ OR ㆮવሶߩ㊀ⶄߪઁߩູ੃㘃ߦᲧߴߡ㕖Ᏹߦዋߥߊ‫ߦ࡯࡝࠻࡯ࡄ࡟ޔ‬ട߃ߡ‫ޔ‬㔤㐳⋡ߢᣂߚߦ߁ߺߛߐࠇࠆ
OR ㆮવሶ߇ᭂ┵ߦዋߥߊߥߞߡ޿ࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬
U) SRV-5 ߩోࠥࡁࡓ㈩೉᳿ቯߣ SRV-5 ․⇣⊛ PCR ᬌᩏᴺߩ㐿⊒
᧻੗ᷕ㧘᧖ᧄᄥ㇢㧘ศᎹ⎸ഥ㧔࠙ࠗ࡞ࠬ⎇ⓥᚲ㧕㧘ችᴛቁᐘ㧔࠙ࠗ࡞ࠬ⎇ⓥᚲ㧕㧘ጟᧄቬ⵨
ⴊዊ᧼ᷫዋ∝ߩේ࿃ߣ⠨߃ࠄࠇࠆࠨ࡞࡟࠻ࡠ࠙ࠗ࡞ࠬ 4 ဳ㧔SRV-4㧕ߦᗵᨴߒߚࡑࠞࠢ୘૕ࠍᬌ಴ߔࠆߚ߼‫ᧃޔ‬
᪳ⴊ↱᧪ߩ DNA ࠍ↪޿ߚ PCR ᬌᩏᴺࠍ⏕┙ߒ‫ޔ‬㔤㐳㘃⎇ⓥᚲߩో㘺⢒୘૕ߩࠬࠢ࡝࡯࠾ࡦࠣࠍⴕߞߡ߈ߚ‫ޕ‬ᣂߚ
ߦ‫ޔ‬ᬌᩏኻ⽎ࠍࠨ࡞࡟࠻ࡠ࠙ࠗ࡞ࠬ 5 ဳ㧔SRV-5㧕ߦ߽᜛ߍࠆߚ߼‫ޔ‬SRV-5 ߩోࠥࡁࡓ㈩೉ࠍ᳿ቯߒ‫ޔ‬SRV-5 ᗵᨴ
୘૕ߩߺࠍᬌ಴ߔࠆ PCR ᴺࠍ㐿⊒ߒߚ‫ ߩߢ߹ࠇߎޔߚ߹ޕ‬SRV-4 ࠍኻ⽎ߣߔࠆᬌᩏߪ SRV-5 ࠍᬌ಴ߔࠆߎߣߪߥ
ߊ‫ ߩࠇߙࠇߘޔ‬SRV ࠲ࠗࡊߦ․⇣⊛ߥᬌᩏᴺߢ޽ࠆߎߣࠍ⏕⹺ߒߚ‫ ࡓࠗ࠲࡞ࠕ࡝ޕ‬PCR ᴺߦࠃࠆ◲ᤃ⊛ߥᗵᐲ⸘
㧙㧙
▚ߦࠃࠆߣ‫ ߩࠄߜߤޔ‬PCR ᬌᩏᴺ߽ᭂ߼ߡ㜞ᗵᐲߢ޽ࠆߎߣ߇ផ᷹ߐࠇߚ‫ޔߚ߹ޕ‬SRV-4‫ޔ‬SRV-5 ᗵᨴ୘૕ࠍ৻ᐲ
ߦᬌ಴ߔࠆ PCR ᬌᩏᴺ߽㐿⊒ߒ‫ߺ⚵ࠍࠄࠇߎޔ‬วࠊߖࠆߎߣߢ‫◲ޔ‬ଢߥᬌᩏߣᔅⷐߦᔕߓߡ SRV ࠲ࠗࡊߩ․ቯࠍ
ⴕ߁ߎߣ߇น⢻ߥᬌᩏࠪࠬ࠹ࡓࠍ⏕┙ߒߚ‫ޕ‬
V) ࡒ࠻ࠦࡦ࠼࡝ࠕࠥࡁࡓߩࠕࡒࡁ㉄⚵ᚑߣઍ⻢ല₸ߦߟ޿ߡߩ⎇ⓥ
ർᷝᐽᒄ㧔㜞⍮ᄢቇ㧕㧘ጯ㊁ᵗਭ㧔᧲੩ᄢቇ㧕㧘㐳⼱Ꮉ᡽⟤㧔ᓳᣤᄢቇ‫⸘⛔ޔ‬ᢙℂ⎇ⓥᚲ㧕㧘᧻੗ᷕ㧘Nick Lane (University
College London)㧘↰ਛ㓷༹㧔᧲੩ㇺ⠧ੱ✚ว⎇ⓥᚲ㧕
↢‛ߩ૕ߩᄢ߈ߐ‫ޔ‬ኼ๮‫ޔ‬ઍ⻢₸ߩ⋧㑐ߪࠃߊ⍮ࠄࠇߡ޿ࠆ߇‫ޔߣࠄࠇߎޔ‬⣄ᬁേ‛ߩࡒ࠻ࠦࡦ࠼࡝ࠕߩ࠲ࡦࡄ
ࠢ⾰ߩ߁ߜ⤑⽾ㅢ㗔ၞߩ⇹᳓ᕈ࡮ⷫ᳓ᕈߩࠕࡒࡁ㉄⚵ᚑߩᄌൻ߇ᒝߊ⋧㑐ߒߡ޿ࠆߎߣࠍ⷗಴ߒߚ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
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Okamoto M, Sako Y, Ito A (2011) Taeniasis/Cyaticercosis in Bali, Indonesia. Southeast Asian Journal of Tropical
Medicine and Public Health 42(4):793-802.
3) ㋈ᧁ᮸ℂ㧘᣿㉿ብᢥ㧘ጟᧄቬ⵨㧘ศ↰෹ᢎ㧘ጟᨋ૒⍮ (2012) SRV-4 ߩ㑐ਈ߇⇼ࠊࠇࠆ࠾ࡎࡦࠩ࡞ߩⴊዊ᧼ᷫ
ዋ∝. ࠝࡌ࡝ࠬࠢ 17:5-10.
ႎ๔
1)
2)
3)
༑ᄙᱜ๺㧘ጟᧄቬ⵨ (2011) ታ㛎േ‛ᗵᨴ∝ߩ⃻⁁ ࠨ࡞࡟࠻ࡠ࠙ࠗ࡞ࠬ 4 ဳ㧔SRV-4㧕 ታ㛎േ‛࠾ࡘ࡯ࠬ
60:32-34.
ጟᧄቬ⵨ (2011) ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ߩේ࿃ⓥ᣿ߦߟ޿ߡ. ታ㛎േ‛කቇ 36:18-19.
ጟᧄቬ⵨ (2011) ੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲߢ⊒↢ߒߚ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ߣߘߩ∛࿃. 㑐⷏ታ㛎േ‛⎇ⓥળ
ળႎ 33:36-40.
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
2)
3)
4)
㋈ᧁ᮸ℂ (2011) ࠨ࡞㘃ߩ∔∛ࠞ࡜࡯ࠕ࠻࡜ࠬ. (ࠨ࡞㘃ߩ∔∛ߣ∛ℂߩߚ߼ߩ⎇ⓥળ✬) p.204 ␠࿅ᴺੱ੍㒐
ⴡ↢දળ⊒ⴕ㧚
Hayakawa T, Varki A (2012) Human-specific Changes in Sialic Acid Biology. (Post-Genome Biology of Primates,
Primatology Monographs) (ed. Hirai H, Imai H, Go Y) p.123-148, Springer.
Hirai H (2012) Evolution and biological meaning genomic wastelands (RCRO): Proposal of hypothesis. (Post-Genome
Biology of primates, Primatology Monographs.) (ed. Hirai H, Imai H, Go Y) p.227-240, Springer.
Matsui A, Hasegawa M (2012) Molecular Phylogeny and Evolution in Primates. (Post-Genome Biology of Primates,
Primatology Monographs) (ed. Hirai H, Imai H, Go Y) p.243-267, Springer.
✬㓸
1)
Hirai H, Imai H, Go Y (2012) Post-Genome Biology of Primates, Primatology Monographs. p.286 Tokyo, Dordrecht,
Heidelberg, London, New York: Springer.
ቇળ⊒⴫
1)
2)
3)
4)
5)
Akari H, Iwasaki Y, Mori K, Ishii K, Maki N, Iijima S, Yoshida T, Okabayashi S, Katakai Y, Young-Jung Lee, Saito A
(2011) Long-term persistent GBV-B infection and development of a progressive chronic hepatitis C-like disease in
marmosets. International Union of Microbiological Society 2011 Congress (2011/09/11-16, Sapporo).
Kusolsuk T, Dekumyoy P, Chaisiri K, Sanguankiat S, Okamoto M, Yanagida T, Sako Y, Komalanisra C, Ito A (2011)
Taenia and Taeniasis: Epidemiological survey and molecular identification in Tha Song Yang district, Tak Province
Thailand. Joint International Tropical Medicine Meeting 2011 (2011/12, Bangkok, Thailand)㧚
Li T, Long C, Nakao M, Chen X, Okamoto M, Giraudoux P, Craig PS, Xiao N, Huang L, Ito A (2011) Usefulness of
pumpkin seeds combining areca nuts in treatment of community-based screened taeniasis carreiers, China. Joint
International Tropical Medicine Meeting 2011 (2011/12, Bangkok, Thailand).
Matsui A, Niimura Y (2011) Comparative evolutionary analyses of olfactory receptor gene repertoires among five
primates and eight non-primate mammal. The 5th International Symposium of the Biodiversity and Evolution Global
COE project "Evolutionary Consequences of Biological Interactions -genome to ecosystem-" (2011/07/09-10, Kyoto,
Japan).
Niimura Y, Matsui A (2011) Diversity of Olfactory Receptor Gene Repertoires among 38 Mammals SMBE annual
meeting (2011/07/26-30, Kyoto, Japan).
㧙㧙
6)
7)
8)
9)
10)
11)
12)
13)
14)
15)
16)
17)
18)
19)
20)
21)
22)
23)
24)
25)
26)
27)
Okamoto M (2011) Hybridization between Taenia asiatica and Taenia saginata and taxonomici status of T. asiatica. Joint
International Tropical Medicine Meeting 201 (2011/12, Bangkok, Thailand).
Saito A, Nomaguchi M, Kono K, Nakayama EE, Shioda T, Yoshida T, Yasutomi Y, Takahashi N, Matano T, Adachi A,
Akari H (2011) Susceptibility of cynomolgus monkeys to monkey-tropic HIV-1 infection is determined by TRIM5
genotypes. 29th Annual Sympojium on Nonhuman Primate Models for AIDS (2011/10/25-28, Seattle)㧚
Saito A, Nomaguchi M, Kono K, Nakayama EE, Shioda T, Yoshida T, Yasutomi Y, Matano T, Adachi A, Akari
H(2011) Genotypic variation of cynomolgus monkey TRIM5 determines the susceptibility to monkey-tropic HIV-1
infection. International Union of Microbiological Society 2011 Congress (2011/09/11-16, Sapporo).
Suda-Hashimoto N, Kaneko A,Kamanaka Y, Okamoto M (2011) Introduction of Management in Open Enclosure and
Case Reports in Group Cage Enrichment for Captive Japanese Macaques. 10th International Conference of
Environmental Enrichment (2011/08/20, Portland US).
Swastica K, Wandra T, Sudarmaja M, Dharmawan NS, Sako Y, Yanagida T, Okamoto M, Sutisna P, Ito A (2011)
Current situation of taeniasis and cysticercosis in Bali, Indonesia. Joint International Tropical Medicine Meeting 2011,
(2011/12, Bangkok, Thailand).
Yoshida T, Okamoto M, Akari H, Suzuki J, Miyabe-Nishiwaki T, Hayakawa T, Imai H, Matsui A, Watanebe A, Kaneko
A, Hirai H (2011) Simian retrovirus-4-associated infectious thrombocytopenia in Japanese macaques. International
Union of Microbiological Society 2011 Congress (2011/09/11-16, Sapporo).
Yoshida T, Okamoto M, Akari H, Suzuki J, Miyabe-Nishiwaki T, Hayakawa T, Imai H, Matsui A, Watanebe A, Kaneko
A, Hirai H (2011) Simian retrovirus-4-associated infectious thrombocytopenia in Japanese macaques. The Unlimited
World Microbes International Union of Microbiological Societies 2011 Congress XV International Congress of
Virology. (2011/09, Sapporo, Japan).
᣿㉿ብᢥ㧘㋈ᧁ᮸ℂ㧘ጟᧄቬ⵨㧘ችㇱ⾆ሶ㧘ᷰㆺᦶ㊁㧘౗ሶ᣿ਭ㧘ᾢፒᷡೣ㧘㒙ㇱ᡽శ㧘㊍ਛᘮᦶ㧘೨↰ౖᒾ㧘
᫪ᧄ⌀ᑿ㧘ᷰㆺ␽ᐔ㧘㗇↰⋥ሶ㧘ᐔ੗໪ਭ㧘᧻ᴛື㇢ (2011) ੩ᄢ㔤㐳⎇ߢ⷗ࠄࠇߚ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝
(1)㧦᭎ⷐႎ๔. ╙ 58 ࿁ᣣᧄታ㛎േ‛ቇળ (2011/05, ᧲੩)㧚
᣿㉿ብᢥ㧘㋈ᧁ᮸ℂ㧘ጟᧄቬ⵨㧘ችㇱ⾆ሶ㧘ᷰㆺᦶ㊁㧘౗ሶ᣿ਭ㧘㒙ㇱ᡽శ㧘㊍ਛᘮᦶ㧘೨↰ౖᒾ㧘᫪ᧄ⌀ᑿ㧘
ᷰㆺ␽ᐔ㧘㗇↰⋥ሶ㧘ᐔ੗໪ਭ㧘᧻ᴛື㇢ (2011) ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ߩ⊒↢ߦ㑐ߔࠆ⚻ㆊ᭎ⷐ. ╙ 152
࿁ᣣᧄ₞කቇળᄢળ (2011/10, ႓㧘ᄢ㒋)㧚
ㇹᐽᐢ㧘⼾↰ᢕ㧘ળᵤᥓᐘ㧘੹੗໪㓶㧘⮮ጊ⑺૒ᄦ㧘ᐔ੗໪ਭ (2011) ࠾ࡎࡦࠩ࡞ࠛࠢ࠰࡯ࡓ⸃ᨆ㨪ታ㛎േ‛
ൻߦ߻ߌߚㆮવ⊛ࡃ࠶ࠢࠣ࡜࠙ࡦ࠼ߩ⸃᣿. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/16-18, ›ጊᏒ)㧚
ᣧᎹᢅਯ㧘Mitra N㧘Wang X㧘Varki N㧘Varki A (2011) ࡅ࠻♽⛔ߦ߅ߌࠆࠪࠕ࡞㉄ฃኈ૕ Siglec-11 ߩㅴൻ. ᣣ
ᧄㅴൻቇળ╙ 13 ࿁ᄢળ (2011/07, ੩ㇺ)㧚
ᐔ੗໪ਭ㧘ᐔ੗⊖วሶ㧘ฎ⾐┨ᒾ㧘㡻Ლବผ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ޽ߞߡࡅ࠻ߦߥ޿ᨴ⦡૕┵ㇱࠥࡁࡓਇᲫ
࿾Ꮺ㧦(1) ሽ࿷᭽ᑼᄌ⇣߆ࠄផ᷹ߐࠇࠆ㕖⋧หᨴ⦡૕㑆ᧃ┵⚵឵߃. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ
(2011/07/16-18, ›ጊᏒ)㧚
ᐔ੗໪ਭ㧘ᐔ੗⊖วሶ㧘ฎ⾐┨ᒾ㧘㡻Ლବผ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ޽ߞߡࡅ࠻ߦߥ޿ᨴ⦡૕┵ㇱࠥࡁࡓਇᲫ
࿾Ꮺ㧦ሽ࿷᭽ᑼᄌ⇣. ╙ 62 ࿁ᨴ⦡૕ቇળ (2011/11/11-13, ᐔႦ)㧚
౗ሶ᣿ਭ㧘ᷰㆺᦶ㊁㧘㗇↰㧔ᯅᧄ㧕⋥ሶ㧘㋈ᧁ᮸ℂ (2011) ૕Ძߦࠃࠅᜰߦ⛉ᛗഃࠍ↢ߓߚ࠾ࡎࡦࠩ࡞੃ఽߩ
∝଀㧚Digit strangulation by hair wrapping in infant Japanese macaques (Macaca fuscata). ╙ 17 ࿁ᣣᧄ㊁↢േ‛කቇ
ળ (2011/09/29-10/02, ᧲੩)㧚
ฎ⾐┨ᒾ㧘ᐔ੗⊖วሶ㧘ᐔ੗໪ਭ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ޽ߞߡࡅ࠻ߦߥ޿ᨴ⦡૕┵ㇱࠥࡁࡓਇᲫ࿾Ꮺ㧦(2) ၭ
㙃⚦⢩ࠍ↪޿ߚ⚵឵߃ߩᬌ಴. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/16-18, ›ጊᏒ)㧚
೨↰ౖᒾ (2011) ‫࠻࠶࠮ࡕ࡯ࡑޟ‬㘺⢒ቶߩᢛ஻‫⥸৻ޠ‬ታ㛎ቶ߆ࠄߩォ឵㧔੐଀ႎ๔㧕. ᣣᧄታ㛎േ‛ᛛⴚ⠪ද
ળ╙ 45 ࿁✚ળ (2011/09/30-10/01, ⋓ጟ)㧚
᧻੗ᷕ㧘Jahan I㧘Ialam MA㧘Rhman ZMM㧘ᐔ੗໪ਭ (2011) ࡒ࠻ࠦࡦ࠼࡝ࠕࠥࡁࡓߦࠃࠆ࠹࠽ࠟࠩ࡞ߩಽሶ♽
⛔ㅴൻ. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/16-18, ›ጊᏒ)㧚
᧻੗ᷕ㧘ᣂ᧛⧐ੱ (2011) 㔤㐳㘃ߩ༦ⷡฃኈ૕ㆮવሶߩ࡟ࡄ࡯࠻࡝࡯ߪዋߥ޿ߩ߆㧫 ᣣᧄㅴൻቇળ╙ 13 ࿁ᄢ
ળ (2011/07/30-31, ੩ㇺ).
ጟᧄቬ⵨㧘ዊ㊁ᢥሶ㧘⮮ᧄᶈੑ㧘㜞㊁ᷕ৻㇢㧘ự㊁ᱜᢘ㧘᫪Ꮉ⨃㧘᳗↰ౖઍ㧘᳓⼱ື਽㧘㈬੗ብᴦ㧘ၳ੗ବብ㧘
ਛደ㓉᣿㧘ਛ᧛᣹ᄥ㧘ችᴛቁᐘ㧘ศᎹ⑍ഥ (2011) ੩ᄢ㔤㐳⎇ߢ⷗ࠄࠇߚ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝(2)㧦ේ࿃࠙
ࠗ࡞ࠬߩหቯߣ⸃ᨆ. ╙ 58 ࿁ᣣᧄታ㛎േ‛ቇળ (2011/05, ᧲੩)㧚
ጟᧄቬ⵨㧘ዊ㊁ᢥሶ㧘⮮ᧄᶈੑ㧘㜞㊁ᷕ৻㇢㧘ự㊁ᱜᢘ㧘᫪Ꮉ⨃㧘᳗↰ౖઍ㧘᳓⼱ື਽㧘㈬੗ብᴦ㧘ၳ੗ବብ㧘
ਛደ㓉᣿㧘ਛ᧛᣹ᄥ㧘ችᴛቁᐘ㧘᧻੗ᷕ (2011) ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ߩේ࿃࠙ࠗ࡞ࠬߩหቯ. ╙ 152 ࿁ᣣ
ᧄ₞කቇળᄢળ (2011/10, ႓㧘ᄢ㒋)㧚
㥱⮮ᥙ㧘ᴡ㊁ஜ㧘ਛጊ⧷⟤㧘⿷┙ᤘᄦ㧘㊁㑆ญ㓷ሶ㧘଻ንᐽብ㧘ୀ㊁ືᦶ㧘Ⴎ↰㆐㓶㧘᣿㉿ብᢥ (2011) ࠨ࡞
ᜰะᕈ HIV-1 ߳ߩᗵฃᕈߦᓇ㗀ࠍਈ߃ࠆࡑࠞࠢࠨ࡞ TRIM5 ㆮવሶߩᄙ᭽ᕈ. ╙ 25 ࿁ᣣᧄࠛࠗ࠭ቇળቇⴚ㓸ળ
(2011/11/30-12/2, ᧲੩)㧚
㗇↰㧔ᯅᧄ㧕⋥ሶ㧘ᄐ⋡ዅᅢ (2011) 㘺⢒ਅ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆᱫఽㆇ៝ߣᱫఽ㘩޿ߩ੐଀. SAGA14 (2011/11/12,
ᾢᧄ)㧚
㧙㧙
28) ㋈ᧁ᮸ℂ㧘᣿㉿ብᢥ㧘ጟᧄቬ⵨㧘ศ↰෹ᢎ㧘ጟᨋ૒⍮ (2011) SRV-4 ߩ㑐ਈ߇⇼ࠊࠇࠆⴊዊ᧼ᷫዋ∝. ╙ 20 ࿁
ࠨ࡞∔∛ࡢ࡯࡚ࠢࠪ࠶ࡊ (2011/07/02, 㤗Ꮣᄢቇ㧔␹ᄹᎹ⋵㧕)㧚
29) 㜞ᯅዏผ㧘㥱⮮ᥙ㧘㊁㑆ญ㓷ሶ㧘᧻ጟ૒❱㧘⿷┙ᤘᄦ㧘᣿㉿ብᢥ㧘ୀ㊁ືᦶ (2011) ࠨ࡞ᜰะᕈ HIV-1 ᗵᨴᘟ
ᕈẜફᦼߩࠞ࠾ࠢࠗࠨ࡞߆ࠄߩᗵᨴᕈ࠙ࠗ࡞ࠬߩ࿁෼. ╙ 25 ࿁ᣣᧄࠛࠗ࠭ቇળቇⴚ㓸ળ (2011/11/30-12/2, ᧲
੩)㧚
30) ┻ౝ᣽↵㧘᧲Ớ◊ᓼ㧘┻ౝ༑ਭሶ㧘’㊁ೋ㖸㧘↰ᴧ⃍ሶ㧘⿷┙ᷕ㧘㜞ᯅ৻ᦶ㧘ᦺ㐳Პ㧘᪢Ỉ᣿ᒄ㧘੉ጟᵗ␭ (2011)
ࡅ࠻㑆⪲♽ᐙ⚦⢩ᩣ㧔UE6E7T-3㧕ߩᒻ⾰ォ឵ㆊ⒟ߦ߅ߌࠆ mRNA ⊒⃻⸃ᨆ. ╙ 34 ࿁ᣣᧄಽሶ↢‛ቇળᐕળ
(2011/12/13-16, ࡄࠪࡈࠖࠦᮮᵿ)㧚
31) ᧲Ớ◊ᓼ㧘ဈᚻ㦖৻㧘㜞ᯅ৻ᦶ㧘⿷┙ᷕ㧘ᦺ㐳Პ㧘଻ንᐽብ㧘੉ጟᵗ␭ (2011) ࠞ࠾ࠢࠗࠩ࡞⊕ⴊ⃿ߦ߅ߌࠆ
⚦⢩ᄖࠞ࡞࡟࠹ࠖࠠࡘ࡝ࡦߦࠃࠆㆮવሶ⊒⃻ߩᄌൻ. ╙ 34 ࿁ᣣᧄಽሶ↢‛ቇળᐕળ (2011/12/13-16, ࡄࠪࡈࠖ
ࠦᮮᵿ)㧚
32) ᷰㆺᦶ㊁㧘౗ሶ᣿ਭ㧘ችㇱ⾆ሶ㧘⷏⣁ᒄ᮸㧔ࠃߒߑ߈േ‛∛㒮㧕㧘㋈ᧁ᮸ℂ㧘⏷๺ᒄ৻㧔ᣣᧄ↢‛⑼ቇ࠮ࡦ࠲
࡯ࢃ㧕 (2011) ‫ޟ‬⣢⤳ߦᄙᢙߩྙ⢩߇⹺߼ࠄࠇߚᘟᕈ⣢ਇోߩࠕࠞࠥࠩ࡞ߩ 1 ଀ߦߟ޿ߡ㧘A case of chronic renal
failure with multiple cysts in a rhesus macaque‫ޠ‬. ╙ 17 ࿁ᣣᧄ㊁↢േ‛කቇળ (2011/09/29-10/02, ᧲੩)㧚
33) ᷰㆺᦶ㊁㧘౗ሶ᣿ਭ㧘ችㇱ⾆ሶ㧘⷏⣁ᒄ᮸㧔ࠃߒߑ߈േ‛∛㒮㧕㧘㋈ᧁ᮸ℂ㧘⏷๺ᒄ৻㧔ᣣᧄ↢‛⑼ቇ࠮ࡦ࠲
࡯ࢃ㧕 (2011) ‫ޟ‬⣢⤳ߦᄙᢙߩྙ⢩߇⹺߼ࠄࠇߚᘟᕈ⣢ਇోߩࠕࠞࠥࠩ࡞ߩ 1 ଀ߦߟ޿ߡ㧘A case of chronic renal
failure with multiple cysts in a rhesus macaque‫ޠ‬. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07/16-18, ›ጊᏒ)㧚
34) ศ↰෹ᢎ㧘ጟᧄቬ⵨㧘᣿㉿ብᢥ㧘੹੗໪㓶㧘᧻੗ᷕ㧘ᣧᎹᢅਯ㧘↢㚤ᥓሶ㧘ષᎹ⟤Ⓞ㧘㥲⮮ᵄሶ㧘ᷰㆺᦶ㊁㧘
౗ሶ᣿ਭ㧘ችㇱ⾆ሶ㧘㋈ᧁ᮸ℂ㧘ᐔ੗໪ਭ (2011) ੩ᄢ㔤㐳⎇ߢ⷗ࠄࠇߚ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝(4)㧦∉ቇ⺞
ᩏ. ╙ 58 ࿁ᣣᧄታ㛎േ‛ቇળ (2011/05, ᧲੩)㧚
35) ศᎹ⑍ഥ㧘૒⮮⧷ᰴ㧘ጟᧄቬ⵨㧘㋈ᧁ᮸ℂ㧘ศ↰෹ᢎ㧘ችᴛቁᐘ (2011) ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝⊒∝ࠩ࡞߆
ࠄߩࠨ࡞࡟࠻ࡠ࠙ࠗ࡞ࠬ 4 ဳߩಽ㔌. ╙ 152 ࿁ᣣᧄ₞කቇળᄢળ (2011/10, ႓㧘ᄢ㒋)㧚
36) દ⮮੫㧘ጟᧄቬ⵨㧘ㄼᐽੳ㧘ᩉ↰ື⍫㧘ਛየ⒤㧘ਛ⼱๺ብ (2012) ࠕࠫࠕߦ߅ߌࠆੱ₞౒ㅢነ↢⯻∛‫∝⯻ྙޔ‬
ኻ╷ߦะߌߚ࿖㓙౒ห⎇ⓥ. ╙ 81 ࿁ᣣᧄነ↢⯻ቇળᄢળ (2012/03, ⷏ች㧘౓ᐶ)㧚
37) ጟᧄቬ⵨ (2012) ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ߩో૕௝. ╙ 1 ࿁ ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ࠪࡦࡐࠫ࠙ࡓ (2012/02,
੩ㇺ)㧚
38) ጟᧄቬ⵨ (2012) 㔤㐳⎇ߩࡑࠞࠢ㘃ߦ⷗ࠄࠇࠆᄌ⇣࡮ⴕേ⇣Ᏹߣࡕ࠺࡞♽⛔ߩ૞⵾. ᐔᚑ 23 ᐕᐲ੩ㇺᄢቇ㔤㐳
㘃⎇ⓥᚲ ౒ห೑↪⎇ⓥળ ⴕേ․ᕈࠍᡰ㈩ߔࠆࠥࡁࡓၮ⋚ߣ⣖ᯏ⢻ߩ⸃᣿ (2012/03, ›ጊ㧘ᗲ⍮)㧚
39) 㥱⮮ᥙ㧘ᴡ㊁ஜ㧘ਛጊ⧷⟤㧘ᣣᩉ┨ᒾ㧘⿷┙ᤘᄦ㧘㊁㑆ญ㓷ሶ㧘଻ንᐽብ㧘ୀ㊁ືᦶ㧘Ⴎ↰㆐㓶㧘ศ↰෹ᢎ㧘
᧲Ớ◊ᓼ㧘↢㚤ᥓሶ㧘Ꮉᧄ⧐㧘㠽ዬ㓉ਃ㧘᣿㉿ብᢥ (2012) ࠞ࠾ࠢࠗࠩ࡞ TRIM5 ㆮવሶࠕ࡝࡞ߩ࿾ℂ⊛ᄙ᭽ᕈ
ߣߘߩᯏ⢻⊛ᗧ⟵. ╙ 153 ࿁ᣣᧄ₞කቇળቇⴚ㓸ળ (2012/03/27-29, ᄢች)㧚
40) ૒⮮⧷ᰴ㧘ศᎹ⑍ഥ㧘ችᴛቁᐘ㧘ศ↰෹ᢎ㧘ጟᧄቬ⵨ (2012) SRV-4 ᗵᨴࠩ࡞ߦ߅ߌࠆ࠙ࠗ࡞ࠬᩭ㉄ߩቯ㊂. ╙
1 ࿁࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ࠪࡦࡐࠫ࠙ࡓ (2012/02/16, ศ↰ᴰᲚ, ੩ㇺ)㧚
41) ศ↰෹ᢎ (2012) ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ߦ߅ߌࠆ∛ℂ⚵❱⸃ᨆ. ╙ 1 ࿁ ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ࠪࡦࡐࠫ࠙
ࡓ (2012/02, ੩ㇺ)㧚
42) ศᎹ⑍ഥ㧘૒⮮⧷ᰴ㧘ጟᧄቬ⵨㧘㋈ᧁ᮸ℂ㧘ศ↰෹ᢎ㧘᣿㉿ብᢥ㧘ਃᶆᥓⴕ㧘ችᴛቁᐘ (2012) ࠾ࡎࡦࠩ࡞ⴊ
ዊ᧼ᷫዋ∝ߩේ࿃࠙ࠗ࡞ࠬߩត⚝. ╙ 153 ࿁ᣣᧄ₞කቇળቇⴚ㓸ળ (2012/03/27-29, ၯ₹)㧚
⻠Ṷ
1)
Kaneko A (2011/05/18-20) Care-taking and veterinarian’s job in KUPRI. Workshop Healthy management for non-human
primates. Faculty of Science, Chulalongkorn University, Bangkok.
2) Kaneko A (2011/05/18-20) Medication, anesthesia and analgesia in nonhuman primates in KUPRI. Workshop Healthy
management for non-human primates. Faculty of Science, ChulalongkornUniversity, Bangkok.
3) Maeda N (2011/05/18-20) Handling of Non-human Primates. Workshop Healthy management for non-human primates.
Faculty of Science, Chulalongkorn University, Bangkok.
4) Maeda N (2011/05/18-20) Housing and Housing condition. Workshop Healthy management for non-human primates.
Faculty of Science, Chulalongkorn University, Bangkok.
5) Okamoto M (2011/05) Bidirectional disease transmission between humans and monkeys and the prevention. Workshop:
Healthy Management for Non-human Primeates. Bangkok, Thailand.
6) Okamoto M (2011/05) Why and how to quarantine the nonhuman primates. Workshop: Healthy Management for
Non-human Primeates. Bangkok, Thailand.
7) ᣿㉿ብᢥ (2011/06/22) ࠛࠗ࠭࠙ࠗ࡞ࠬߩኋਥㆡวᚢ⇛. ੩ㇺᄢቇ࠙ࠗ࡞ࠬ⎇ⓥᚲ࡮࠙ࠗ࡞ࠬ⎇ⓥߩầᵹࠪ࡝࡯
࠭࠮ࡒ࠽࡯. ੩ㇺ㧚
8) ᣿㉿ብᢥ (2011/10/7) 㔤㐳㘃ࡕ࠺࡞േ‛ࠍ↪޿ߚ࠙ࠗ࡞ࠬᗵᨴ∝⎇ⓥ. ᧲੩ක⑼ᱤ⑼ᄢቇ࡮㔍ᴦ∔ᖚ౒ห⎇ⓥ
᜚ὐ⎇ⓥ㓸ળ. ᧲੩㧚
9) ጟᧄቬ⵨ (2011/07) ࠨ࡞ߩነ↢⯻ߣࡅ࠻ߩነ↢⯻㧦౒ㅴൻߣኋਥォ឵. ᐔᚑ 23 ᐕᐲ ੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲ ›
ጊ౏㐿⻠ᐳ 㨬㔤㐳㘃ቇߩᗢߒߺ㨭㧚 ›ጊ, ᗲ⍮㧚
10) ጟᧄቬ⵨ (2011/12) SRV ᗵᨴ∝ኻ╷ߦߟ޿ߡႎ๔ߣߎࠇ߆ࠄߩ⺖㗴. ╙ 8 ࿁౏㐿ࠪࡦࡐࠫ࠙ࡓ‫ࠗࡃ࡞ࠩᧄޟ‬
㧙㧙
ࠝ࡝࠰࡯ࠬࡊࡠࠫࠚࠢ࠻╙ 2 ᦼߩᚑᨐߣ዁᧪ዷᦸ‫ޠ‬. ᧲੩㧚
11) ጟᧄቬ⵨ (2011/12) ࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ߩ∛ᘒߣߘߩ∛࿃. ╙ 4 ࿁ࠨ࡞ࠪࡦࡐࠫ࠙ࡓ. ᄢᵤ, ṑ⾐㧚
12) ᐔ੗໪ਭ (2012/03/19) Chimpanzee chromosomes and gibbon oversea project. Linconl University, UK㧚
࿖㓙౒หవ┵⎇ⓥ࠮ࡦ࠲࡯
㧨⎇ⓥ᭎ⷐ㧪
A) Comparative Wildlife Biology, Conservation, and the Evolution of Social Systems
FB Bercovitch
Our analysis of koala population size, density, and distribution revealed that ecological mathematical models estimating
population abundance were making assumptions that conflict with the available data, so presented erroneous conclusions. The
analysis of giraffe social systems revealed that kinship and sex of giraffe had a significant impact on herd composition.
B) Behaviour, Ecology and Conservation of Forest Bats
DA Hill
Research activity focussed on the development and application of the Autobat acoustic lure for surveying forest bat
diversity at various sites, including Aichi-ken, Yakushima, South Korea, Malaysia and Thailand. I completed two experiments:
an assessment of the effect of trap location (Yakushima) and effectiveness of the lure for enhancing surveys of tropical rain
forest bat diversity (Bukit Panchor, Malaysia). I also made preliminary surveys in South Korea and in rain forest fragments in
southern Thailand. In South Korea we captured five individuals of Murina ussuriensis at three sites. There was only one
previous record of this species for Korea, made in 1956.
C) ࠴ࡦࡄࡦࠫ࡯ߩᲧセ⹺⍮⊒㆐⎇ⓥ
⿷┙ᐞ⏴
࠴ࡦࡄࡦࠫ࡯‫ࠍఽ੃࡞ࠩࠢࠞࡑޔ‬ኻ⽎ߦ‫␠ޔ‬ળ⊛⹺⍮⢻ജ‫ߦߊߣޔ‬㗻⍮ⷡ᭽ᑼ࡮୘૕ᖱႎߩⷞ⡬ⷡ⛔วߦ߆߆ࠊ
ࠆᲧセ⊒㆐⎇ⓥࠍ߅ߎߥߞߚ‫⺖࠲࡯ࡘࡇࡦࠦߩဳ✵⸠ޕ‬㗴ߦࠃࠆᚑ❣⹏ଔ߅ࠃ߮‫ޔ‬ฦ⒳ߩⷞⷡೝỗឭ␜ᤨߩᵈⷞⴕ
േߩಽᨆⷞ✢ߩ⸘᷹ࠍ߅ߎߥߞߚ‫ޕ‬
D) േ‛࿦ߩ࠴ࡦࡄࡦࠫ࡯ߩ⍮ᕈߩ⎇ⓥ
⿷┙ᐞ⏴
ฬฎደᏒߩ᧲ጊേ‛࿦ߩ࠴ࡦࡄࡦࠫ࡯1 ⟲ 5 ୘૕ࠍኻ⽎ߦ‫ޔ‬ደᄖㆇേ႐ߢߩ␠ળⴕേࠍⷰኤ⸥㍳ߒߚ‫ޔߚ߹ޕ‬
‫ࡄޟ‬
ࡦ࡜ࡏ‫ߣޠ‬ฬߠߌࠄࠇߚࡉ࡯ࠬߦ߅޿ߡ‫ޔ‬㆏ౕ૶↪߿ࠦࡦࡇࡘ࡯࠲⺖㗴ࠍߣ߅ߒߡᓐࠄߩ⍮ᕈࠍಽᨆߒߚ‫ޕ‬
E) Captive Chimpanzee Group Formation and Integration of Individuals into Existing Social Groups
M Seres
Continue working at the Kumamoto Sanctuary where I have been assisting in several ongoing group formation projects.
Collecting behavioral data using tablet PC on these newly formed and assembled groups. Additionally to our ongoing
Sanctuary projects, I have been acting as an adviser at various Zoo projects keeping and reassembling their Chimpanzee
groups in Japan as well as on International level in the U.S.A., the Netherlands, Spain, Poland and in Hungary.
F) Sources of Variation across Individual Primate Hosts in Parasite Infection
AJJ MacIntosh
I visited Yakushima in September, 2011, to collect fecal samples and have since begun testing for variation in infection in
relation to social network position, fecal cortisol, fecal testosterone, and fecal antibodies specific to nematode species
infecting Japanese macaques.
G) Fractal Analysis of Behavior as an Indicator of Individual and Environmental Quality
AJJ MacIntosh
I visited the Institut Pluridisciplinaire Hubert Curien of the Centre National de la Recherche Scientifique at the Université
de Strasbourg, France, to conduct collaborative research analyzing penguin behavior using fractal techniques.
H) ࡉ࡯࠲ࡦߦ㑐ߔࠆ⺞ᩏ⎇ⓥ
⷏Ỉ๺ሶ
ࡉ࡯࠲ࡦ₺࿖ߩ National Referral Hospital ߦߡᣂ↢ఽ⸻≮ߦ៤ࠊࠅ‫⃻ޔ‬࿾ߩᣂ↢ఽක≮ߩ⃻⁁ߣ⺖㗴ߦߟ߈⺞ᩏ⎇
ⓥࠍⴕߞߚ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
2)
Adachi I, Hampton RR (2011) Rhesus monkeys see who they hear: Spontaneous cross-modal memory for familiar
conspecifics. PLoS ONE 6(8):1-8, doi:10.1371/journal.pone.0023345.
Ellis W, Bercovitch FB (2011) Body size and sexual selection in the koala. Behavioral Ecology and Sociobiology
65:1229-1235.
㧙㧙
3)
4)
5)
6)
7)
8)
9)
Ellis W, Bercovitch FB, FitzGibbon S, Roe P, Wimmer J, Melzer A, Wilson R (2011) Koala bellows and their
association with the spatial dynamics of free ranging koalas. Behavioral Ecology 22:372-377.
Higgins AL, Bercovitch FB, Tobey JR, Andrus CH (2011) Dietary specialization and Eucalyptus species preferences in
Queensland koalas. Zoo Biology 30:52-58.
Ludwig V*, Adachi I*, Matsuzawa T (2011) Can you see sounds? Chimpanzees associate high auditory pitch with
visual lightness. Proceedings of the National Academy of Sciences 108(51):20661-20665, doi:10.1073/pnas.1112605108.
* These authors contributed equally to this work and co-corresponding authors of the paper.
MacIntosh AJJ, Alados CL, Huffman MA (2011) Fractal analysis of behavior in a wild primate: behavioral complexity
in health and disease. Journal of the Royal Society Interface 8(63):497-509.
Rothwell ES, Bercovitch FB, Andrews J, Anderson MJ (2011) Estimating daily walking distance of captive African
elephants using an accelerometer. Zoo Biology 30:579-591.
MacIntosh AJJ, Huffman MA, Nishiwaki K, Miyabe-Nishiwaki T (2012) Urological screening of a wild group of
Japanese macaques (Macaca fuscata yakui): investigating trends in nutrition and health. International Journal of
Primatology 33:460-478.
⷏Ỉ๺ሶ (2012) ࡉ࡯࠲ࡦ₺࿖ߦ߅ߌࠆᣂ↢ఽක≮ߩ⃻⁁ߣ⺖㗴㧦㊀∝ᣂ↢ఽ 3 ଀ߩᴦ≮⚻㛎ࠍㅢߒߡ. ࡅࡑ
࡜ࡗቇ⹹ 13:254-264.
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
Huffman MA, MacIntosh AJJ (2012) Plant-food diet of the Arashiyama Japanese macaques and its potential medicinal
value. (The Monkeys of Stormy Mountain: 60 Years of Primatological Research on the Japanese Macaques of
Arashiyama) (ed. Leca JB, Huffman MA, Vasey P) p.356-432 Cambridge University Press.
ߘߩઁߩၫ╩
1)
2)
3)
⷏Ỉ๺ሶ (2012) ᣂ↢ఽ⑼කᏧ㧘㔗㦖ߩ࿖߳ ᐘߖߩ࿖ࡉ࡯࠲ࡦߢ⿒ߜ߾ࠎߣ↢߈ࠆ. ࡀࠝࡀࠗ࠲࡞ࠤࠕ 25(1)
p.102-103 ࡔ࠺ࠖࠞ಴ 㧚
⷏Ỉ๺ሶ (2012) ᣂ↢ఽ⑼කᏧ㧘㔗㦖ߩ࿖߳ ᐘߖߩ࿖ࡉ࡯࠲ࡦߢ⿒ߜ߾ࠎߣ↢߈ࠆ. ࡀࠝࡀࠗ࠲࡞ࠤࠕ 25(2)
p.64-65 ࡔ࠺ࠖࠞ಴ 㧚
⷏Ỉ๺ሶ (2012) ᣂ↢ఽ⑼කᏧ㧘㔗㦖ߩ࿖߳ ᐘߖߩ࿖ࡉ࡯࠲ࡦߢ⿒ߜ߾ࠎߣ↢߈ࠆ. ࡀࠝࡀࠗ࠲࡞ࠤࠕ 25(3)
p.54-55 ࡔ࠺ࠖࠞ಴ 㧚
ቇળ⊒⴫
1)
2)
3)
4)
5)
6)
7)
8)
9)
10)
11)
12)
13)
14)
Bercovitch FB, Berry PSM (2011) Social associations and kinship in the Thornicroft’s giraffe of Zambia. 1st ‘Wild’
Giraffe Indaba (2011/07/04-07, Etosha, Namibia)㧚
Fukui D, Hill DA, Matsumura S (2011) Roosting behaviour of nursing females in the Ussurian tube-nosed bat Murina
ussureinsis. 2nd International South-East Asian Bat Conference (2011/06/06-09, Bogor, Indonesia).
Hill DA (2011) Application of an acoustic lure to enhance forest surveys of bats. 2nd International South-East Asian Bat
Conference (2011/06/06-09, Bogor, Indonesia).
MacIntosh AJJ, Jacobs A, Huffman MA, Hernandez AD (2011) Parasite transmission through social networks of
Japanese macaques: a cost of grooming? The 27th Congress Primate Society of Japan (2011/07/16-18, Inuyama).
Seres M (2011) Chimpanzee introductions and group formations. The past, the present, with suggestions for the future.
SAGA14 (2011/11/12-13, Kumamoto).
⿷┙ᐞ⏴ (2011) ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆࠨ࠶࠴ࡖ࡯㍲ⷞ. ᣣᧄᔃℂቇળ╙ 75 ࿁ᄢળ (2011/09/15-17, ᣣᧄᄢቇ)㧚
⿷┙ᐞ⏴ (2011) ࡅ࠻ߣ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ“౒ᗵⷡ”ߩᲧセಽᨆ 2. ANIMAL2011 (2011/09/08-11, ᘮᙥ⟵Ⴖ
ᄢቇ)㧚
⿷┙ᐞ⏴㧘෹᳗㓷Ꮖ㧘᧻ᴛື㇢ (2011) ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ㗻ో૕ಣℂߩ⊒㆐. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ
(2011/07/16-18, ›ጊ࿖㓙ⷰశ࠮ࡦ࠲࡯ࡈࡠࠗ࠺)㧚
⿷┙ᐞ⏴㧘෹᳗㓷Ꮖ㧘᧻ᴛື㇢ (2011) ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ㗻⍮ⷡ᭽ᑼߩ⊒㆐. ᣣᧄ⿒ߜ߾ࠎቇળ╙ 11 ࿁ቇⴚ
㓸ળ (2011/05/07-08, ਛㇱቇ㒮ᄢቇ)㧚
ᧁ᧛రᄢ㧘ᰞᐸ㓁ሶ㧘Ꮢ㊁ᖝሶ㧘ፉ↰߆ߥ߃㧘㋈ᧁஜᄥ㧘ᷰㆻߺߥߺ㧘ㄭ⮮⵨ᴦ㧘ᧁ᧛ᐘ৻㧘⿷┙ᐞ⏴ (2011)
᧲ጊേ‛࿦ߩ࠴ࡦࡄࡦࠫ࡯࠲ࡢ࡯೑↪⁁ᴫߩ⛮⛯⺞ᩏ. SAGA14 (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
᳓㊁૫✜㉿㧘⿷┙ᐞ⏴ (2011) ᣣᧄߦ߅ߌࠆ࠱࠙ߩၞᄖ଻ోߩ੹ᓟ㨪ో࿖ߩ㘺⢒ਅ࠱࠙ߩಽᏓ࿑߆ࠄ㨪.
SAGA14 (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
ፉ↰߆ߥ߃㧘ᰞᐸ㓁ሶ㧘Ꮢ㊁ᖝሶ㧘ᧁ᧛రᄢ㧘㋈ᧁஜᄥ㧘ᷰㆺߺߥߺ㧘ㄭ⮮␭ᴦ㧘ᧁ᧛ᐘ৻㧘⿷┙ᐞ⏴ (2011)
ዊဳዷ␜ᣉ⸳“ࡄࡦ࡜ࡏ”ߩ⹏ଔ㧙࠴ࡦࡄࡦࠫ࡯ߩ೑↪₸࡮ⴕേߩᄌൻࠍㅢߒߡ㧙. SAGA14 (2011/11/12-13, ᾢ
ᧄᏒേᬀ‛࿦)㧚
㋈ᧁஜᄥ㧘ᰞᐸ㓁ሶ㧘Ꮢ㊁ᖝሶ㧘ᧁ᧛రᄢ㧘ፉ↰߆ߥ߃㧘ᷰㆺߺߥߺ㧘ㄭ⮮␭ᴦ㧘ᧁ᧛ᐘ৻㧘⿷┙ᐞ⏴ (2011)
᧲ጊേ‛࿦ߩ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ⹺⍮ታ㛎ෳട₸ߩᄌേ. SAGA14 (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
ᷰㆺߺߥߺ㧘Ꮢ㊁ᖝሶ㧘ᧁ᧛రᄢ㧘ᰞᐸ㓁ሶ㧘ፉ↰߆ߥ߃㧘㋈ᧁஜᄥ㧘ㄭ⮮␭ᴦ㧘ᧁ᧛ᐘ৻㧘⿷┙ᐞ⏴ (2011)
᧲ጊേ‛࿦ߢߩ࠴ࡦࡄࡦࠫ࡯ߩ⍮ᕈዷ␜. SAGA14 (2011/11/12-13, ᾢᧄᏒേᬀ‛࿦)㧚
㧙㧙
⻠Ṷ
1)
2)
3)
4)
5)
6)
7)
Hill DA (2011/11/09) Applications of an acoustic lure for surveying and studying bats in woodlands. Wildlife Research
Center, Kyoto University, Kyoto.
MacIntosh AJJ (2011/06/11) Epidemiology of nematode parasite infection among wild Japanese macaques:
heterogeneity in the external and internal environments. The 12th Annual Japanese Macaque Symposium, Primate
Research Institute Inuyama.
MacIntosh AJJ (2011/09/04) Of worms and monkeys: the secret struggle for health in the wild. “Yakushima Kenkyuu
Koza”, Yakushima㧚
Seres M (2011/11/22) Chimpanzee (Pan troglodytes) introductions and group formations in captivity. Yerkes National
Primate Research Center, Emory University Atlanta, USA.
Adachi I (2012/01/06) Social recognition in nonhuman primates. International Conference Looking Within:
Interdisciplinary Approaches to Consciousness, National Institute of Advanced Studies, Bangalore, India.
Bercovitch FB (2012/03/16) Giraffe society and human evolution. The City University of New York New York, USA.
MacIntosh AJJ (2012/02/02) A fractal ethos for ethology: revealing behavioral stereotypies in stress and disease. German
Primate Center, Gottingen, Germany.
⊕⋲ࡊࡠࠫࠚࠢ࠻
㧨⎇ⓥ᭎ⷐ㧪
A) ⴫ᖱࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߦߟ޿ߡߩታ㛎ᔃℂቇ⊛⎇ⓥ
૒⮮ᒎ
⴫ᖱ߿ⷞ✢ߦࠃࠆኻੱࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߦ߅ߌࠆᖱႎಣℂㆊ⒟ࠍ‫ޔ‬෻ᔕ⸥㍳࡮ࡆ࠺ࠝ㍳↹࡮╭㔚࿑⸘᷹ߥߤߦࠃ
ࠅᬌ⸛ߒߚ‫ޕ‬ቯဳ⊒㆐⠪߅ࠃ߮⊒㆐㓚ኂ⠪ࠍኻ⽎ߣߒߚ‫ޕ‬
B) ⴫ᖱࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߦߟ޿ߡߩ␹⚻⑼ቇ⊛⎇ⓥ
૒⮮ᒎ㧘ᴡౝጊ㓉♿㧘Ỉ↰⃍ሶ
⴫ᖱ߿ⷞ✢ߦࠃࠆኻੱࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦ⺖㗴ࠍㆀⴕਛߩ␹⚻ᵴേࠍ‫ޔ‬fMRI࡮ᷓㇱ⣖ᵄߥߤࠍ↪޿ߡ⸘᷹ߒߚ‫ޕ‬
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
2)
3)
4)
5)
✚⺑
1)
2)
3)
ႎ๔
1)
2)
3)
4)
Sato W, Kochiyama T, Uono S, Matsuda K, Usui K, Inoue Y, Toichi M (2011) Rapid amygdala gamma oscillations in
response to eye gaze. PLoS One 6:e28188.
Uono S, Sato W, Toichi M (2011) The specific impairment of fearful expression recognition and its atypical
development in pervasive developmental disorder. Social Neuroscience 6(5-6):452-463.
Fujimura T, Sato W, Okanoya K (2012) Subcategories of positive emotion. Psychologia 55(1):1-8.
Okada T, Sato W, Kubota Y, Toichi M, Murai T (2012) Right hemispheric dominance and interhemispheric cooperation
in reflexive attentional shift by gaze. Psychiatry and Clinical Neurosciences 66(2):97-104.
Sato W, Kochiyama T, Uono S, Matsuda K, Usui K, Inoue Y, Toichi M (2012) Temporal profile of amygdala gamma
oscillations in response to faces. Journal of Cognitive Neuroscience 24(6):1420-1433.
ᴡౝጊ㓉♿ (2011) ᯏ⢻⊛⏛᳇౒㡆↹௝ᴺߦࠃࠆ㗔ၞ㑆⚿วಽᨆ. ␹⚻ᔃℂቇ 27(1):35-46.
૒⮮ᒎ (2011) 㗻ࠍಣℂߔࠆ⣖ᵴേߩᤨⓨ㑆ࡄ࠲ࡦ. ၮ␆ᔃℂቇ⎇ⓥ 29(2):171-175.
૒⮮ᒎ (2011) り૕ㆇേ⍮ⷡߦ߅ߌࠆᓟㇱ਄஥㗡Ḵߩ߰ࠆ߹޿. ࡌࡆ࡯ࠨࠗࠛࡦࠬ 10:19-20.
ᴡౝጊ㓉♿㧘↰ㆻብ᮸ (2011) ⸘▚⺰⊛⸃೬ቇࠍ೑↪ߒߚൻ⍹⣖㗡⬄㛽߆ࠄ⣖ታ⾰ߩౣ᭴ᚑ. Proceedings of the
5th Conference on the Replacement of Neanderthals by Modern Humans (RNMH) project: 88-89.
ᴡ ౝ ጊ 㓉 ♿ 㧘 ↰ ㆻ ብ ᮸ (2011) ⸘ ▚ ⺰ ⊛ ⸃ ೬ ቇ ࠍ ೑ ↪ ߒ ߚ ൻ ⍹ ⣖ 㗡 ⬄ 㛽 ߆ ࠄ ⣖ ታ ⾰ ߩ ౣ ᭴ ᚑ ߩ ⹜ ߺ.
Proceedings of the 3rd Conference on the Replacement of Neanderthals by Modern Humans (RNMH) project: 56-57.
ᴡ ౝ ጊ 㓉 ♿ 㧘 ↰ ㆻ ብ ᮸ (2011) ⸘ ▚ ⺰ ⊛ ⸃ ೬ ቇ ࠍ ೑ ↪ ߒ ߚ ൻ ⍹ ⣖ 㗡 ⬄ 㛽 ߆ ࠄ ⣖ ታ ⾰ ߩ ౣ ᭴ ᚑ ߩ ⹜ ߺ.
Proceedings of the 4th Conference on the Replacement of Neanderthals by Modern Humans (RNMH) project: 134-136.
↰ㆻብ᮸㧘ᴡౝጊ㓉♿ (2011) ⃻ઍੱ⣖ᯏ⢻࿾࿑ߩൻ⍹⣖߳ߩ౮௝ᴺ߳ߩ╙৻ᱠ. Proceedings of the 2nd
Conference on the Replacement of Neanderthals by Modern Humans (RNMH) project: 56-57.
ቇળ⊒⴫
1)
2)
Aizawa E, Kochiyama T, Sato Y, Morishita J, Sekiguchi A, Kotozaki Y, Miyazaki A, Kano M, Kanazawa M, Sugiura M,
Kawashima R, Mushiake H, Fukudo S (2011) Increased Activation of ventromedial Prefrontal Cortex during Decision.
╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/17, ᮮᵿ).
Aizawa E, Sato Y, Kochiyama T, Morishita J, Kano M, Kanazawa M, Hongo M, Fukudo S (2011) Neural Substrates of
Decision Making in Irritable Bowel Syndrome. Digestive Disease Week 2011 (2011/05/07, Chicago, USA).
㧙㧙
3)
Kitada R, Okamoto Y, Sasaki AT, Kochiyama T, Miyahara M, Lederman S, Sadato N (2011) Brain network involved in
the recognition of facial expressions of emotion in the early blind. 12th International Multisensory Research Forum
(2011/10/19, Fukuoka, Japan).
4) Okamoto Y, Kosaka H, Kitada R, Tanabe HC, Munesue T, Ishitobi M, Hayashi JM, Saito DN, Yanaka HT, Kochiyama
T, Omori M, Wada Y, Okazawa H, Sadato N (2011) The EBA dysfunction in the ASD; as a " comparator " of self and
other's action during reciprocal imitation. ╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ (2011/09/17, ᮮᵿ).
5) Sato W (2011) Temporal profile of amygdala activities in response to faces and emotional facial expressions.
International Workshop on "Perception and Expression of Emotions" (2011/07/25, Kyoto, Japan).
6) Sato W (2011) Temporal profile of amygdala activity in response to emotional facial expressions. International Society
for Research on Emotion 2011 (2011/07/28, Kyoto, Japan).
7) Yanaka H, Saito DN, Kochiyama T, Fujii T, Kosaka H, Shimada T, Asai T, Okazawa H (2011) Inhibition load changes
the brain activation irrespective of attentional differences: a functional magnetic resonance imaging study. ╙ 34 ࿁ᣣᧄ
␹⚻⑼ቇᄢળ (2011/09/17, ᮮᵿ)㧚
8) ૒⮮ᒎ (2011) േ⊛⴫ᖱࠍಣℂߔࠆᔃℂ࡮␹⚻ࡔࠞ࠾࠭ࡓ. ╙ 75 ࿁ᣣᧄᔃℂቇળ (2011/09/15, ᧲੩)㧚
9) ጟᧄᖘሶ㧘ዊဈᶈ㓉㧘ർ↰੫㧘↰ㆻብ᮸㧘᫟ዬବᄦ㧘⍹㘧ା㧘ᨋᱜ㆏㧘㥲⮮ᄢテ㧘ᴡౝጊ㓉♿㧘⼱ਛਭ๺㧘
ᄢ᫪᥏ᄦ㧘๺↰᦭ม㧘ጟᴛ⑲ᒾ㧘ቯ⮮ⷙᒄ (2011) ⥄㐽∝ࠬࡍࠢ࠻࡜ࡓߦ߅ߌࠆ EBA ߩᯏ⢻ૐਅ㧙⋧੕ᮨ୮
ᤨߩ⥄ઁߩേ૞ߩᲧセߦ㑐ࠊࠆ␹⚻ၮ⋚. ╙ 13 ࿁ᣣᧄࡅ࠻⣖ᯏ⢻ࡑ࠶ࡇࡦࠣቇળ (2011/09/02, ੩ㇺ)㧚
10) ૒‫┨ᧁޘ‬ብ㧘ᴡౝጊ㓉♿㧘᧖ᶆర੫㧘↰ㆻብ᮸㧘ቯ⮮ⷙᒄ (2011) ⷞⷡㆇേ⊛ߥേ૞⴫⽎ࠍᜂ߁⣖ౝࡀ࠶࠻ࡢ
࡯ࠢ. ╙ 13 ࿁ᣣᧄࡅ࠻⣖ᯏ⢻ࡑ࠶ࡇࡦࠣቇળ (2011/09/01, ੩ㇺ)㧚
11) ศ↰ఝ⟤ሶ㧘↰ㆻብ᮸㧘ᨋᱜ㆏㧘ᴡౝጊ㓉♿㧘ቯ⮮ⷙᒄ (2011) ੍๔ലᨐߦ߅ߌࠆ⵬⿷ㆇേ೨㊁ࠍਛᔃߣߔࠆ
ࡀ࠶࠻ࡢ࡯ࠢ㧦ᯏ⢻⊛ MRI ⎇ⓥ. ╙ 13 ࿁ᣣᧄࡅ࠻⣖ᯏ⢻ࡑ࠶ࡇࡦࠣቇળ (2011/09/01, ੩ㇺ)
⻠Ṷ
1) ᴡౝጊ㓉♿ (2011/07/23) SPM ߦࠃࠆ⣖ᯏ⢻↹௝࠺࡯࠲⸃ᨆ. ╙ 14 ࿁ᣣᧄశ⣖ᯏ⢻ࠗࡔ࡯ࠫࡦࠣ⎇ⓥળ. ᧲
੩࡮ᤊ㒺ળ㙚㧚
2) ᴡౝጊ㓉♿ (2011/10/30) ⣖ࠍ⺞ߴࠆᣂᛛⴚ㨪⣖ᵴേߩ႐ᚲࠍ․ቯߒࠃ߁. ╙ 26 ࿁࿖᳃ᢥൻ⑂࡮੩ㇺ 2011࡮᣿
ᣣߩ᥵ࠄߒߩᢥൻዷ. ੩ㇺ࡮ߌ޿ߪࠎߥࡊ࡜ࠩ㧚
3) ᴡౝጊ㓉♿ (2011/11/20) ↢૕⸘᷹࠺࡯࠲ߩᤨ♽೉⸃ᨆ. ᧲ർᄢቇᄢቇ㒮⢇૕ਇ⥄↱ቇಽ㊁࠮ࡒ࠽࡯. ઄บ࡮᧲
ർᄢቇ㧚
4) ᴡౝጊ㓉♿ (2011/12/05) ᯏ⢻⊛⏛᳇౒㡆↹௝ᴺߣ༦ⷡ࡮๧ⷡ⎇ⓥ߳ߩᔕ↪. ᔕ↪⣖⑼ቇࠦࡦ࠰࡯ࠪࠕࡓ. ᧲
੩࡮ࠕࠨࡅࡆ࡯࡞๋ᆄᯅࡆ࡞㧚
ࡏࡁࡏ㧔ᨋේ㧕⎇ⓥㇱ㐷㧦2011 ᐕ 8 ᦬߹ߢ
ࡅ࠻⑼ 3 ⒳Ყセ⎇ⓥࡊࡠࠫࠚࠢ࠻㧦2011 ᐕ 9 ᦬㨪
㧨⎇ⓥ᭎ⷐ㧪
A) ㊁↢࠴ࡦࡄࡦࠫ࡯ߩⴕേ⺞ᩏ
᧻ᴛື㇢㧘ጊᧄ⌀਽
ࠡ࠾ࠕ౒๺࿖ࡏ࠶࠰࠙᧛ߦߡ‫ޔ‬㊁↢࠴ࡦࡄࡦࠫ࡯ߩⴕേߣ↢ᘒࠍ⺞ᩏߒߚ‫ޕ‬㆏ౕ૶↪ⴕേ‫␲ޔ‬Უߦࠃࠆሶ⢒ߡද
ജ‫ޔ‬㓸࿅ߢߩදജ࡮ᓎഀಽᜂ‫ޔ‬ᬀ‛ߩ⮎↪೑↪ⴕേߥߤࠍ⸥㍳ߒ‫⸃ޔ‬ᨆࠍ߅ߎߥߞߚ‫ޕ‬
B) ㊁↢ࡏࡁࡏߩⴕേ⺞ᩏ
ጊᧄ⌀਽
ࠦࡦࠧ᳃ਥ౒๺࿖ࡢࡦࡃ᧛ߦߡ‫ޔ‬㊁↢ࡏࡁࡏߩⴕേߣ↢ᘒࠍ⺞ᩏߒߚ‫ޕ‬㘩‛ಽ㈩ࠍ฽߻୘૕㑆㑐ଥ‫ޔ‬ㆊ଻⼔੐଀
ࠍ฽߻Უሶ⊒㆐‫ޔ‬㓸࿅ߢߩදജ࡮ᓎഀಽᜂߥߤࠍ⸥㍳ߒ‫⸃ޔ‬ᨆࠍ߅ߎߥߞߚ‫ޕ‬
C) ㊁↢ࠧ࡝࡜ߩⴕേ⺞ᩏ
᧻ᴛື㇢㧘ᐔ↰⡡
࡞ࡢࡦ࠳౒๺࿖ࡧࠜ࡞ࠞࡦ࿖┙౏࿦ߦߡ‫ޔ‬㊁↢ࠧ࡝࡜ߩⴕേࠍ⺞ᩏߒߚ‫ޕ‬න㓶ⶄ㓽ߩ᭴ᚑߢ޽ࠆ⸘ 3 ⟲ࠍㅊ〔ⷰ
ኤߒ‫ޔ‬୘૕㑆੤ᷤ߿ណ㘩ⴕേߦߟ޿ߡ⸥㍳ߔࠆߣหᤨߦ‫ߡ޿ߟߦ⁁⃻ߩࡓ࠭࡝࡯࠷ࠦࠛޔ‬ᖱႎ෼㓸ߒߚ‫ޕ‬
D) 㘺⢒࠴ࡦࡄࡦࠫ࡯ࠍኻ⽎ߣߒߚᲧセ⹺⍮⑼ቇ⊛⎇ⓥ
᧻ᴛື㇢㧘ᐔ↰⡡㧘ጊᧄ⌀਽
㔤㐳㘃⎇ⓥᚲߩ 14 ୘૕‫ޔ‬ᨋේ㘃ੱ₎⎇ⓥ࠮ࡦ࠲࡯ߩ 8 ୘૕‫ޔ‬ᾢᧄࠨࡦࠢ࠴ࡘࠕ࡝ߩ 51 ୘૕ߩ࠴ࡦࡄࡦࠫ࡯ࠍኻ
⽎ߦ‫⹺ߚ޿↪ࠍ࡯࠲࠾ࡕ࡞ࡀࡄ࠴࠶࠲ޔ‬⍮⎇ⓥ‫ޔ‬㕖᜔᧤ဳࠕࠗ࠻࡜࠶ࠞ࡯ࠍ↪޿ߚⷞ✢ࡄ࠲࡯ࡦߩ⸥㍳‫ޔ‬㆏ౕ૶↪
ⴕേߩታ㛎࡮ⷰኤ‫ޔ‬୘૕㑆␠ળ੤ᷤߩታ㛎࡮ⷰኤߥߤࠍ߅ߎߥߞߚ‫ޕ‬
E) ᣣᧄ߳ߩࡏࡁࡏዉ౉ߦะߌߚḰ஻‫⾗ޔ‬ᢱ෼㓸‫ޔ‬ᶏᄖ㘺⢒ࡏࡁࡏ୘૕⟲ߩⷰኤ
ᐔ↰⡡㧘ጊᧄ⌀਽
㧙㧙
ᣣᧄߢߩ㘺⢒ࡏࡁࡏࠍኻ⽎ߦߒߚ⹺⍮⎇ⓥࠍ┙ߜ਄ߍࠆḰ஻ߣߒߡ‫ޔ‬ዉ౉ᚻ⛯߈ߩ⺞ᩏ‫ޔ‬ᶏᄖߢߩࡏࡁࡏ㘺⢒࡮
ታ㛎⎇ⓥߩ⃻႐ⷞኤ‫ޔ‬㘺⢒ࡏࡁࡏ୘૕ߦ߆ࠎߔࠆᖱႎ෼㓸ࠍ߅ߎߥߞߚ‫ޕ‬
㧨᧻ᴛື㇢ߩᬺ❣ߦߟ޿ߡߪᕁ⠨⸒⺆ಽ㊁ࠍෳᾖ㧪
㧨⎇ⓥᬺ❣㧪
ේ⪺⺰ᢥ
1)
2)
3)
4)
5)
6)
✚⺑
1)
Hirata S, Fuwa K, Sugama K, Kusunoki K, Takeshita H (2011) Mechanism of birth in chimpanzees: humans are not
unique among primates. Biology Letters 7:686-688 (doi: 10.1098/rsbl.2011.0214).
Hirata S, Matsuda G, Ueno A, Fuwa K, Sugama K, Kusunoki K, Fukushima H, Hiraki K, Tomonaga M, Hasegawa T
(2011) Event-related potentials in response to subjects’ own names: A comparison between humans and a chimpanzee.
Communicative & Integrative Biology 4(3):321-323(doi: 10.4161/cib.4.3.14841).
Kano F, Hirata S, Call J, Tomonaga M (2011) The visual strategy specific to humans among hominids: A study using the
gap–overlap paradigm. Vision Research 51:2348-2355(doi: 10.1016/j.visres.2011.09.006).
Bril B, Smaers J, Steele J, Rein R, Nonaka T, Dietrich G, Biryukova E, Hirata S, Roux V (2012) Functional mastery of
percussive technology in nut-cracking and stone-flaking actions: experimental comparison and implications for the
evolution of the human brain. The Philosophical Transactions of the Royal Society B: Biological Sciences,
367:59-74(doi: 10.1098/rstb.2011.0147).
Myowa-Yamakoshi M, Scola C, Hirata S (2012) Humans and chimpanzees attend differently to goal-directed actions.
Nature Communications 3:693 (doi: 10.1038/ncomms1695).
Yamamoto S, Humle T, Tanaka M (2012) Chimpanzees’ flexible targeted helping based on an understanding of
conspecifics’ goals. Proceedings of the National Academy of Sciences USA 109(9):3588-3592 (doi:
10.1073/pnas.1108517109).
ጊᧄ⌀਽ (2011) ೑ઁ࡮දജⴕേߩࡔࠞ࠾࠭ࡓߣ␠ળߩㅴൻ. 㔤㐳㘃⎇ⓥ 27(2):95-109.
ߘߩઁߩၫ╩
1)
2)
ᐔ↰⡡ (2011) ࠴ࡦࡄࡦࠫ࡯ߩࡔ࠲⹺⍮ታ㛎. p.128:96-104 ⊒㆐㧚
ጊᧄ⌀਽ (2011) ࠴ࡦࡄࡦࠫ࡯࡮ࡏࡁࡏߦߺࠆ‫ޟ‬ᓼ‫⿠ߩޠ‬Ḯ. p.7:20-23 ߎߎࠈߩᧂ᧪㧚
ቇળ⊒⴫
1)
2)
3)
4)
5)
6)
7)
⻠Ṷ
1)
2)
3)
4)
Yamamoto S (2011) Evolution of altruism, reciprocity, and cooperation: suggestions from chimpanzees and bonobos.
The 27th Annual Meeting of Japanese Society of Population Ecology. Symposium “The evolution of animal societies:
generality and specificity of the systems” (2011/10/16, Okayama)㧚
Yamamoto S (2011) Nature in Bhutan and its environmental education. Welcome Meeting for The Honorable
Chairperson of the National Council of Bhutan and His Delegation (2011/09/30, Kyoto).
ᐔ↰⡡ (2011) ࠴ࡦࡄࡦࠫ࡯ߩࡔ࠲⹺⍮㧦⥄りߩ⍮⼂⁁ᘒߩ⹺⍮ߦ㑐ߔࠆታ㛎⊛⎇ⓥ. ᣣᧄᔃℂቇળ╙ 75 ࿁ᄢ
ળ (2011/09/17, ᧲੩)㧚
ᐔ↰⡡ (2011) ᄢੱ࠴ࡦࡄࡦࠫ࡯ߪߤߩߊࠄ޿࠽࠶࠷ഀࠅࠍⷡ߃ࠄࠇࠆ߆. ╙ 14 ࿁ SAGA ࠪࡦࡐࠫ࠙ࡓ
(2011/11/12-13, ᾢᧄ)㧚
ጊᧄ⌀਽ (2011) ㊁↢ࡏࡁࡏߦ߅ߌࠆᨐታಽ㈩㨪㘩‛ಽ㈩ߩㅴൻ࡮ࡔࠞ࠾࠭ࡓߦ߆ࠎߔࠆౣᬌ⸛㨪. Animal
2011 (ᣣᧄേ‛ᔃℂቇળ㧔╙ 71 ࿁㧕࡮ᣣᧄേ‛ⴕേቇળ㧔╙ 30 ࿁㧕࡮ᔕ↪േ‛ⴕേቇળ/ᣣᧄኅ⇓▤ℂቇળ
(2011/09/11, ᧲੩)㧚
ጊᧄ⌀਽㧘᧻ᴛື㇢ (2011) ࠴ࡦࡄࡦࠫ࡯࡮ࡏࡁࡏߦ߅ߌࠆ㆏ᷰࠅᤨߩ㓸࿅දജⴕേ. ╙ 4 ࿁ᣣᧄੱ㑆ⴕേㅴ
ൻቇળ (2011/11/19, ᧅᏻ)㧚
Yamamoto S (2012) Mechanisms of cooperation in our evolutionary relatives. The 3rd meeting of International Institute
of Advanced Studies (2012/01/29, Kyoto).
ᐔ↰⡡ (2011/05/09) ࠴ࡦࡄࡦࠫ࡯ߩᔃߣⴕേ. ੩ጊ㐳ኼ↢߈߇޿࠮ࡒ࠽࡯. ጟጊ⋵ጟጊᏒ࡮੩ጊ౏᳃㙚. ⡬ⴐ
⚂ 30 ฬ㧚
ᐔ↰⡡ (2011/07/18) ࠴ࡦࡄࡦࠫ࡯ߩ⍮ᕈࠍតࠆ. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ ౏㐿ࠪࡦࡐࠫ࠙ࡓ‫ࡄࡦ࠴ߣੱޟ‬
ࡦࠫ࡯ߩ㑆‫ޠ‬. ᗲ⍮⋵›ጊᏒ࡮ࡈࡠࠗ࠺㧚⡬ⴐ⚂ 200 ฬ.
ጊᧄ⌀਽ (2011/10/15) දജ␠ળߩㅴൻ ࿖㓙㜞╬⎇ⓥᚲ⎇ⓥࡊࡠࠫࠚࠢ࠻‫ޟ‬ᔃߩ⿠Ḯ‫ޠ‬2011 ᐕᐲ╙ 2 ࿁⎇ⓥળ
੩ㇺᐭ⋧ᭉ㇭♖⪇↸࡮࿖㓙㜞╬⎇ⓥᚲ. ⡬ⴐ⚂ 30 ฬ㧚
ጊᧄ⌀਽ (2012/03/17) ⷐ᳞ߦᔕ߃ࠆ࠴ࡦࡄࡦࠫ࡯‫ߦ⊛⊒⥄ޔ‬ഥߌࠆࡅ࠻ 㨪೑ઁⴕേߩ߅ߌࠆઁ⠪ℂ⸃ߣߘߩ
ㅴൻ㨪. ർᶏ㆏ᄢቇࠣࡠ࡯ࡃ࡞ COE‫ޟ‬ᔃߩ␠ળᕈߦ㑐ߔࠆᢎ⢒⎇ⓥ᜚ὐ‫✚ޠ‬᜝ࠪࡦࡐࠫ࠙ࡓ ᔃߪ‫ޔޠߗߥޟ‬
‫ߤޟ‬
ߩࠃ߁ߦ‫␠ޠ‬ળ⊛߆㧫㨪ࡈࡠࡦ࠹ࠖࠕߣࠕࠫࠚࡦ࠳㨪. ᧲੩ㇺජઍ↰඙࡮ቇⴚ✚ว࠮ࡦ࠲࡯. ⡬ⴐ⚂ 200 ฬ㧚
㧙㧙
㐳ᦼ㊁ᄖ⎇ⓥࡊࡠࠫࠚࠢ࠻
㧨⎇ⓥ᭎ⷐ㧪
A) ᧲ධࠕࠫࠕᾲᏪᨋߩ㔤㐳㘃ߩ␠ળ↢ᘒቇ⊛⎇ⓥ
᧻↰৻Ꮧ㧘ඨ⼱๋㇢㧔↢ᘒ଻ో㧕㧘ᄢ⼱ᵗ੺㧔↢ᘒ଻ో㧕
2005 ᐕࠃࠅ‫ࡃࠨࠕࠪ࡯࡟ࡑޔ‬Ꮊߩࠬࠞ࠙᧛ࠍ᜚ὐߣߒߚ࠹ࡦࠣࠩ࡞ߩ㐳ᦼⷰኤࡊࡠࠫࠚࠢ࠻ࠍⴕߞߡ޿ࠆ‫ᧄޕ‬
ࡊࡠࠫࠚࠢ࠻ߢߪ‫␠ߩ࡞ࠩࠣࡦ࠹ޔ‬ળ↢ᘒ‫ޔ‬ណ㘩↢ᘒ‫ⴕޔ‬േ↢ᘒߩⷰὐ߆ࠄ⎇ⓥࠍㅴ߼ߡ޿ࠆ‫࡞ࠩࠣࡦ࠹ޔߚ߹ޕ‬
ߣหᚲ⊛ߦ↢ᕷߒߡ޿ࠆઁߩᤤⴕᕈ㔤㐳㘃㧔ࠝ࡜ࡦ࠙࡯࠲ࡦ‫࡯ࡃ࡞ࠪޔ࡞ࠩࠝ࠲ࡉޔ࡞ࠩࠗࠢ࠾ࠞޔ࡞ࠩࠟ࠽࠹ޔ‬
࡜ࡦࠣ࡯࡞㧕߿࿾਄ᕈູ੃㘃㧔ࡅࠥࠗࡁࠪࠪ‫ߤߥࠞࠫࡔࡑޔ࡯ࡃࡦࠨޔ‬㧕ߩၮ␆⊛ߥ↢ᘒ࡮␠ળߩ⎇ⓥ߽หᤨߦⴕ
ߞߡ޿ࠆ‫ޕ‬
㘩‛⾗Ḯ㊂ߩᄌേࠍ⺞ߴࠆ⋡⊛ߢ‫᧛࠙ࠞࠬޔ‬ㄭ㇠ߦ޽ࠆࠠ࠽ࡃ࠲ࡦࠟࡦᎹߩᡰᵹࡑ࠽ࡦࠣ࡞Ꮉߦ⸳⟎ߒߡ޿ࠆᬀ
↢⺞ᩏ඙ߢ‫ޔ‬Ფ᦬৻࿁ߩᬀ‛ࡈࠚࡁࡠࠫ࡯⺞ᩏࠍⴕߞߚ‫ࠍ࡞ࠩࠣࡦ࠹ޔߚ߹ޕ‬฽߻㔤㐳㘃 6 ⒳ߩ୘૕⟲േᘒࠍ᣿ࠄ
߆ߦߔࠆߚ߼ߦ‫ࠆࠃߦ࠻࡯ࡏޔ‬㔤㐳㘃࠮ࡦࠨࠬࠍᲤㅳ৻࿁ⴕߞߚ‫࠙ࠞࠬߚ߈ߡߞⴕߢ߹੹ޕ‬ㄭ㇠ߩᎹㄝᨋߣߪ⇣ߥ
ࠆᬀ↢ߢ޽ࠆ‫ࡉ࡯ࡠࠣࡦࡑޔ‬ᨋߦ↢ᕷߔࠆ࠹ࡦࠣࠩ࡞ߩ੍஻⺞ᩏ߽ⴕߞߚ‫ߩ࡞ࠩࠝ࠲ࡉߩ᧛࠙ࠞࠬޕ‬ၮ␆⊛ߥ↢ᘒ‫ޔ‬
␠ળࠍ᣿ࠄ߆ߦߔࠆ⋡⊛ߢ‫੍ߚ޿↪ࠍࠬࠨࡦ࠮࠻࡯ࡏޔ‬஻⺞ᩏࠍⴕߞߚ‫ޕ‬
B) ࠞ࡝ࡦ࠭᫪ᨋ଻⼔඙ߦ᫮ᕷߔࠆ㊁↢㔤㐳㘃ߩ⎇ⓥ
દᏀᴦ⟤ᄹ㧘ᯅᧄජ⛗㧔↢ᘒ଻ో㧕㧘ᳯፉବ㧔↢ᘒ଻ో㧕㧘ฎᏒ೰ผ㧔␠ળㅴൻ㧕㧘ጟᧄቬ⵨㧔ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ
࠮ࡦ࠲࡯㧕
࠙ࠟࡦ࠳౒๺࿖ࠞ࡝ࡦ࠭᫪ᨋ଻⼔඙ߦ↢ᕷߔࠆ㊁↢࠴ࡦࡄࡦࠫ࡯2 㓸࿅ࠍኻ⽎ߦ‫ޔ‬㓸࿅㑆ߩ಴ળ޿ߩ੤ᷤ‫␠ޔ‬ળ
ⴕേߩ㆑޿‫ޔ‬ណ㘩ⴕേߦߟ޿ߡߩ㐳ᦼ⊛࠺࡯࠲ࠍ෼㓸ߒߚ‫ޕ‬ᨐታ㊂ߦߟ޿ߡ߽᦬ 1 ࿁࠺࡯࠲ࠍߣߞߚ‫ੱޕ‬₞౒ㅢᗵ
ᨴ∝ߩ⎇ⓥࠍㅴ߼ࠆߚ߼ߦ‫⹜♮ޔ‬ᢱߦࠃࠆነ↢⯻ߩ⺞ᩏࠍⴕߞߚ߶߆‫ޔ‬ᗵᨴߩጁᱧࠍ⺞ߴࠆߚ߼ߩ♮࡮ዩ⹜ᢱࠍ෼
㓸ߒߚ‫ߩࡓ࠭࡝࡯࠷ࠦࠛޔߦࠄߐޕ‬ᓇ㗀ࠍ⺞ߴࠆߚ߼ߦ‫ⷰޔ‬శቴߦኻߔࠆ࠴ࡦࡄࡦࠫ࡯ߩⴕേߩ࠺࡯࠲ࠍ෼㓸ߒߚ‫ޕ‬
C) ㊁↢ࡏࡁࡏߩ㓸࿅᭴ㅧߩ↢ᚑߦߟ޿ߡߩ⎇ⓥ
ဈᏎື਽㧘ฎᏒ೰ผ㧔␠ળㅴൻ㧕
㊁↢ࡏࡁࡏߩᲫߠߊࠈ޿੤ᷤ‫ޔ‬㓸࿅ࡔࡦࡃ࡯ߩ಴ળ޿ߩ੤ᷤ‫ޔ‬⒖౉ࡔࠬߩ␠ળ㑐ଥߩᄌㆫߦ㑐ߔࠆ࠺࡯࠲෼㓸ߣ
ಽᨆ‫⚿ޔ‬ᨐߩ౏⴫ࠍㅴ߼ߚ‫ޕ‬㓸࿅㑆㑐ଥࠍ࠹࡯ࡑߦ‫ޔ‬ᣂߚߥ⺞ᩏ㓸࿅ߩㅊ〔ⷰኤ‫ޔߌߠੱޔ‬୘૕⼂೎ࠍ⛮⛯ߒߚ‫ޕ‬
ⴕേߩᄙ᭽ᕈࠍ࠹࡯ࡑߦ‫ޔ‬㓞ធ୘૕⟲ߩ⺞ᩏ߽⛮⛯ߒߚ‫ޕ‬
㧨⎇ⓥᬺ❣ 㧪
ේ⪺⺰ᢥ
1)
2)
3)
4)
5)
6)
Bernard H, Matsuda I, Hanya G, Ahmad AH (2011) Characteristics of Proboscis Monkey (Nasalis larvatus) Night
Sleeping-trees in Sabah, Malaysia. International Journal of Primatology 32:259-26.
Bernard H, Matsuda I, Hanya G, Ahmad AH (2011) Effects of river width on the selection of sleeping-site by proboscis
monkeys (Nasalis larvatus) in Sabah, Malaysia. Journal of Tropical Biology and Conservation 8:9-12.
Matsuda I, Tuuga A, Bernard H (2011) Riverine refuging by proboscis monkeys (Nasalis larvatus): implications for
adaptive benefits in riverine habitat. Mammalian Biology 76:165-171.
Matsuda I, Murai T, Clauss M, Yamada T, Tuuga A, Bernard H, Higashi S (2011) Regurgitation and remastication in the
foregut-fermenting proboscis monkey (Nasalis larvatus). Biology Letters 7:786-789.
Matsuda I, Tuuga A, Bernard H, Furuichi T (2012) Inter-individual relationships in proboscis monkeys: a preliminary
comparison with other non-human primates. Primates 53:13-23.
Sakamaki T (2011) Submissive pant-grunt greeting of female chimpanzees in Mahale Moutains National Park, Tanzania.
African Study Monographs 32:25-41.
✚⺑
1)
᧻↰৻Ꮧ (2011) ࠹ࡦࠣࠩ࡞߆ࠄ⚌⸃ߊࠦࡠࡉࠬ੝⑼ߩᄙ᭽ߥ↢ᘒߣ␠ળ. 㔤㐳㘃⎇ⓥ 27:75-93.
⪺ᦠ㧔ಽᜂၫ╩㧕
1)
2)
Furuichi T, Idani G, Ihobe H, Hashimoto C, Tashiro Y, Sakamaki T, Mulavwa BN, Yangozene K, Kuroda S (2011)
Long-term studies on wild bonobos at Wamba, Luo Scientific Reserve, D.R. Congo: towards the understanding of female
life history in a male-philopatric species. (Long-term field studies of primates) (ed. Kappeler P, Watts D) p.413-433,
Springer-Verlag, Berlin, Heidelberg.
Sha J, Matsuda I, Bernard H (2011) The Natural History of Proboscis Monkey. (Natural History Publications. Kota
Kinabalu) p128.
⪺ᦠ㧔න⪺㧕
1)
᧻↰৻Ꮧ (2012) ࠹ࡦࠣࠩ࡞̆ᴡߣ↢߈ࠆࠨ࡞. ᧲ᶏᄢቇ಴ ળ. p.146.
ቇળ⊒⴫
1)
᧻↰৻Ꮧ㧘Tuuga A㧘Bernard H㧘ฎᏒ೰ผ (2011) ࠹ࡦࠣࠩ࡞␠ળߩ․ᓽ㧦ࡔࠬߩ⒖☋᭽ᑼߣ୘૕㑆㑐ଥ. ╙ 27
㧙㧙
࿁ᣣᧄ㔤㐳㘃ቇળᄢળ (2011/07㧘›ጊ).
2) ᧻↰৻Ꮧ㧘᧛੗഼⵨㧘Clauss M㧘ጊ↰᦮⟤㧘Tuuga A㧘Bernard H㧘᧲ᱜ೰ (2011) 㔤㐳㘃ߩ෻⧖ⴕേߩ⊒⷗㧦࠹
ࡦࠣࠩ࡞ߩ੐଀. 2011 ᐕᐲᣣᧄູ੃㘃ቇળᄢળ (2011/09, ችፒ).
⻠Ṷ
1)
2)
3)
4)
Matsuda I (2011/05) Evolution of Primate Society. Lecture - Studying and Monitoring Primate Behavior: A Workshop for
Asian Conservation Professionals. ࠪࡦࠟࡐ࡯࡞.
᧻↰৻Ꮧ (2011/07) ࡈࠖ࡯࡞࠼ࡢ࡯ࠢߩน⢻ᕈ㧙࠹ࡦࠣࠩ࡞⎇ⓥߣ⑳㧙. ╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ 㜞ፉ
⾨ฃ⾨⻠Ṷ. ›ጊ.
᧻↰৻Ꮧ (2011/11) ࡜ࡏ߆ࠄࡈࠖ࡯࡞࠼߳㧦ࡏ࡞ࡀࠝߩࠫࡖࡦࠣ࡞ߢ࠹ࡦࠣࠩ࡞ࠍㅊ߁. หᔒ␠ᄢቇήᯏൻቇ
⎇ⓥቶ ୃ⚡ળ. ੩ㇺ.
Matsuda I (2011/12) Proboscis Moneky -Big Nose of Borneo. University of Zurich ਥ௅ Lecture series on "Indonesian
wildlife". ࠴ࡘ࡯࡝࠶ࡅ.
2. ੤ᵹදቯ
2012 ᐕ 3 ᦬ 31 ᣣ⃻࿷
ቇⴚ੤ᵹදቯ
දቯ࿖
දቯవ
දቯవ㧔ࠕ࡞ࡈࠔࡌ࠶࠻⴫⸥㧕
දቯᐕ᦬ᣣ
ᦼ㑆
ࠡ࠾ࠕ
ࠡ࠾ࠕ⑼ቇᛛⴚᐡ
La Direction Nationale de la Recherche Scientifique et
Technique
2004.1.28
5 ᐕ㑆
㧔⥄േ⛮⛯㧕
ࠡ࠾ࠕ
ࡏ࠶࠰࠙ⅣႺ⎇ⓥᚲ
L'Institut de Recherche Environnementale de Bossou
(IREB)
2004.1.4
5 ᐕ㑆
㧔⥄േ⛮⛯㧕
ࠬ࡝ࠫࡖࡗࡢ࡞࠺ࡀࡊ࡜ᄢቇ
␠ળቇੱ㘃ቇᢎቶ
࿖┙ዳ᧲⑼ᛛᄢቇ
㊁↢േ‛଻ోቇ⎇ⓥᚲ
University of Sri Jayawardenepura (Faculty of Arts,
Department of Sociology and Anthropology)
2005.8.18
10 ᐕ㑆
Institute of Wildlife Conservation National Pingtung
University of Science and Technology
2008.1.18
10 ᐕ㑆
࠲ࠗ₺࿖േ‛࿦දળ
The Zoological Park Organization, Thailand
2009.9.16
5 ᐕ㑆
(2009.10.1㨪)
࠰࠙࡞ᄢ౏࿦㧔࠰࠙࡞േ‛࿦㧕
Seoul Grand Park (Seoul Zoo)
2010.4.28
ࠬ࡝࡜ࡦࠞ
บḧ
࠲ࠗ
ᄢ㖧᳃࿖
࠲ࠗ
ࠦࡦࠧ᳃ਥ
౒๺࿖
ࠗࡦ࠼ࡀ
ࠪࠕ
ࡃࡦࠣ࡜
࠺ࠪࡘ
࠲ࠗ
࠴ࡘ࡜ࡠࡦࠦࡦᄢቇℂቇㇱ
Faculty of Science, Chulalongkorn University
2010.5.24
5 ᐕ㑆
㧔⥄േ⛮⛯㧕
↢ᘒ᫪ᨋ⎇ⓥᚲ
The Research Center for Ecology and Forestry,
Democratic Republic of Congo
2010.9.15
5 ᐕ㑆
ࠕࡦ࠳࡜ࠬᄢቇℂቇㇱ
Department of Bioloby, Faculty of Science, Andalas
University
2011.4.27
㊁↢↢‛଻⼔ዪ
The Wildlife Trust of Bangladesh, Bangladesh
2011.3.12
ࠞ࠮ࠨ࡯࠻ᄢቇℂቇㇱ
Faculty of Science, Kasetsart University
2011.4.25
5 ᐕ㑆
(2011.4.1㨪)
5 ᐕ㑆
(2011.4.1㨪)
5 ᐕ㑆
(2011.5.1㨪)
౒ห⎇ⓥදቯ
ᣣᧄ
ᣣᧄ
⽷࿅ᴺੱฬฎደߺߥߣᝄ⥝⽷࿅
㧔ฬฎደ᷼᳓ᣖ㙚㧕
⽷࿅ᴺੱ㜞⍮⋵
ߩ޿ߜേ‛౏࿦දળ
2009.7.3
2010.8.12
2013.3.31
ࡑ࡟࡯ࠪࠕ
ࠝ࡜ࡦ࠙࡯࠲ࡦፉ⽷࿅
Orang Utan Foundation
2010.11.1
ࡑ࡟࡯ࠪࠕ
ࡊ࡜࠙ࡃࡦ࠺ࠖࡦࠣ⽷࿅
Pulau Banding Foundation
2010.11.1
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅‫࡚ࠪ࠽࡮ࡓࠕ࡝ࡠ࠻ࡍޔ‬
࠽࡞ᩣᑼળ␠㧔ࡍ࠻ࡠ࠽ࠬ㧕‫ࠨޔ‬
ࡃᎺ᫪ᨋዪ‫ࡃࠨޔ‬Ꮊ㊁↢↢‛ዪ‫ޔ‬
ࡑ࡟࡯ࠪࠕ⑼ቇࠕࠞ࠺ࡒ࡯
Yayasan Sabah, Petroliam Nasional Berhad(Petronas),
Sabah Forestry Departmant, Sabah Wildlife Department,
Academy of Sciences, Malaysia
2011.6.29
3. ቇ૏ขᓧ⠪ߣ⺰ᢥ㗴⋡
੩ㇺᄢቇඳ჻㧔ℂቇ㧕
⑔ፉ⟤๺㧔⺖⒟㧕㧦Reading and Writing Support for Children with Developmental Disorders: The Possibility of Using
e-Learning㧔⊒㆐㓚ኂఽߩ⺒ߺᦠ߈ቇ⠌ᡰេ㧦e-Learning ࠍขࠅ౉ࠇߚᡰេߩน⢻ᕈ㧕
㧙㧙
ዊୖආବ㧔⺖⒟㧕㧦Cognitive foundations of environmental enrichment for Japanese macaques㧔࠾ࡎࡦࠩ࡞ߩⅣႺࠛࡦ࡝
࠶࠴ࡔࡦ࠻ߦ߅ߌࠆ⹺⍮⊛ၮ⋚㧕
દ ⮮ Პ 㧔 ⺖ ⒟ 㧕㧦 External and internal craniofacial morphology of Asian macaques and its evolutionary and
paleobiogeographic implications㧔ࠕࠫࠕ↥ࡑࠞࠢዻ㔤㐳㘃ߦ߅ߌࠆ㗡㛽㗻㕙ㇱߩᄖㇱߣౝㇱߩᒻᘒ߅ࠃ߮ߘߩㅴൻ
⊛࡮ฎ↢‛࿾ℂቇ⊛ᗧ⟵㧕
౗ሶፄ᣿㧔⺖⒟㧕㧦The perception of one’s own actions in chimpanzees and humans㧔࠴ࡦࡄࡦࠫ࡯ߣࡅ࠻ߦ߅ߌࠆ⥄Ꮖ
ߩㆇേߩ⹺⼂㧕
⁚㊁ᢥᶈ㧔⺖⒟㧕
㧦A comparative eye-tracking study in great apes and humans: the pattern of eye movements for scenes and
faces㧔㘃ੱ₎ߣࡅ࠻ߦ߅ߌࠆᲧセࠕࠗ࡮࠻࡜࠶ࠠࡦࠣ⎇ⓥ㧦㘑᥊ߣ㗻ߦኻߔࠆ⌒⃿ㆇേߩࡄ࠲࡯ࡦ㧕
ᯫ၂ዊ⊖ว㧔⺖⒟㧕
㧦Molecular aspects of brain aging in female macaques㧔ࡔࠬࡑࠞࠢࠩ࡞ߦ߅ߌࠆ⣖ߩ⠧ൻߩಽሶ⊛
஥㕙㧕
Christopher Flynn Martin㧔⺖⒟㧕
㧦Chimpanzee social intelligence: information use and strategic interaction in an automated
testing environment㧔࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ␠ળ⊛⍮ᕈ㧦ో⥄േ࠹ࠬ࠻႐㕙ߦ߅ߌࠆᖱႎ೑↪ߣᚢⴚ⊛␠ળ੤ᷤ㧕
੩ㇺᄢቇୃ჻㧔ℂቇ㧕
⏷᧛᦮ሶ㧦⥄㐽∝ఽߦ߅ߌࠆᖱേ⹺⍮ߦ㑐ࠊࠆⷞⷡត⚝⎇ⓥ
૒⮮᧙ᄹ㧦㔤㐳㘃ߩ੃ఽ↹௝ߦኻߔࠆⷞⷡ⊛ᵈᗧ࡮ᑯ೎࡮ㆬᅢᕈߦ㑐ߔࠆታ㛎⊛⎇ⓥ
ᣧᎹථᔒ㧦࠴ࡦࡄࡦࠫ࡯੝⒳㑆ߦ߅ߌࠆ⧰๧ฃኈ૕ㆮવሶߩ࿾ၞಽൻ
᧛᧻᣿Ⓞ㧦࠴ࡦࡄࡦࠫ࡯ߣࡅ࠻ߦ߅ߌࠆ૞ᬺ⸥ᙘߩᲧセ
࡙ ࡝࡜㧦࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆห⺞ⴕേߩታ㛎⊛⎇ⓥ
4. ᄖ࿖ੱ⎇ⓥຬ
᜗߳޿ᄖ࿖ੱቇ⠪࡮ᄖ࿖ੱ౒ห⎇ⓥ⠪
ᒛ 㢈㧔ਛ⪇ੱ᳃౒๺࿖‫ޔ‬ਛጊᄢቇ␠ળቇਈੱ㘃ቇቇ㒮࡮⻠Ꮷ㧕(2010.8.1㨪2011.7.31) ฃ౉ᢎຬ㧦ᷰㆻ㇌ᄦ
⎇ⓥ㗴⋡㧦࠾ࡎࡦࠩ࡞␠ળ᭴ㅧߩᲧセ⎇ⓥ
H Bouchet㧔ࡈ࡜ࡦࠬ౒๺࿖‫৻╙࠿ࡦ࡟ޔ‬ᄢቇ࡮࡝ࠨ࡯࠴ࠕࠪࠬ࠲ࡦ࠻㧕(2010.9.7㨪2011.9.6) ฃ౉ᢎຬ㧦ᱜ㜞ା↵
⎇ⓥ㗴⋡㧦࠾ࡎࡦࠩ࡞ߩ⊒ᖱჿߣᕈᚢ⇛ߦ㑐ߔࠆታ㛎⊛⎇ⓥ
CD Dahl㧔࠼ࠗ࠷ㅪ㇌౒๺࿖‫⎇ࠬࠢࠖ࠹ࡀࡃࠗࠨ‛↢ࠢࡦ࡜ࡊࠬࠢ࠶ࡑޔ‬ⓥᚲ࡮⎇ⓥຬ㧕(2010.10.1㨪2012.9.30) ฃ౉
ᢎຬ㧦෹᳗㓷Ꮖ
⎇ⓥ㗴⋡㧦㔤㐳㘃ߦ߅ߌࠆ㗻⹺⍮߅ࠃ߮␠ળ⊛⹺⍮ߦ㑐ߔࠆᲧセ⹺⍮⑼ቇ⊛⎇ⓥ
C Sueur㧔ࡈ࡜ࡦࠬ౒๺࿖‫ࡦ࠻ࠬࡦ࡝ࡊޔ‬ᄢቇ࡮⎇ⓥຬ㧕(2010.10.13㨪2011.8.3) ฃ౉ᢎຬ㧦ᷰㆻ㇌ᄦ
⎇ⓥ㗴⋡㧦࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ㓸࿅ߣߒߡߩᗧᔒ᳿ቯᯏ᭴
ME Kret㧔ࠝ࡜ࡦ࠳₺࿖‫ࠣ࡯ࡃ࡞ࠖ࠹ޔ‬ᄢቇ࡮PhD Student㧕(2011.2.4㨪2011.7.29) ฃ౉ᢎຬ㧦᧻ᴛື㇢
⎇ⓥ㗴⋡㧦ᖱേ⴫಴ߣ⹺⍮ߩᄙ᭽ᕈߦ㑐ߔࠆᲧセ⹺⍮⑼ቇ⊛⎇ⓥ
PA Pebsworth㧔ࠕࡔ࡝ࠞวⴐ࿖‫ޔ‬Wildcliff Nature Reserve࡮Site Biologist㧕(2011.2.6㨪2011.12.16) ฃ౉ᢎຬ㧦MA Huffman
⎇ⓥ㗴⋡㧦࠴ࡖࠢࡑࡅࡅߩ⥄Ꮖᴦ≮ⴕേߣነ↢⯻ᗵᨴߩ↢ᘒቇߦߟ޿ߡ
5. ᣣᧄੱ⎇ⓥຬ࡮⎇ୃຬ
ᣣᧄቇⴚᝄ⥝ળ․೎⎇ⓥຬ㧔㧼㧰㧕
ᧁ႐␞ሶ㧔2010.4.1㨪2012.3.31㧕 ฃ౉ᢎຬ㧦ਛ᧛స᮸
⎇ⓥ㗴⋡㧦㔤㐳㘃ߦ߅ߌࠆᕈߩ⹺⍮ߣᕈࡎ࡞ࡕࡦߩᓇ㗀
᧻↰৻Ꮧ㧔2011.4.1㨪2011.10.31㧕 ฃ౉ᢎຬ㧦ඨ⼱๋㇢
⎇ⓥ㗴⋡㧦㔤㐳㘃␠ળߩ㊀ጀ᭴ㅧߩ⸃᣿㧦࠹ࡦࠣࠩ࡞ߩ⒳ౝᄌ⇣
ᩊፒోᒄ㧔2011.4.1㨪2014.3.31㧕 ฃ౉ᢎຬ㧦ᱜ㜞ା↵
㧙㧙
⎇ⓥ㗴⋡㧦ࡅ࠻ߩਇ቟㓚ኂߩࡔࠞ࠾࠭ࡓ⸃᣿ߦะߌߚേ‛ࡕ࠺࡞ߩ᭴▽
㕖Ᏹൕ⎇ⓥຬ
᧻੗ ᷕ㧔2011.4.1㨪2012.3.31㧕ੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯
⎇ⓥ㗴⋡㧦࠾ࡎࡦࠩ࡞ⴊዊ᧼ᷫዋ∝ߩ⊒∝ᯏᐨߩ⸃᣿ߣㆮવሶ⸻ᢿᴺߩ㐿⊒
᦯ㇱ⵨ሶ㧔2011.4.1㨪2012.3.31㧕ᕁ⠨⸒⺆ಽ㊁
⎇ⓥ㗴⋡㧦␠ળ⊛⹺⍮ߩၮ⋚ߣߒߡߩห⺞ⴕേߦ㑐ߔࠆᲧセ⹺⍮⎇ⓥ
᧻ේ ᐙ㧔2011.4.1㨪2012.3.31㧕␠ળㅴൻಽ㊁
⎇ⓥ㗴⋡㧦࠾ࠪࡠ࡯࡜ࡦ࠼ࠧ࡝࡜ߩⴕേ⊒㆐ߣ⟲ࠇ᭴ㅧߩ㑐ㅪ
6. ⎇ⓥ㓸ળ
ᚲౝ⺣⹤ળ
╙ 1 ࿁㧦2011 ᐕ 12 ᦬ 8 ᣣ(ᧁ)
Michael P. Muehlenbein (Evolutionary Physiology and Ecology Laboratory, Department of Anthropology, Indiana University)
‫ޟ‬Balancing the costs and benefits of primate-based tourism‫ޠ‬
╙ 2 ࿁㧦2012 ᐕ 3 ᦬ 8 ᣣ(ᧁ)
Rachel Wu (Centre for Brain and Cognitive Development, University of London)
‫ޟ‬Learning (to learn) from attention cues during infancy‫ޠ‬
╙ 3 ࿁㧦2012 ᐕ 3 ᦬ 9 ᣣ(㊄)
Yin Lijie (Center for Nature and Society, School of Life Sciences, Peking University)
‫ޟ‬Research on white-headed leaf monkeys in Guangxi, China‫ޠ‬
╙ 4 ࿁㧦2012 ᐕ 3 ᦬ 13 ᣣ(Ἣ)
Badrul Munir Md Zain (School of Environmental and Natural Resource Sciences, Faculty of Science and Technology,
Universiti Kebangsaan)
‫ޟ‬Phylogenetic position of Bornean banded langur and nuisance problems caused by long-tailed macaques in Malaysia‫ޠ‬
㧔ᢥ⽿㧦᧻ᧄᱜᐘ㧕
7. 2011 ᐕ 㔤㐳㘃ቇ✚ว࠯ࡒ࠽࡯࡞
The Interdisciplinary Seminar on Primatology 2011
ᣣᤨ㧦2011 ᐕ 12 ᦬ 9 ᣣ(㊄)
ળ႐㧦੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲᧄ᫟ᄢળ⼏ቶ
⊒⴫㧦31 ઙ㧔ญ㗡 19 ઙ‫࡯࠲ࠬࡐޔ‬12 ઙ㧕
㔤㐳㘃ቇ✚ว࠯ࡒ࠽࡯࡞ߪ‫ޔ‬ᚲౝ࠹ࠖ࡯࠴ࡦࠣ࡮ࠕࠪࠬ࠲ࡦ࠻ࠍਛᔃߣߒߚᄢቇ㒮↢߇ડ↹ߒ‫ޔ‬ቇ↢߿ᚲౝ⎇ⓥ
ຬߩ⎇ⓥ੤ᵹࠍଦㅴߔࠆߎߣࠍ⋡⊛ߣߔࠆ‫ߪ࡞࡯࠽ࡒ࠯ᧄޔߚ߹ޕ‬㔤㐳㘃ቇ♽ߩᱜᑼߥࠞ࡝ࠠࡘ࡜ࡓߩ৻ߟߣߒߡ
⹺ቯߐࠇߡ޿ࠆ‫ᧄޕ‬ᐕᐲߪ‫߁޿ߣ߁ⴕߢ⺆⧷ߡߴߔࠍ⺰⼏߮ࠃ߅⴫⊒ޔ‬ᣂߚߥ⹜ߺߦ᜸ᚢߒߚ‫ޕ‬ቇ↢߅ࠃ߮⧯ᚻ⎇
ⓥ⠪ߦࠃࠆ⎇ⓥ⊒⴫ߦട߃‫ୃޔ‬჻ 1 ᐕ↢ߩ⎇ⓥ㧔⸘↹㧕⊒⴫߿਎⇇ฦ࿾ߩࡈࠖ࡯࡞࠼⚫੺ߩᨒࠍ⸳ߌࠆߥߤ‫ޔ‬ౝኈ
ߦ߽Ꮏᄦࠍขࠅ౉ࠇߚ‫ޕ‬ᒰᣣߪ‫ࠅ⋓߽⺰⼏ޔ‬਄߇ࠅ‫⴫⊒ޔ‬⠪࡮ෳട⠪ߦߣߞߡᄢᄌ᦭ᗧ⟵ߥ⎇ⓥળߣߥߞߚ‫ޕ‬
㧨ญ㗡⊒⴫࡮⥄↱Ṷ㗴㧪
1)
ዊᎹ ⹞ਫ㧔⹺⍮ቇ⠌ಽ㊁࡮ᄢቇ㒮↢㧕
Learning support for children with developmental disorder
⊒㆐㓚ኂఽ߳ߩቇ⠌ᡰេታ〣ߩ⚫੺
2)
Y Kim㧔ᕁ⠨⸒⺆ಽ㊁࡮ᄢቇ㒮↢㧕
Changes in estradiol levels and sexual behavior in captive Sichuan snub-nosed monkeys (Rhinopithecus roxellana)
following a male replacement
㓶੤ઍߦ઻߁㘺⢒ࠠࡦࠪࠦ࠙㧔Rhinopithecus roxellana㧕ߩࠛࠬ࠻࡜ࠫࠝ࡯࡞࡟ࡌ࡞ߩᄌൻߣᕈⴕേ
3)
P Pomchote㧔ㅴൻᒻᘒಽ㊁࡮⎇ⓥ↢㧕
Genetic variation and bone morphology of newts genus Tylototriton in Thailand
࠲ࠗߦ߅ߌࠆ Tylototriton ዻࠗࡕ࡝ߩㆮવ⊛ᄌ⇣ߣ㛽ᒻᘒ
㧙㧙
4)
RS Cicalise Takeshita㧔␠ળㅴൻಽ㊁࡮ᄢቇ㒮↢㧕
Hematological, hepatic, and renal evaluation in Aotus azarai infulatus
Aotus azarai infulatus ߦ߅ߌࠆⴊᶧ࡮⢄ᯏ⢻࡮⣢ᯏ⢻ߩ⹏ଔ
5)
᧛᧻ ᣿Ⓞ㧔ᕁ⠨⸒⺆ಽ㊁࡮ᄢቇ㒮↢㧕
Ongoing experiment in SPR
SPR ߢᦨㄭⴕߞߡ޿ࠆታ㛎
6)
AJJ MacIntosh㧔࿖㓙౒หవ┵⎇ⓥ࠮ࡦ࠲࡯࡮⎇ⓥຬ㧕
Bio-Logging for fractal patterns in penguin behavior: diving into the world of bio-complexity
ࡍࡦࠡࡦⴕേߩࡈ࡜ࠢ࠲࡞ࡄ࠲࡯ࡦߩࡃࠗࠝࡠࠡࡦࠣ㧦↢‛ⶄ㔀ᕈߩ਎⇇߳ߩ࠳ࠗࡆࡦࠣ
㧨ญ㗡⊒⴫࡮ࡢ࡯࡚ࠢࠪ࠶ࡊ㧦ୃ჻ 1 ᐕ↢ߦࠃࠆ⎇ⓥ㧔⸘↹㧕⊒⴫㧪
1)
ጊ↰ ᥓሶ㧔⹺⍮ቇ⠌ಽ㊁࡮ᄢቇ㒮↢㧕
Research planning: Bilingual children have difficulty in Hearing in noise test
⎇ⓥ⸘↹㧦ࡃࠗ࡝ࡦࠟ࡞ఽ┬ߩ㖸ჿ⍮ⷡߦ㑐ߔࠆ⎇ⓥ
2)
ᅏ↰ ᵏᒄ㧔⛔ว⣖ࠪࠬ࠹ࡓಽ㊁࡮ᄢቇ㒮↢㧕
Development of viral vector systems for pathway-specific gene regulation in the primate nervous system
㔤㐳㘃␹⚻♽ߦ߅ߌࠆ⚻〝ㆬᛯ⊛ㆮવሶ⊒⃻೙ᓮ࠙ࠗ࡞ࠬࡌࠢ࠲࡯ࠪࠬ࠹ࡓߩ㐿⊒
3)
ᬀ↰ ᗐ㧔ᕁ⠨⸒⺆ಽ㊁࡮ᄢቇ㒮↢㧕
Proposal for research theme
ୃ჻⎇ⓥ⸘↹
4)
↰ਛ ⟤ㇺ㧔⹺⍮ቇ⠌ಽ㊁࡮ᄢቇ㒮↢㧕
Do autistic children have altered gaze patterns in an imitation task including single-person (or two-person)? : An
eye-tracking analysis
ੑ⠪㑆㧔߽ߒߊߪਃ⠪㑆㧕ᮨ୮⺖㗴ߦ߅ߌࠆ⥄㐽∝ఽߩⷞ✢ࡄ࠲࡯ࡦߩಽᨆ㧦ࠕࠗ࠻࡜࠶ࠠࡦࠣࠍ↪޿ߚታ㛎
5)
ේ ᥰ㧔ࠥࡁࡓᄙ᭽ᕈಽ㊁࡮ᄢቇ㒮↢㧕
Survey of active DNA transposons for Hoolock hoolock and Hylobates lar
ࡈ࡯ࡠ࠶ࠢ࠹࠽ࠟࠩ࡞‫ߩߢ࡞ࠩࠟ࠽࠹࠹ࡠࠪޔ‬ᵴᕈဳ DNA ࠻࡜ࡦࠬࡐ࠱ࡦߩត⚝
6)
ᰞᐸ 㓁ሶ㧔ᕁ⠨⸒⺆ಽ㊁࡮ᄢቇ㒮↢㧕
PC tasks as a cognitive enrichment and a rehabilitation for the handicapped chimpanzee
㓚ኂࠍᜬߟ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆࡄ࠰ࠦࡦ⺖㗴ࠍ↪޿ߚ⹺⍮ࠛࡦ࡝࠶࠴ࡔࡦ࠻ߣ࡝ࡂࡆ࡝߳ߩᔕ↪
㧨ญ㗡⊒⴫࡮ࠪࡦࡐࠫ࠙ࡓ㧦㔤㐳㘃ߩࡈࠖ࡯࡞࠼⚫੺㧪
1)
ᣧᎹ ථᔒ㧔ㆮવሶᖱႎಽ㊁࡮ᄢቇ㒮↢㧕
Eastern chimpanzee (Pan troglodytes schweinfurthii) in the Mahale Mountains National Park, Tanzania
࠲ࡦࠩ࠾ࠕ࡮ࡑࡂ࡟ጊ႙࿖┙౏࿦ߩࡅࠟࠪ࠴ࡦࡄࡦࠫ࡯
2)
ጊ᪸ ⵨⟤㧔ᕁ⠨⸒⺆ಽ㊁࡮ᄢቇ㒮↢㧕
Western chimpanzee (Pan troglodytes verus) in Bossou, Guinea
ࠡ࠾ࠕ࡮ࡏ࠶࠰࠙ߩ࠾ࠪ࠴ࡦࡄࡦࠫ࡯
3)
દ⮮ Პ㧔♽⛔⊒↢ಽ㊁࡮ᄢቇ㒮↢㧕
Mammalian fossils from the Late Neogene sediments (e.g.,Trachypithecus sp.) in Chaingzauk, Myanmar and mammalian
fossils from the Plio-Pleistocene cave deposits (e.g.,Macaca sp.) in Beijing University Chongzuo Biodiversity Research
Institute, China
ࡒࡖࡦࡑ࡯࡮࠴ࡖࠗࡦࠩ࠙ࠢߩᣂ╙ਃ♿࡮ൻ⍹ູ੃㘃෸߮ਛ࿖࡮ፏᏀߩ㞲ᣂ਎-ᦝᣂ਎࡮ൻ⍹ູ੃㘃
4)
㋈ᧁ ධ⟤㧔ㆮવሶᖱႎಽ㊁࡮ᄢቇ㒮↢㧕
White-headed langur (Trachypithecus leucocephalus) in Beijing University Chongzuo Biodiversity Research Institute,
China
ਛ࿖࡮ፏᏀߩࡂࠢ࠻࡛࠙࠙ࠦ࠙
5)
ᳯፉ ବ㧔↢ᘒ଻ోಽ㊁࡮ᄢቇ㒮↢㧕
Primates in Kalinzu Forest Reserve, Uganda
࠙ࠟࡦ࠳࡮ࠞ࡝ࡦ࠭᫪ᨋߩ㔤㐳㘃
6)
ᓼጊ ᄹᏔሶ㧔␠ળㅴൻಽ㊁࡮ᄢቇ㒮↢㧕
Bonobo (Pan paniscus) in Wamba, Luo Scientific Reserve, Democratic Republic of the Congo
ࠦࡦࠧ᳃ਥ౒๺࿖࡮࡞ࠝ⑼ቇ଻⼔඙࡮ࡢࡦࡃߩࡏࡁࡏ
㧙㧙
7)
ᄢ⼱ ᵗ੺㧔↢ᘒ଻ోಽ㊁࡮ᄢቇ㒮↢㧕
Yakushima macaque (Macaca fuscata yakui) in Yakushima, Japan and pig-tailed macaque (Macaca nemestrina) in
Sukau, Sabah, Malaysia
ᣣᧄ࡮ደਭፉߩ࠾ࡎࡦࠩ࡞෸߮ࡑ࡟࡯ࠪࠕ࡮ࠨࡃᎺ࡮ࠬࠞ࠙ߩࡉ࠲ࠝࠩ࡞
㧨ࡐࠬ࠲࡯⊒⴫㧪
1)
⏷᧛ ᦮ሶ㧔⹺⍮ቇ⠌ಽ㊁࡮ᄢቇ㒮↢㧕
Visual Search in children with ASD
⥄㐽∝ఽߦ߅ߌࠆⷞⷡត⚝⎇ⓥ
2)
ᄢ⼱ ᵗ੺㧔↢ᘒ଻ోಽ㊁࡮ᄢቇ㒮↢㧕
Seasonal variation of males’ temporal isolation behavior of Japanese macaque
ࡗࠢࠪࡑࠩ࡞ߩࠝࠬߪቄ▵ߦࠃߞߡ⇣ߥࠆ⋡⊛ߢ㓸࿅ࠍ㔌ࠇࠆ
3)
౗ሶ ፄ᣿㧔ᕁ⠨⸒⺆ಽ㊁࡮ᄢቇ㒮↢㧕
Loss of visual field caused by an arachnoid cyst in a chimpanzee
࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆྙ⢩ߦࠃࠆⷞ㊁ᰳ៊
4)
㡨ᳰ ⩿଻㧔㜞ᰴ⣖ᯏ⢻ಽ㊁࡮ᄢቇ㒮↢㧕
Working memory of rhythm retained in fronto-parietal motor network
࡝࠭ࡓ૞ᬺ⸥ᙘߦ߅ߌࠆ೨㗡㗡㗂ㆇേࡀ࠶࠻ࡢ࡯ࠢߩᓎഀ
5)
ᧁ႐ ␞ሶ㧔㜞ᰴ⣖ᯏ⢻ಽ㊁࡮⎇ⓥຬ㧕
Characteristics of serial order learning in common marmosets (Callithrix jacchus)
ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ߦࠃࠆ㗅ᐨቇ⠌
6)
ᮭ↰ ᓀ㧔ጘ㒂ᄢቇᄢቇ㒮࡮ᄢቇ㒮↢㧕
Quantitative analysis of taste signal transduction molecules in the gastrointestinal tract of common marmosets
ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ߦ߅ߌࠆᶖൻ▤ߢߩ๧ⷡᖱႎવ㆐‛⾰ߩቯ㊂⊛⸃ᨆ
7)
㋈ᧁ ධ⟤㧔ㆮવሶᖱႎಽ㊁࡮ᄢቇ㒮↢㧕
Region specific dysfunction of bitter taste receptor TAS2R38 in Japanese macaques
࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ࿾ၞ․⇣⊛ߥ⧰๧ᗵฃᕈᄌ⇣
8)
ᓼጊ ᄹᏔሶ㧔␠ળㅴൻಽ㊁࡮ᄢቇ㒮↢㧕
Why do Japanese macaques perform redirected aggression?
࠾ࡎࡦࠩ࡞ߪߥߗੑᰴ᡹᠄ࠍⴕ߁ߩ߆
9)
ੑች ᄥᐔ㧔⛔ว⣖ࠪࠬ࠹ࡓಽ㊁࡮⎇ⓥຬ㧕
Segregated pathways carrying top-down signals from frontal cortex to visual areas MT and V4 in macaques
ࡑࠞࠢࠩ࡞ߦ߅ߌࠆ೨㗡⪲߆ࠄ㜞ᰴⷞⷡ㊁ MT ߅ࠃ߮ V4 ߳ߩᄙࠪ࠽ࡊࠬᕈ౉ജ᭽ᑼ
10) ⑧භ 㓷ผ㧔㜞ᰴ⣖ᯏ⢻ಽ㊁࡮ᄢቇ㒮↢㧕
Differential activities of monkey lateral prefrontal neurons between decisions based on short-term and long-term
memories
ࠨ࡞⢛஥೨㗡೨㊁␹⚻⚦⢩ߦ߅ߌࠆ⍴ᦼ⸥ᙘ࡮㐳ᦼ⸥ᙘߦၮߠߊᗧᕁ᳿ቯᤨߩᓎഀ
11) ᣧᎹ ථᔒ㧔ㆮવሶᖱႎಽ㊁࡮ᄢቇ㒮↢㧕
To investigate the food taste of wild chimpanzees (Pan troglodytes): tasting, chemistry, and genetics
㊁↢࠴ࡦࡄࡦࠫ࡯ߩ㘩ߴ‛ߩ๧ࠍ⺞ߴࠆ㧦⹜㘩‫ޔ‬ൻቇಽᨆ‫ޔ‬ㆮવሶ⸃ᨆ
12) ጊ᪸ ⵨⟤㧔ᕁ⠨⸒⺆ಽ㊁࡮ᄢቇ㒮↢㧕
Cortisol analysis from hair samples of captive chimpanzees (Pan troglodytes)
㘺⢒࠴ࡦࡄࡦࠫ࡯㧔Pan troglodytes㧕ߩᲫਛࠦ࡞࠴࠱࡞᷹ቯ
㧙㧙
㧔ᢥ⽿㧦ᣧᎹථᔒ㧕
Φ. ᢎ⢒ᵴേ
ࠣࡠ࡯ࡃ࡞ COE ߣߒߡߩᵴേ
⧷⺆࠻࡟࡯࠾ࡦࠣࠦ࡯ࠬߩ㐿௅
Friendly Scientific Debate Training Course㧔FSDTC1㧕
ࠣࡠ࡯ࡃ࡞ COE ߪ‫ޔ‬㒮↢ߩ࿖㓙ൻࡊࡠࠫࠚࠢ࠻ࠍផㅴߒߡ޿ࠆ‫ߣࡦ࡚ࠪ࡯࠹ࡦ࠯࡟ࡊࠆࠃߦ⺆⧷޿઻ߦࠇߎޕ‬
࠺ࠖࡌ࡯࠻߇ߢ߈ࠆࠃ߁ߦߔࠆߚ߼ߩ࠻࡟࡯࠾ࡦࠣ࡮ࠦ࡯ࠬ Friendly Scientific Debate Training Course (FSDTC1)߇‫ޔ‬
߅߅ࠃߘ᦬ 1 ᐲߩࡍ࡯ࠬߢ㐿௅ߐࠇ‫ޔ‬㔤㐳㘃⎇ⓥᚲ߆ࠄ߽Ფ࿁⊒⴫⠪ 1 ฬ‫⺰⸛ޔ‬⠪ 1 ฬ߇ෳടߒߚ‫ޕ‬
╙ 42 ࿁
5 ᦬ 12 ᣣ㧔ᧁ㧕11:30㨪17:00 ℂቇ⎇ⓥ⑼ 1 ภ㙚 214 ภቶ‫⴫⊒ޔ‬⠪㧦⷏ጟ ૓৻㇢ ⸛⺰⠪㧦ጊ᪸ ⵨⟤
Extinction of Japanese voles in Shikoku Island, western Japan
╙ 43 ࿁
6 ᦬ 9 ᣣ㧔ᧁ㧕11:30㨪17:00 ℂቇ⎇ⓥ⑼ 1 ภ㙚 214 ภቶ‫⴫⊒ޔ‬⠪㧦㡨ᳰ ⩿଻‫⺰⸛ޔ‬⠪㧦㋈ᧁ ධ⟤
Different contributions of frontal, parietal, and temporal cortices in working memory of rhythm
╙ 44 ࿁
7 ᦬ 7 ᣣ㧔ᧁ㧕11:30㨪17:00 ℂቇ⎇ⓥ⑼ 1 ภ㙚 104 ภቶ‫⴫⊒ޔ‬⠪㧦ศ↰ᒎ↢㧔㊁↢⎇࠮ࡦ࠲࡯㧕‫⺰⸛ޔ‬⠪㧦ᬀ↰ᗐ
Acoustic behavior of Commerson’s dolphins (ephalorhnchus commersonii) in Fitz Roy strait, southern Chile: Comparison
between wild and captive dolphins
╙ 45 ࿁
8 ᦬ 18 ᣣ㧔ᧁ㧕11:30㨪17:00 ℂቇ⎇ⓥ⑼ 1 ภ㙚 214 ภቶ‫⴫⊒ޔ‬⠪㧦౗ሶ ፄ᣿‫⺰⸛ޔ‬⠪㧦દ⮮ Პ
Relative contributions of kinematic information and goal representations for perception of self-agency in chimpanzees and
humans
╙ 46 ࿁
9 ᦬ 8 ᣣ㧔ᧁ㧕13:30㨪16:00 ℂቇ⎇ⓥ⑼ 1 ภ㙚 214 ภቶ‫⴫⊒ޔ‬⠪㧦㋈ᧁ ධ⟤‫⺰⸛ޔ‬⠪㧦ዊᎹ ⹞ਫ
Identification of non-taster Japanese macaques for a specific bitter taste
╙ 47 ࿁
10 ᦬ 13 ᣣ㧔ᧁ㧕11:30㨪17:00 ℂቇ⎇ⓥ⑼ 1 ภ㙚 104 ภቶ‫⴫⊒ޔ‬⠪㧦ᄢ⼱ ᵗ੺‫⺰⸛ޔ‬⠪㧦ศ↰ᒎ↢㧔㊁↢⎇࠮ࡦ
࠲࡯㧕
Temporal isolation of male Japanse macaques in Yakushima
╙ 48 ࿁
11 ᦬ 10 ᣣ㧔ᧁ㧕11:30㨪17:00 ℂቇ⎇ⓥ⑼ 1 ภ㙚 104 ภቶ‫⴫⊒ޔ‬⠪㧦቟੗ᣧ♿㧔㊁↢⎇࠮ࡦ࠲࡯㧕
‫⺰⸛ޔ‬⠪㧦ߥߒ
Personality assessment and factores influencing personality in captive elephants
╙ 49 ࿁
12 ᦬ 8 ᣣ㧔ᧁ㧕11:30㨪17:00 ℂቇ⎇ⓥ⑼ 1 ภ㙚 214 ภቶ‫⴫⊒ޔ‬⠪㧦᧻Ꮉ޽߅޿㧔㊁↢⎇࠮ࡦ࠲࡯㧕‫⺰⸛ޔ‬⠪㧦ᷡ
㐳 ⼾
Ecology of porcupines living rain forests of Borneo, Malaysia
╙ 50 ࿁
1 ᦬ 13 ᣣ㧔㊄㧕11:30㨪17:00 ℂቇ⎇ⓥ⑼ 1 ภ㙚 214 ภቶ‫⴫⊒ޔ‬⠪㧦ዊᎹ⹞ਫ‫⺰⸛ޔ‬⠪㧦㘵↰ᕺℂሶ㧔㊁↢⎇࠮ࡦ
࠲࡯㧕
Variation of characteristics of reading and writing difficulties in children with developmental disorders
╙ 51 ࿁
2 ᦬ 9 ᣣ㧔ᧁ㧕11:30㨪17:00 ℂቇ⎇ⓥ⑼ 1 ภ㙚 214 ภቶ‫⴫⊒ޔ‬⠪㧦㘵↰ᕺℂሶ㧔㊁↢⎇࠮ࡦ࠲࡯㧕
‫⺰⸛ޔ‬⠪㧦ߥߒ
Ecology of bush hyrax in a miombo woodland, Ugalla, Tanzania
㧔ᢥ⽿㧦ᱜ㜞ା↵㧕
㧙㧙
Χ. ᄢဳࡊࡠࠫࠚࠢ࠻
1. ⧯ᚻ⎇ⓥ⠪ࠗࡦ࠲࡯࠽࡚ࠪ࠽࡞࡮࠻࡟࡯࠾ࡦࠣ࡮ࡊࡠࠣ࡜ࡓ㧔ITP-HOPE㧕
ᣣᧄቇⴚᝄ⥝ળߩ‫⧯ޟ‬ᚻ⎇ⓥ⠪ࠗࡦ࠲࡯࠽࡚ࠪ࠽࡞࡮࠻࡟࡯࠾ࡦࠣ࡮ࡊࡠࠣ࡜ࡓ‫ࠆࠃߦޠ‬ഥᚑ੐ᬺߢ޽ࠆ‫ޕ‬ᱜᑼ
ߥ੐ᬺฬߪ‫ޔ‬๺ᢥ‫ੱޟ‬㘃ㅴൻߩ㔤㐳㘃⊛⿠Ḯߩ⸃᣿ߦะߌߚ⧯ᚻ⎇ⓥ⠪⢒ᚑ࿖㓙ࡊࡠࠣ࡜ࡓ HOPE‫⧷ޔޠ‬ᢥ
‫ޟ‬International Training Program for Young Researchers: Primate Origins of Human Evolution (HOPE)‫⇛ޕޠ‬⒓ࠍ‫ޔ‬
ITP-HOPE ߣߒߡ޿ࠆ‫ޕ‬
㔤㐳㘃⎇ⓥᚲ߇ਥ௅ߔࠆ㔤㐳㘃ቇߩ࿖㓙ㅪ៤ࠍ߼ߑߒߚ੤ᵹ੐ᬺࠍ HOPE ߣ✚⒓ߒߡ߈ߚ‫ޕ‬HOPE ߪ‫ੱޟޔ‬㑆ߩ
ㅴൻߩ㔤㐳㘃⊛⿠Ḯ‫ࠍޠ‬ᗧ๧ߔࠆ⧷ᢥฬ⒓‫ޟ‬Primate Origins of Human Evolution‫ߩޠ‬㗡ᢥሼߩࠕ࠽ࠣ࡜ࡓ㧔ਗߴᦧ߃
ߚ߽ߩ㧕ߢ޽ࠆ‫⧯ޕ‬ᚻ⎇ⓥ⠪ߦቇ໧ߩ⊒ዷߩ዁᧪ࠍ⸤ߒߚ޿‫ޟߩ߳᧪ᧂ߁޿߁ߘޕ‬Ꮧᦸ‫ߚ߼ߎࠍޠ‬๮ฬߢ޽ࠆ‫ޕ‬
HOPE ੐ᬺߪ‫ޔ‬ᐔᚑ 16 ᐕ 3 ᦬ߦ‫ޔ‬ᣣᧄቇⴚᝄ⥝ળߩ‫ޟ‬వ┵⎇ⓥ᜚ὐ੐ᬺ‫ߩޠ‬ណᛯ╙ 1 ภߣߒߡ⊒⿷ߒߚ‫ߩߎޕ‬
వ┵⎇ⓥ᜚ὐ੐ᬺ HOPE ߪ‫ޔ‬ᐔᚑ 16-20 ᐕᐲߩ 5 ᐕ㑆⛮⛯ߒߚ‫ޕ‬ᒰೋߪᣣ⁛ߩㅪ៤ߔߥࠊߜ੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲ
ߣࡑ࠶ࠢࠬࡊ࡜ࡦࠢㅴൻੱ㘃ቇ⎇ⓥᚲߣߩ 2 ࿖㑆ㅪ៤ߣߒߡᆎ߹ߞߚ‫ߩߘޕ‬ᓟ‫⧷☨ޔ‬દ੽ߣ㗅ᰴടࠊߞߡ‫ޔ‬5 ᐕ㑆
ࠍ߆ߌߡవㅴ 6 ߆࿖ߩㅪ៤૕೙߇᭴▽ߐࠇߚ‫ޕ‬
వ┵⎇ⓥ᜚ὐ੐ᬺ HOPE ߩᓟ⛮߇‫ޔ‬ITP-HOPE ੐ᬺߢ޽ࠆ‫ޕ‬ᐔᚑ 21-25 ᐕᐲߩ 5 ᐕ㑆ណᛯߐࠇߡ޿ࠆ‫ޕ‬ᐔᚑ 20
ᐕᐲߦ‫ޔ‬㔤㐳㘃⎇ⓥᚲ߇Უ૕ߣߥߞߡ‫ޔ‬੩ㇺᄢቇߦ㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯WRC ߣ޿߁ᣂߚߥㇱዪ߇⺀↢ߒߚ‫ߘޕ‬
ߎߢ ITP-HOPE ߢߪ 2 ߟߩᆌᆂㇱዪ߇ㅪ៤ߒߡ‫⧯ޔ‬ᚻ⎇ⓥ⠪ߩࠗࡦ࠲࡯࠽࡚ࠪ࠽࡞࡮࠻࡟࡯࠾ࡦࠣ࡮ࡊࡠࠣ࡜ࡓߩ
⏕┙ࠍ⋡ᜰߒߚ‫ޕ‬೨りߩవ┵⎇ⓥ᜚ὐ੐ᬺ HOPE ߣߩ㆑޿ߪ‫ޔ‬ਥߦ 3 ὐߦⷐ⚂ߢ߈ࠆ‫ޕ‬Ԙ㔤㐳㘃ߛߌߢߥߊߘࠇએ
ᄖߩേ‛ߩ⎇ⓥ߽ኻ⽎ߦߒߚ‫ޕ‬ԙᄖ࿖ߩࡄ࡯࠻࠽࡯ᯏ㑐߇వㅴ⻉࿖ߛߌߢߥߊ‫࡮ࠞ࡝ࡈࠕޔ‬ਛධ☨࡮᧲ධࠕࠫࠕߥ
ߤߩ⊒ዷㅜ਄࿖ߦᐢ߇ߞߚ‫ޕ‬ԚITP ੐ᬺߩߘ߽ߘ߽ߩ೙⚂ߣߒߡ‫ޔ‬2 ߆᦬㑆એ਄ߩᄖ࿖ṛ࿷ࠍᔅⷐߣߔࠆ‫ޕ‬
ᐔᚑ 23 ᐕᐲߪ‫ޔ‬ITP-HOPE ੐ᬺߩ 3 ᐕ⋡ߛߞߚ‫ޕ‬એਅߦ 3 ߟߩࡊࡠࠣ࡜ࡓߩ᭎ⷐࠍ⹦ㅀߔࠆ‫ޕ‬
1) ⎇ⓥᯏ㑐੤ᵹᢎ⢒ࡊࡠࠣ࡜ࡓ㧔Inter-Laboratory Training Program㧕
એਅߩ⎇ⓥᯏ㑐╬ߦ⧯ᚻ⎇ⓥ⠪ 4 ฬࠍᵷ㆜ߒ‫ޔ‬ታ㛎ቶ߅ࠃ߮㊁ᄖ⺞ᩏ࿾ߢߩ౒ห⎇ⓥࠍⴕߞߚ‫ࡊࠗ࡜ޔ࠷ࠗ࠼ޕ‬
࠴࠶ࡅ‫ࠢࡦ࡜ࡊࠬࠢ࠶ࡑޔ‬ㅴൻੱ㘃ቇ⎇ⓥᚲ㧔Ỉᩙ㧕‫࠼ࡦࠗߪࠇߎޕ‬㜞ේߢߩ࠼࡯࡞ߩ⎇ⓥࠍ㐿ᆎߔࠆߚ߼ߦᔅⷐ
ߥࠬ࠹࠶ࡊߢ޽ࠆ‫ࠕ࠾࡞ࠜࡈ࡝ࠞޔࠞ࡝ࡔࠕޔߚ߹ޕ‬ᄢቇࠨࡦ࠺ࠖࠛࠧᩞ㧔ࡂࡦ࡮࠰ࠫࡘࡦ‫ޔ‬ศ↰㧕‫ࠛࠖ࠺ࡦࠨޕ‬
ࠧേ‛࿦ߣㅪ៤ߒߡ‫ޔࠞ࡞ࠗޔࡏࡁࡏޔ‬᳓᫮ູ੃㘃ߩ⎇ⓥࠍ㐿ᆎߔࠆߚ߼ߩᔅⷐߥࠬ࠹࠶ࡊߢ޽ࠆ‫ࡦࠧ࡟ࠝߚ߹ޕ‬
㔤㐳㘃࠮ࡦ࠲࡯ߢേ‛⑔␩ቇߩ੤ᵹࡊࡠࠣ࡜ࡓߣߒߡ‫⧯ޔ‬ᚻ⎇ⓥ⠪ࠍᵷ㆜ߒߚ㧔ዊୖ㧕‫⎇ޔ߅ߥޕ‬ⓥᯏ㑐੤ᵹࡊࡠ
ࠣ࡜ࡓߩ౞Ṗߥታᣉߩߚ߼ߦᢎ⡯ຬࠍࡄ࡯࠻࠽࡯ᯏ㑐ߦᵷ㆜ߒߚ‫ࠬࠢ࠶࠮ࠨޕ‬ᄢቇߣࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇࠨࡦ࠺ࠖ
ࠛࠧᩞߦ‫ޔ‬ᤓᐕᐲߦ߭߈ߟߠ߈ᢎຬ 2 ฬ㧔ࡈ࡟࠶࠼࡮ࡌ࡞ࠦࡆ࠶࠴ߣ࠺ࠗࡆ࠶࠼࡮ࡅ࡞㧕ࠍᵷ㆜ߒߚ‫ࠕࠪ࡯࡟ࡑޕ‬
⑼ቇᄢቇ㧔᧻ᴛ㧕
‫ࠕ࠾ࠩࡦ࠲ޔ‬㊁↢േ‛⎇ⓥᚲ㧔ኋベ㧕‫ޔ‬᪸⧎ᅚሶᄢቇ㧔౎ᧁ㧘ਤ⠀㧕
‫ޔ‬ᐢ⷏Ꮷ▸ᄢቇ㧔㜞੗㧕
‫ߚ߹ޔ‬
࠲ࠗߩ࠴ࡘ࡜ࡠࡦࠦࡦᄢቇਥ௅ߩ㔤㐳㘃ࠪࡦࡐࠫ࠙ࡓߦ‫ޔ‬ᢎ⡯ຬࠍᵷ㆜ߒߚ㧔ự↰㧘ጟᧄ㧘਄၂㧕‫ޕ‬
2) ౒ห㊁ᄖ⺞ᩏࡊࡠࠣ࡜ࡓ㧔Collaborative On-site Research Program㧕
એਅߩ㊁ᄖ⺞ᩏ࿾ߦ⧯ᚻ⎇ⓥ⠪ 9 ฬࠍᵷ㆜ߒ‫࡯࠽࠻࡯ࡄޔ‬ᯏ㑐ߩ⎇ⓥ⠪ߣㅪ៤ߒߚ⎇ⓥࠍⴕߞߚ‫ޕ‬᜝ᒐౝߪ‫ޔ‬ㅪ
៤⎇ⓥࠍⴕߞߚࡄ࡯࠻࠽࡯ᯏ㑐ߢ޽ࠆ‫ޕ‬
ࠡ࠾ࠕ࡮ࡏ࠶࠰࠙㧔ࡏ࠶࠰࠙ⅣႺ⎇ⓥᚲ㧕‫⮮ޔ‬Ỉ
ࠦࡦࠧ࡮ࡢࡦࡃ㧔↢ᘒ᫪ᨋ⎇ⓥ࠮ࡦ࠲࡯㧕‫ޔ‬ጊᧄ
ࠦࡦࠧ࡮ࡢࡦࡃ㧔↢ᘒ᫪ᨋ⎇ⓥ࠮ࡦ࠲࡯㧕‫ࡦࠫࡦࡈ࡮࡯ࡘ࡝ޔ‬
ࠦࡦࠧ࡮࡞ࠝ࡯Ꮉቇⴚ଻⼔඙㧔↢ᘒ᫪ᨋ⎇ⓥ࠮ࡦ࠲࡯㧕‫ޔ‬ဈᏎ
࠲ࡦࠩ࠾ࠕ࡮ࠞ࠲ࡆ࿖┙౏࿦㧔࠲ࡦࠩ࠾ࠕ㊁↢േ‛⎇ⓥᚲ)‫ޔ‬㥲⮮
࠲ࠗ㧔࠴ࡘ࡜ࡠࡦࠦࡦᄢቇ㧕‫ޔ‬቟੗
ࡑ࡟࡯ࠪࠕ࡮࠲ࡆࡦ㊁↢↢‛଻⼔඙㧔ࡑ࡟࡯ࠪࠕ࡮ࠨࡃᄢቇ㧕‫ޔ‬ਛᨋ
ࠗࡦ࠼ࡀࠪࠕ࡮ࡄࠟࡦ࠳࡜ࡦ࿖┙౏࿦㧔ࠕࡦ࠳࡜ࠬᄢቇ㧕‫ޔ‬ㄞ
࠴࡝࡮ࡊࠛ࡞࠻࡮ࡕࡦᴪጯ࡮ࡑ࠯࡜ࡦᶏጽᴪጯ㧔ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇࠨࡦ࠺ࠖࠛࠧᩞ㧕‫ޔ‬᳓ญ
3) ࿖㓙ࡢ࡯࡚ࠢࠪ࠶ࡊ㧔Annual International Workshop㧕
ᐔᚑ 23 ᐕᐲ߽ 2 ߟߩ࿖㓙ࡢ࡯࡚ࠢࠪ࠶ࡊࠍ߅ߎߥߞߚ‫ࠕࠪ࡯࡟ࡑޔߪߟߣ߭ޕ‬㗔ࡏ࡞ࡀࠝߩࡑ࡝ࠝࡌࠗ࠭ࡦ࡮
ࡈࠖ࡯࡞࠼࠮ࡦ࠲࡯ߣࠗࡦࡃࠢࠠࡖ࠾ࠝࡦ࡮ࡈࠖ࡯࡞࠼࠮ࡦ࠲࡯ߩᵴ↪ࠍߐߋࠆ࿖㓙ࡢ࡯࡚ࠢࠪ࠶ࡊߢ޽ࠆ‫ޕ‬6 ᦬
ߦ‫⑼ࠕࠪ࡯࡟ࡑޔ‬ቇᄢቇߣࡑ࡟࡯ࠪࠕ࡮ࠨࡃᄢቇߣߩ౒௅ߢ߅ߎߥߞߚ‫ޕ‬೎ㅜ⚻⾌ߢߩភ⟎ࠍ฽߼ߡ‫ߦࠇߎޔ‬ᢎຬ࡮
ᄢቇ㒮↢╬ 21 ฬࠍᵷ㆜ߒߚ‫ ߪߟߣ߭߁߽ޕ‬7 ᦬ߦ‫߽ࠇߎޔ‬೎ㅜ⚻⾌ߢ‫ޔ‬੩ㇺᄢቇࡉ࡯࠲ࡦ෹ᅢࡊࡠࠣ࡜ࡓࠍታᣉ
ߒ‫ޔ‬7 ᦬ߩ╙ 5 ⸰໧࿅߇‫ޟߢࡦ࠲࡯ࡉޔ‬ஜᐽ࡮ᢥൻ࡮↢ᘒ♽‫ߚߒߦࡑ࡯࠹ࠍޠ‬࿖㓙ࡢ࡯࡚ࠢࠪ࠶ࡊࠍ㐿௅ߒߚ‫޿ޕ‬
ߕࠇ߽ࡄ࡯࠻࠽࡯ᯏ㑐ߣߩ౒ห㐿௅ࡊࡠࠣ࡜ࡓߣߒߡᶏᄖߢታᣉߔࠆ߽ߩߢ޽ࠆ‫ޔߚ߹ޕ‬࿖ౝ㐿௅ߩ࿖㓙ࡢ࡯ࠢࠪ
࡚࠶ࡊߣߒߡߪ‫ ╙ޔ‬15 ࿁࿖㓙ᗧ⼂⑼ቇળࠍ੩ㇺߢ㐿௅ߒߚ‫ߚߒ߁ߎޕ‬࿖㓙ࡢ࡯࡚ࠢࠪ࠶ࡊࠍㅢߓߡ‫ޔ‬᰷☨ߥࠄ߮
ߦ⊒ዷㅜ਄࿖ߩ⎇ⓥ⠪ߣᵴ⊒ߥᗧ⷗੤឵ࠍ߅ߎߥߞߚ‫ޕ‬
ᐔᚑ 23 ᐕᐲߩว⸘ߪએ਄ߩ‫⧯ޔ‬ᚻ⎇ⓥ⠪ 13 ઙ෸߮ᢎ⡯ຬ 13 ઙߩ⸘ 26 ઙߢ޽ࠆ‫ޕ‬ᵷ㆜ᣣᢙߪ‫⧯ޔ‬ᚻ⎇ⓥ⠪ 1,404
㧙㧙
ᣣ෸߮ᢎ⡯ຬ 124 ᣣߩ⸘ 1,528 ᣣߦߥߞߚ‫ޕ‬ᵷ㆜ታ❣ࠍએਅߦ␜ߔ‫ޕ‬
ᵷ㆜⠪᳁ฬ
ᚲዻ
⡯ฬ╬
ᵷ㆜వ࿖ฬ
ᵷ㆜వᯏ㑐࡮⚵❱╬ฬ
ᵷ㆜㐿ᆎᣣ
ᵷ㆜⚳ੌᣣ
ᵷ㆜
ᣣᢙ
ጊᧄ⌀਽
㔤㐳㘃⎇ⓥᚲ
․ቯഥᢎ
ࠦࡦࠧ
↢ᘒ᫪ᨋ⎇ⓥ࠮ࡦ࠲࡯
H23.6.14
H23.8.22
70
ਛᨋ㓷
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻⺖⒟ 2 ᐕ
ࡑ࡟࡯ࠪࠕ
ࡑ࡟࡯ࠪࠕ࡮ࠨࡃᄢቇ
H23.5.19
H23.10.11
146
H23.8.18
H23.12.20
125
Ỉᩙ⑲ᄥ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻⺖⒟ 2 ᐕ
࠼ࠗ࠷
ࡑ࠶ࠢࠬࡊ࡜ࡦࠢ
ੱ㘃ㅴൻ⎇ⓥᚲ
㥲⮮⟤଻
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻⺖⒟ 2 ᐕ
࠲ࡦࠩ࠾ࠕ
࠲ࡦࠩ࠾ࠕ㊁↢േ‛⎇ⓥᚲ
H23.7.1
H23.11.4
127
H23.9.19
H24.3.16
180
S Han
㔤㐳㘃⎇ⓥᚲ
ୃ჻⺖⒟ 1 ᐕ
ࠕࡔ࡝ࠞ
ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇ
ࠨࡦ࠺ࠖࠛࠧᩞ
H Ryu
㔤㐳㘃⎇ⓥᚲ
ୃ჻⺖⒟ 1 ᐕ
ࠦࡦࠧ
↢ᘒ᫪ᨋ⎇ⓥ࠮ࡦ࠲࡯
H23.8.5
H24.2.7
187
H23.9.29
H23.12.4
67
61
᳓ญᄢテ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻⺖⒟ 1 ᐕ
ࠕࡔ࡝ࠞ
ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇ
ࠨࡦ࠺ࠖࠛࠧᩞ
ㄞᄢ๺
㔤㐳㘃⎇ⓥᚲ
ഥᢎ
ࠗࡦ࠼ࡀࠪࠕ
ࡏࠧ࡯࡞ㄘ⑼ᄢቇ
H23.11.10
H24.1.9
⮮Ỉ㆏ሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
․ቯഥᢎ
ࠡ࠾ࠕ
ࡏ࠶࠰࠙ⅣႺ⎇ⓥᚲ
H23.10.25
H23.12.26
63
ဈᏎື਽
㔤㐳㘃⎇ⓥᚲ
⎇ⓥຬ
ࠦࡦࠧ
↢ᘒ᫪ᨋ⎇ⓥ࠮ࡦ࠲࡯
H23.10.12
H24.2.28
140
ዊୖආବ
㔤㐳㘃⎇ⓥᚲ
ᢎോ⵬૒ຬ
ࠕࡔ࡝ࠞ
ࠝ࡟ࠧࡦ࿖┙㔤㐳㘃
⎇ⓥ࠮ࡦ࠲࡯
H23.12.3
H24.3.1
90
቟੗ᣧ♿
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻⺖⒟ 1 ᐕ
࠲ࠗ
࠴ࡘ࡜ࡠࡦࠦࡦᄢቇ
H23.12.11
H24.3.1
82
ศ↰ᒎ↢
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻⺖⒟ 2 ᐕ
ࠕࡔ࡝ࠞ
ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇ
H24.1.25
H24.3.30
66
ጟᧄቬ⵨
㔤㐳㘃⎇ⓥᚲ
ᢎ᝼
࠲ࠗ
࠴ࡘ࡜ࡠࡦࠦࡦᄢቇ
H23.5.18
H23.5.21
4
ự↰ⓛ
㔤㐳㘃⎇ⓥᚲ
ᢎ᝼
࠲ࠗ
࠴ࡘ࡜ࡠࡦࠦࡦᄢቇ
H23.5.16
H23.5.23
8
਄၂ᵏᶈ
㔤㐳㘃⎇ⓥᚲ
⎇ⓥഥᚑដ㐳
࠲ࠗ
࠴ࡘ࡜ࡠࡦࠦࡦᄢቇ
H23.5.16
H23.5.22
7
DA Hill
㔤㐳㘃⎇ⓥᚲ
ቴຬᢎ᝼
⧷࿖
ࠨ࠮࠶ࠢࠬᄢቇ
H23.9.6
H23.9.23
18
ዊ㊁৻ઍ
㔤㐳㘃⎇ⓥᚲ
✚ോដ㐳
ࡉ࡯࠲ࡦ
ࡉ࡯࠲ࡦ₺┙ᄢቇ
H23.7.5
H23.7.13
9
ኋベࡑࡒ
㔤㐳㘃⎇ⓥᚲ
․ቯ⡯ຬ
࠲ࡦࠩ࠾ࠕ
࠳࡞ࠛࠬࠨ࡜࡯ࡓᄢቇ
H23.8.17
H23.8.26
10
᧻ᴛື㇢
㔤㐳㘃⎇ⓥᚲ
ᢎ᝼
ࡑ࡟࡯ࠪࠕ
ࡑ࡟࡯ࠪࠕ⑼ቇᄢቇ
H23.9.5
H23.9.19
15
FB Bercovitch
㔤㐳㘃⎇ⓥᚲ
․ቯᢎ᝼
ࠕࡔ࡝ࠞ
ࠞ࡝ࡈࠜ࡞࠾ࠕᄢቇ
H23.11.16
H23.12.1
16
౎ᧁቯⴕ
㔤㐳㘃⎇ⓥᚲ
੐ോ㐳
㖧࿖
᪸⧎ᅚሶᄢቇ
H23.10.20
H23.10.22
3
ਤ⠀⟤Ꮤ
㔤㐳㘃⎇ⓥᚲ
․ቯ⡯ຬ
㖧࿖
᪸⧎ᅚሶᄢቇ
H23.10.20
H23.10.22
3
㜞੗ᱜᚑ
㔤㐳㘃⎇ⓥᚲ
ᢎ᝼
ਛ࿖
ᐢ⷏Ꮷ▸ᄢቇ
H24.3.12
H24.3.15
4
ᐘፉม㇢
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ᢎ᝼
ࡑ࡟࡯ࠪࠕ
ࠨࡃᄢቇ
H23.6.17
H23.7.1
15
↰ਛᱜਯ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ಎᢎ᝼
ࡑ࡟࡯ࠪࠕ
ࠨࡃᄢቇ
H23.6.20
H23.7.1
12
ᐔᚑ 23 ᐕᐲߩว⸘ߪએ਄ߩ‫⧯ޔ‬ᚻ⎇ⓥ⠪ 13 ઙ෸߮ᢎ⡯ຬ 13 ઙߩ⸘ 26 ઙߢ޽ࠆ‫ޕ‬ᵷ㆜ᣣᢙߪ‫⧯ޔ‬ᚻ⎇ⓥ⠪ 1,404
ᣣ෸߮ᢎ⡯ຬ 124 ᣣߩ⸘ 1,528 ᣣߦߥߞߚ‫✚⾌⚻ޕ‬㗵ߪ‫ޔ‬16,400,000 ౞ߛߞߚ‫ޕ‬
㧔ᢥ⽿㧦᧻ᴛື㇢࡮਄၂ᵏᶈ㧕
2. ⚵❱⊛ߥ⧯ᚻ⎇ⓥ⠪╬ᶏᄖᵷ㆜ࡊࡠࠣ࡜ࡓ㧔AS-HOPE㧕
ᣣᧄቇⴚᝄ⥝ળߩ‫⧯ߥ⊛❱⚵ޟ‬ᚻ⎇ⓥ⠪ᶏᄖᵷ㆜ࡊࡠࠣ࡜ࡓ‫ࠆࠃߦޠ‬ഥᚑ੐ᬺߢ޽ࠆ‫ޕ‬ᱜᑼߥ੐ᬺฬߪ‫ޔ‬๺ᢥ‫ੱޟ‬
㑆ߩᧄᕈߩㅴൻ⊛⿠Ḯߦ㑐ߔࠆవ┵⎇ⓥ‫⧷ޔޠ‬ᢥ‫ޟ‬The advanced studies on the evolutionary origins of human nature‫ޕޠ‬
⇛⒓ࠍ‫ޔ‬AS-HOPE ߣߒߡ޿ࠆ‫ޕ‬੐ᬺታᣉᦼ㑆ߪ‫ޔ‬ᐔᚑ 22 ᐕ㧔2010 ᐕ㧕3 ᦬ 1 ᣣ߆ࠄ‫ޔ‬ᐔᚑ 25 ᐕ㧔2013 ᐕ㧕2 ᦬
28 ᣣ߹ߢߩ 3 ᐕ㑆ߢ޽ࠆ‫ޕ‬੐ᬺታᣉ⚻⾌✚㗵㧔ഥᚑ᳿ቯ㗵㧕ߪ‫ޔ‬77,819,000 ౞ߢ޽ࠆ‫ޕ‬
㔤㐳㘃ቇߩ࿖㓙ㅪ៤ࠍ߼ߑߒߚ੐ᬺࠍ HOPE ߣ✚⒓ߒߡ޿ࠆ‫ޕ‬11 ߆᦬వⴕߒߡᆎ߹ߞߚ ITP-HOPE ੐ᬺ㧔ᐔᚑ
21-25 ᐕᐲߩ 5 ᐕ㑆ណᛯ㧕ߣ⋧੕⵬ቢ⊛ߥ੐ᬺߣ૏⟎ઃߌࠄࠇࠆ‫ޕ‬੩ㇺᄢቇ㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯WRC ߣㅪ៤ߒߡ‫ޔ‬
ቇㇱቇ↢߽฽߼ߚ⧯ᚻ⎇ⓥ⠪╬ߩᶏᄖᵷ㆜ࠍ߅ߎߥ߁‫ޕ‬ITP-HOPE ੐ᬺߣߩᄢ߈ߥ㆑޿ߪ‫ޔ‬ਥߦ 4 ὐߦⷐ⚂ߢ߈ࠆ‫ޕ‬
Ԙ⧯ᚻ⎇ⓥ⠪ߩኻ⽎ࠍᐢߍߡቇㇱቇ↢߽ᷰ⥶߇น⢻ߣߒߚ‫ޕ‬੩ㇺᄢቇߢ޽ࠇ߫ᚲዻቇㇱࠍ໧ࠊߥ޿‫ޕ‬ԙᄢቇ㒮↢ߪ
㔤㐳㘃⎇ⓥᚲߥ޿ߒ㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯ߩᚲዻࠍේೣߣߔࠆ߇‫⑼‛↢ޔ‬ቇኾ᡹ߩࠣࡠ࡯ࡃ࡞ COE ੐ᬺ㧔㒙ᒻᷡ
๺ઍ⴫㧕ߣㅪ៤ߒߡ޿ࠆߎߣߦ㐓ߺߡ‫ޔ‬ℂቇ⎇ⓥ⑼ߩ↢‛⑼ቇኾ᡹ߩᄢቇ㒮↢࡮ቇㇱ↢ߢ޽ࠇ߫ෳടࠍ⹺߼ࠆ‫ޕ‬Ԛ
ITP ੐ᬺߩࠃ߁ߥᵷ㆜ᦼ㑆ߩߒ߫ࠅ߇ߥߊ‫ޔ‬2 ߆᦬એ਄߇ផᅑߐࠇࠆ߇‫߽ࠅࠃࠇߘޔ‬⍴ᦼߩᶏᄖᵷ㆜߽น⢻ߢ޽ࠆ‫ޕ‬
ԛ⧯ᚻ⎇ⓥ⠪߇ਥኻ⽎ߢ޽ࠆߎߣߦᄌࠊࠅߪߥ޿߇‫ޔ‬੐ᬺࠍ౞Ṗߦㅴ߼ࠆߚ߼ߩ⎇ⓥㅪ⛊߿‫ޔ‬ቇㇱቇ↢ߩᶏᄖታ⠌
㧙㧙
ߩᒁ₸ߩߚ߼ߦ‫ޔ‬หⴕߔࠆᢎ᝼߿ಎᢎ᝼ߩᷰ⥶߽⹺߼ࠆ‫ޕ‬
ᐔᚑ 21 ᐕᐲᧃߦᆎ߹ࠅᐔᚑ 22 ᐕᐲߦ⛮⛯ߐࠇ‫ޔ‬ᐔᚑ 23 ᐕᐲߪᧄᩰ⊛ߦⒿ௛ߔࠆᄙ↥ߥ 1 ᐕߛߞߚ‫ޕ‬ᒰೋ⸘↹
ㅢࠅ‫ޔ‬5 ߟߩၮᧄࡊࡠࠣ࡜ࡓࠍ⛮⛯ߒߚ‫ޕ‬Ԙ‫౒ޟ‬ห㊁ᄖ⺞ᩏࡊࡠࠣ࡜ࡓ‫ޔޠ‬ԙ‫⎇ޟ‬ⓥᯏ㑐੤ᵹᢎ⢒ࡊࡠࠣ࡜ࡓ‫ޔޠ‬
Ԛ‫ޟ‬࿖㓙ࡢ࡯࡚ࠢࠪ࠶ࡊ‫ޔޠ‬ԛ‫ޟ‬ㅢᐕ⺞ᩏࡊࡠࠣ࡜ࡓ‫ޔޠ‬Ԝ‫ޟ‬ቇㇱቇ↢⍴ᦼ㊁ᄖ⺞ᩏࡊࡠࠣ࡜ࡓ‫߁ߩߎޕࠆ޽ߢޠ‬
ߜ‫ޔ‬ԘߪᲤᐕᐲᄌࠊࠆᄙ᭽ߥ⎇ⓥߢ޽ࠆ‫ޕ‬ᐔᚑ 23 ᐕᐲߪ‫ޔ‬ධ☨ߩࡉ࡜ࠫ࡞ߣ࠴࡝ߩ 2 ߆࿖‫࡮࠳ࡦࠟ࠙ߩࠞ࡝ࡈࠕޔ‬
ࠤ࠾ࠕ࡮࠲ࡦࠩ࠾ࠕ࡮ࠡ࠾ࠕߩ 4 ߆࿖‫ ߩࠗ࠲࡮ࡦ࠲࡯ࡉ࡮ࡓ࠽࠻ࡌ࡮ࠕࠪ࡯࡟ࡑߩࠕࠫࠕޔ‬4 ߆࿖ߦ⧯ᚻ⎇ⓥ⠪ࠍ
ᵷ㆜ߒߡ౒ห㊁ᄖ⺞ᩏࠍ߅ߎߥߞߚ‫ޕ‬ԙߦߟ޿ߡߪ࠼ࠗ࠷ߩࡑ࠶ࠢࠬࡊ࡜ࡦࠢㅴൻੱ㘃ቇ⎇ⓥᚲࠍਛᩭㅪ៤ᯏ㑐ߣ
ߒߡᲤᐕታᣉߔࠆ߽ߩߢ޽ࠆ‫ޕ‬ᐔᚑ 23 ᐕᐲ߽‫⧯ޔ‬ᚻ⎇ⓥ⠪߇ࡑ࠶ࠢࠬࡊ࡜ࡦࠢㅴൻੱ㘃ቇ⎇ⓥᚲߢߣߊߦᗵⷡฃ
ኈ૕ߩࠥࡁࡓ⸃ᨆߦߟ޿ߡߩ੤ᵹࠍ߅ߎߥߞߚ‫ࡏ࡯ࡍ࡮࠹ࡦࡃࠬޕ‬ᢎ᝼ߣ౒หߢ‫ޔ‬ᒁ߈⛯޿ߡ๧ⷡ࡮༦ⷡߩฃኈㆮ
વሶߩ⎇ⓥࠍ߅ߎߥߞߚ‫ࡠ࠮ࡑ࠻࡮࡞ࠤࠗࡑޕ‬ᢎ᝼ߣߪ‫ੱޔ‬㑆એᄖߩ㔤㐳㘃ࠍኻ⽎ߣߒߚ␠ળ⊛⍮ᕈߣߊߦᗧ࿑ᕈ
ߩℂ⸃߿੕ᕺᕈߩ⎇ⓥࠍ߅ߎߥߞߡ޿ߡ੹ᐕᐲߩᵷ㆜ߪߥ߆ߞߚ߇‫ޔ‬೎ㅜ⾌↪ߢ᜗⡜߇޽ࠅ‫⋧ޔ‬੕੤ᵹ߇ㅴࠎߛ‫ޕ‬
߹ߚ‫ߩઁߩߘޔ‬࿖‫߽ߢޘ‬ห᭽ߩ⎇ⓥᯏ㑐੤ᵹࠍ߅ߎߥߞߚ‫ߦߊߣޕ‬᰷☨ߩవㅴ 6 ߆࿖‫☨ޔ‬࿖࡮࠼ࠗ࠷࡮ࠗࠡ࡝ࠬ࡮
ࡈ࡜ࡦࠬ࡮ࠬࠗࠬ࡮ࠝ࡯ࠬ࠻࡝ࠕߩ⎇ⓥᯏ㑐ߣߩ੤ᵹࠍ߅ߎߥߞߚ‫ޕ‬Ԛߦߟ޿ߡߪᶏᄖࡄ࡯࠻࠽࡯⎇ⓥᯏ㑐ߣ⋧⺣
ߒߡታᣉߔࠆ߽ߩߢ޽ࠆ‫ߣࡓ࡜ࠣࡠࡊߩઁޕ‬૬↪ߒߡ‫ߢࠕࠪ࡯࡟ࡑޔ‬㊁↢଻ోߦ㑐ߔࠆ࿖㓙ࡢ࡯࡚ࠢࠪ࠶ࡊࠍ߅ߎ
ߥߞߚ‫౒ߩߢࡦ࠲࡯ࡉߚ߹ޕ‬ห⎇ⓥࠍ㐿ᆎߒஜᐽ࡮ᢥൻߦὶὐࠍ޽ߡߚ࿖㓙ࡢ࡯࡚ࠢࠪ࠶ࡊࠍ߅ߎߥߞߚ‫ޕ‬ԛߦߟ
޿ߡߪ‫࡮ࠕࠪ࡯࡟ࡑޔ࠙࠰࠶ࡏߩࠕ࠾ࠡޔࡃࡦࡢߩࠧࡦࠦޔ࠭ࡦ࡝ࠞߩ࠳ࡦࠟ࠙ޔ࡜ࠟ࠙࡮࡟ࡂࡑߩࠕ࠾ࠩࡦ࠲ޔ‬
ࡏ࡞ࡀࠝߩ࠳࠽ࡓ࡮ࡃ࡟ࠗߣ޿߁ 5 ߆࿖ߩ⺞ᩏၮ࿾ࠍኻ⽎ߦ 3 ᐕ㑆⛮⛯ߒߡ߅ߎߥ߁㐳ᦼ⛮⛯⺞ᩏࡊࡠࠣ࡜ࡓߢ޽
ࠆ‫ޕ‬㔤㐳㘃߅ࠃ߮ߘߩઁߩ㊁↢േ‛ߩ㊁ᄖ⎇ⓥࠍታᣉߒߚ‫ޕ‬ㅢᐕߩ⛮⛯⎇ⓥ૕೙ߣ⧯ᚻ⎇ⓥ⠪ߩᶏᄖᵷ㆜ߦࠃߞߡ
㘧べ⊛ߥ⎇ⓥᚑᨐߩะ਄߇ᓧࠄࠇࠆߣᦼᓙߒߡ޿ࠆ‫౒ߩߢࠕࠪ࡯࡟ࡑޔߪ߈ߴߔ⸥․ޕ‬ห⎇ⓥߩ⪺ߒ޿⌕ታߥㅴዷ
ߢ޽ࠆ‫࡯࡟ࡑޕ‬ඨፉߢ㊁↢଻ోߣ㊁↢ᓳᏫߩࡊࡠࠣ࡜ࡓ߇ࡑ࠶࠴ࡦࠣࡈࠔࡦ࠼ߩ♖␹ߢᆎേߒ‫ࡓ࠽࠳ߩࠝࡀ࡞ࡏޔ‬
ࡃ࡟ࠗߢߪᐔᚑ 22 ᐕߦ⺞ᩏዊደ߇㐿⸳ߐࠇߡ㐳ᦼ⛮⛯⎇ⓥߩ㆏߇㐿ߌߚ‫⿠ࠍࠇߎޕ‬ὐߦࡑ࡟࡯ࠪࠕ࡮ࠨࡃ⽷࿅ߩ
ᡰេߢ⎇ⓥ੤ᵹ߇ᩰᲑߦߔߔࠎߛ‫ޕ‬వᣇߩฃߌ౉ࠇᯏ㑐߇ṛ࿷⾌ࠍ⽶ᜂߔࠆࡑ࠶࠴ࡦࠣᣇᑼ߇⏕┙ߒ‫ߥ߆ߕࠊޔ‬ᷰ
⥶⾌↪ߛߌߢᄙᢙߩ⧯ᚻ⎇ⓥ⠪ࠍㅍࠅㄟ߼ࠆࠃ߁ߦߥߞߚ‫ޕ‬ᐔᚑ 23 ᐕᐲߪ‫ౕޔ‬૕⊛ߦߪࡏ࡞ࡀࠝߩࡑ࡝ࠕ࠙࡮ࡌ
ࠗ࠭ࡦߩ⎇ⓥ߇ㅴࠎߛ‫ޕ‬Ԝߩቇㇱቇ↢⍴ᦼ㊁ᄖ⺞ᩏࡊࡠࠣ࡜ࡓߦߟ޿ߡߪቇㇱ㧝ᐕ↢߆ࠄෳടߢ߈ࠆᢎ⢒ࡊࡠࠣ࡜
ࡓߢ޽ࠆ‫ޕ‬࿖ᄖߩ⎇ⓥၮ࿾ࠍၮ⋚ߦዷ㐿ߒߚ‫ౕޕ‬૕⊛ߦߪ‫⺞ߩࠗ࡟ࡃࡓ࠽࠳ߩࠝࡀ࡞ࡏޔ‬ᩏၮ࿾ࠍ೑↪ߒߚᲧセⴕ
േቇߩታ⠌ߦቇㇱ↢ࠍෳടߐߖߚ‫⑼ࡓࡁࠥޔߚ߹ޕ‬ቇ㑐ㅪߩቇㇱቇ↢੤ᵹߣߒߡ࠼ࠗ࠷ߩ࠼࡟ࠬ࠺ࡦᄢቇߢࠥࡁࡓ
⑼ቇታ⠌ࠍታᣉߒ‫ޔ‬ቇㇱቇ↢ߦᣧᦼߦቇ໧ߣᶏᄖ⚻㛎ࠍⓍ߹ߖࠆߎߣ߇ߢ߈ߚ‫ᦨޕ‬ᓟߦ‫ޟޔ‬੩ㇺᄢቇࡉ࡯࠲ࡦ෹ᅢ
ࡊࡠࠣ࡜ࡓ‫ᧄ߇ޠ‬ቇߦ⊒⿷ߒߚߎߣࠍฃߌߡ‫ߩߢࡦ࠲࡯ࡉޔ‬ቇㇱ↢ߩᵴേࠍᡰេߒ‫ޔ‬ᣂߒ޿㊁ᄖ⎇ⓥࡈࠖ࡯࡞࠼ߩ
⏕┙ߦነਈߒߚ‫ޕ‬
ᐔᚑ 23 ᐕᐲߩᶏᄖᵷ㆜ᢙߩว⸘ߪ 74 ઙߢ޽ࠆ‫ޕ‬ᵷ㆜ᣣᢙߩว⸘ߪ 1,880 ᣣߦߥߞߚ‫✚⾌⚻ޕ‬㗵ߪ‫ޔ‬25,860,424
౞ߛߞߚ‫ޔ߅ߥޕ‬ᐔᚑ 22 ᐕᐲߩว⸘ߪ 78 ઙ‫ޔ‬ᵷ㆜ᣣᢙߪ 1,620 ᣣ‫✚⾌⚻ޔ‬㗵ߪ‫ޔ‬22,604,187 ౞ߛߞߚߩߢ‫߷߶ޔ‬
หⷙᮨߩᵷ㆜߇ߢ߈ߚߣ޿߃ࠆ‫ޕ‬ᵷ㆜⠪࡝ࠬ࠻ߪ‫ޔ‬એਅߩߣ߅ࠅߢ޽ࠆ‫ޕ‬
ᵷ㆜⠪╬
᳁ฬ
ᚲዻ
⡯ ૏
ᷰ⥶వߩ࿖╬
ᵷ㆜వߩ⎇ⓥ
ᯏ㑐࡮⚵❱╬ฬ
ᵷ㆜
㐿ᆎᣣ
ᵷ㆜
⚳ੌᣣ
ᵷ㆜
ᣣᢙ
᧻↰৻Ꮧ
㔤㐳㘃⎇ⓥᚲ
ࡐࠬ࠼ࠢ
ࡑ࡟࡯ࠪࠕ
ࠨࡃᄢቇ‫᧛࠙ࠞࠬޔ‬๟ㄝ࿾ၞ
H23.5.10
H23.7.10
62
㊄᫪ᦺሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ࡐࠬ࠼ࠢ
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯᫪ᨋ଻⼔඙
H23.6.7
H23.7.1
25
ㄞᄢ๺
㔤㐳㘃⎇ⓥᚲ
ഥᢎ
ࡌ࠻࠽ࡓ
ࡈࠛᄢቇ࠰ࡦ࠴ࡖ⥄ὼ଻⼔඙
H23.6.15
H23.7.21
37
ㄞᄢ๺
(਄Ბߣห৻ߩ⠪)
㔤㐳㘃⎇ⓥᚲ
ഥᢎ
ࠗࡦ࠼ࡀࠪࠕ
ࡏࠧ࡯࡞ㄘ⑼ᄢቇ‫ࡦ࡜࠳ࡦࠟࡄޔ‬
⥄ὼ଻⼔඙
H23.7.22
H23.8.20
30
᳓ญᄢテ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.17
H23.7.1
15
ዊᨋବ੺
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.17
H23.7.3
17
᧻Ꮉ޽߅޿
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
࠲ࡆࡦ㊁↢↢‛଻⼔඙
H23.6.20
H23.12.4
168
ਛ᧛⟤⍮ᄦ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ᜂᒰᢎຬ
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.20
H23.7.1
12
᧖ᶆ⑲᮸
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ᜂᒰᢎຬ
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.20
H23.7.1
12
૒‫ᧁޘ‬෹♿ሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.20
H23.7.1
12
Ỉᩙ⑲ᄥ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.20
H23.7.1
12
㘵↰ᕺℂሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.20
H23.7.1
12
ᬀ↰ᓀኈሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.20
H23.7.1
12
↰ፉ⍮ਯ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.20
H23.7.3
14
ỳ↰㘧㠽
㔤㐳㘃⎇ⓥᚲ
ୃ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠠ࠽ࡃ࠲ࡦࠟࡦᎹᵹၞ‫ࡃࡓ࠽࠳ޔ‬
࡟࡯ࠦࡦࠨ࡯ࡌ࡯࡚ࠫࡦࠛ࡝ࠕ
H23.6.22
H23.6.30
9
᧛ጊ⟤Ⓞ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ᜂᒰᢎຬ
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.24
H23.7.1
8
㤥㠽⧷ବ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.24
H23.7.1
8
㧙㧙
ศ↰ᒎ↢
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃ⽷࿅
H23.6.24
H23.7.1
8
ਭ਎Ớሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ࡐࠬ࠼ࠢ
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯⥄ὼ଻⼔඙‫ޔ‬
ࠨࡃᄢቇ
H23.6.28
H23.7.9
12
ዊ㒋⧎᪸
ㄘቇㇱ
ቇㇱቇ↢
ࡉ࡯࠲ࡦ
ࡉ࡯࠲ࡦ₺┙ᄢቇ
H23.7.5
H23.7.13
9
H23.7.15
H23.8.13
30
቟੗ᣧ♿
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
࠲ࠗ
ࠧ࡯࡞࠺ࡦ࠻࡜ࠗࠕࡦࠣ࡞ࠕࠫ
ࠕ࠱࠙⽷࿅
቟੗ᣧ♿
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
ࠗࡦ࠼
ࠗࡦ࠼⑼ቇᄢቇ
H23.8.14
H23.9.16
34
㔤㐳㘃⎇ⓥᚲ
ᜂᒰᢎຬ
࠲ࠗ
࠴ࡘ࡜ࡠࡦࠦ࡯ࡦᄢቇ
H23.9.5
H23.9.13
9
౎዆๺਽
ᢥቇㇱ
ቇㇱቇ↢
࠲ࠗ
࠴ࡘ࡜ࡠࡦࠦ࡯ࡦᄢቇ
H23.9.5
H23.9.13
9
᳓⼱ᒄ᮸
⚻ᷣቇㇱ
ቇㇱቇ↢
࠲ࠗ
࠴ࡘ࡜ࡠࡦࠦ࡯ࡦᄢቇ
H23.9.5
H23.9.13
9
⮮ᧄବᐔ
ㄘቇㇱ
ቇㇱቇ↢
࠲ࠗ
࠴ࡘ࡜ࡠࡦࠦ࡯ࡦᄢቇ
H23.9.5
H23.9.13
9
ዊᨋବ੺
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠞࡆ࡝࡯࠮ࡇࡠࠢ᫪ᨋ଻⼔඙
H23.9.6
H23.12.2
88
ᐘፉม㇢
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ᜂᒰᢎຬ
ࡑ࡟࡯ࠪࠕ
ࡑ࡝ࠕ࠙ࡌࠗࠬࡦ⥄ὼ଻⼔඙
H23.9.6
H23.9.19
14
H23.9.6
H23.9.13
8
(਄Ბߣห৻ߩ⠪)
ự↰ⓛ
ખỈિሶ
ℂቇㇱ
ቇㇱቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃᄢቇ‫ࡦࠬࠗࡌ࠙ࠕ࡝ࡑޔ‬
⥄ὼ଻⼔඙
᧻↰৻Ꮧ
㔤㐳㘃⎇ⓥᚲ
ࡐࠬ࠼ࠢ
ࡑ࡟࡯ࠪࠕ
ࠨࡃᄢቇ‫᧛࠙ࠞࠬޔ‬๟ㄝ࿾ၞ
H23.9.15
H23.11.15
62
෹᳗㓷Ꮖ
㔤㐳㘃⎇ⓥᚲ
ᜂᒰᢎຬ
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯᫪ᨋ଻⼔඙
H23.11.21
H23.11.29
9
ጊᧄ⌀਽
㔤㐳㘃⎇ⓥᚲ
ഥᢎ
ࡉ࡯࠲ࡦ
ࡉ࡯࠲ࡦ₺┙ᄢቇ
H23.11.22
H23.12.9
18
↰๺ఝሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻ቇ↢
ࡉ࡯࠲ࡦ
ࡉ࡯࠲ࡦ₺┙ᄢቇ
H23.11.22
H23.12.9
18
ᩙේᵗ੺
ℂቇㇱ
ቇㇱቇ↢
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯᫪ᨋ଻⼔඙
H23.11.22
H23.11.29
8
ựญఝ
කቇㇱ
ቇㇱቇ↢
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯᫪ᨋ଻⼔඙
H23.11.22
H23.11.29
8
⾐Ꮉߎߎࠈ
ᴺቇㇱ
ቇㇱቇ↢
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯᫪ᨋ଻⼔඙
H23.11.22
H23.11.29
8
ᩉỈ┨ᐢ
ℂቇㇱ
ቇㇱቇ↢
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯᫪ᨋ଻⼔඙
H23.11.22
H23.11.29
8
ጊᧄ⧷ታ
ᢎ⢒ቇㇱ
ቇㇱቇ↢
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯᫪ᨋ଻⼔඙
H23.11.22
H23.11.29
8
ਛጊᄢテ
Ꮏቇㇱ
ቇㇱቇ↢
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯᫪ᨋ଻⼔඙
H23.11.22
H23.11.29
8
ጟㇱጪੱ
ㄘቇㇱ
ቇㇱቇ↢
ࡉ࡯࠲ࡦ
ࡉ࡯࠲ࡦ₺┙ᄢቇ
H23.11.22
H23.12.9
18
㊄᫪ᦺሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ࡐࠬ࠼ࠢ
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯᫪ᨋ଻⼔඙
H23.12.2
H23.12.13
12
ᄢ⼱ᵗ੺
㔤㐳㘃⎇ⓥᚲ
ඳ჻ቇ↢
ࡑ࡟࡯ࠪࠕ
ࠨࡃᄢቇ‫᧛࠙ࠞࠬޔ‬๟ㄝ࿾ၞ
H24.1.20
H24.3.7
48
ਭ਎Ớሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ࡐࠬ࠼ࠢ
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯⥄ὼ଻⼔඙‫ࡃࠨޔ‬ᄢቇ
H24.2.12
H24.2.19
8
ਭ਎Ớሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ࡐࠬ࠼ࠢ
ࡑ࡟࡯ࠪࠕ
࠳࠽ࡓࡃ࡟࡯⥄ὼ଻⼔඙‫ࡃࠨޔ‬ᄢቇ
H24.3.5
H24.3.11
7
᧛਄ᄩᒎ
ㄘቇㇱ
ቇㇱቇ↢
ਛ࿖
ർ੩ᄢቇፏᏀ↢‛ᄙ᭽ᕈ⎇ⓥᚲ
H24.3.16
H24.3.25
10
ዊᴰᢘᒾ
ℂቇㇱ
ቇㇱቇ↢
ਛ࿖
ർ੩ᄢቇፏᏀ↢‛ᄙ᭽ᕈ⎇ⓥᚲ
H24.3.16
H24.3.25
10
㋈ᧁධ⟤
㔤㐳㘃⎇ⓥᚲ
ඳ჻ቇ↢
ਛ࿖
ർ੩ᄢቇፏᏀ↢‛ᄙ᭽ᕈ⎇ⓥᚲ
H24.3.17
H24.3.24
8
੹੗໪㓶
㔤㐳㘃⎇ⓥᚲ
ᜂᒰᢎຬ
ਛ࿖
ർ੩ᄢቇፏᏀ↢‛ᄙ᭽ᕈ⎇ⓥᚲ
H24.3.17
H24.3.24
8
㊄᫪ᦺሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ࡐࠬ࠼ࠢ
ࡑ࡟࡯ࠪࠕ
ࠬࠞ࠙᧛৻Ꮺ
H24.3.24
H24.3.29
6
ᳯፉବ
㔤㐳㘃⎇ⓥᚲ
ୃ჻ቇ↢
࠙ࠟࡦ࠳
ࠞ࡝ࡦ࠭᫪ᨋ଻⼔඙
H23.7.1
H23.7.21
21
H23.7.30
H23.8.29
31
ᄢ᦬੫ᄥ
ℂቇㇱ
ቇㇱቇ↢
ࠤ࠾ࠕ
࡜࠼ࠟ࡯ࠬᄢቇ‫ޔ‬
࠲࡯࠽Ꮉ᫪ᨋ଻⼔඙
ᓼጊᄹᏔሶ
㔤㐳㘃⎇ⓥᚲ
ୃ჻ቇ↢
ࠦࡦࠧ
↢ᘒ᫪ᨋ⎇ⓥᚲ
࡞ࠝ࡯ቇⴚ଻⼔඙
H23.8.5
H23.9.27
54
㘵↰ᕺℂሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻ቇ↢
࠲ࡦࠩ࠾ࠕ
࠲ࡦࠩ࠾ࠕ㊁↢േ‛⎇ⓥᚲ‫ޔ‬
࠙ࠟ࡜࿾ၞࡦࠣࠛ⺞ᩏ୯
H23.8.8
H23.12.21
136
ᰞᐸ㓁ሶ
㔤㐳㘃⎇ⓥᚲ
ୃ჻ቇ↢
࠲ࡦࠩ࠾ࠕ
ࠧࡦࡌ࿖┙౏࿦‫ޔ‬
࠮࡟ࡦࠥ࠹ࠖ࿖┙౏࿦
H23.8.17
H23.8.26
10
M Seres
㔤㐳㘃⎇ⓥᚲ
ࡐࠬ࠼ࠢ
࠲ࡦࠩ࠾ࠕ
ࠧࡦࡌ࿖┙౏࿦‫ޔ‬
࠮࡟ࡦࠥ࠹ࠖ࿖┙౏࿦
H23.8.17
H23.8.26
10
㥲⮮੝⍫
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ഥᢎ
࠲ࡦࠩ࠾ࠕ
ࠧࡦࡌ࿖┙౏࿦‫ޔ‬
࠮࡟ࡦࠥ࠹ࠖ࿖┙౏࿦
H23.8.17
H23.8.26
10
ᬀ↰ᗐ
㔤㐳㘃⎇ⓥᚲ
ୃ჻ቇ↢
࠲ࡦࠩ࠾ࠕ
ࠧࡦࡌ࿖┙౏࿦‫ޔ‬
࠮࡟ࡦࠥ࠹ࠖ࿖┙౏࿦
H23.8.17
H23.8.26
10
㧙㧙
ቲ↰৻ノ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ୃ჻ቇ↢
࠙ࠟࡦ࠳
ࡉ࠙ࠖࡦ࠺ࠖේ↢࿖┙౏࿦
H23.10.5
H23.10.18
14
ጊ᪸⵨⟤
㔤㐳㘃⎇ⓥᚲ
ඳ჻ቇ↢
ࠡ࠾ࠕ
ࡏ࠶࠰࠙ⅣႺ⎇ⓥᚲ
H23.12.10
H24.3.15
97
ᳯፉବ
㔤㐳㘃⎇ⓥᚲ
ୃ჻ቇ↢
࠙ࠟࡦ࠳
ࠞ࡝ࡦ࠭᫪ᨋ଻⼔඙
H24.1.4
H24.2.29
57
⷏ᳯੳᓼ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ࡐࠬ࠼ࠢ
࠲ࡦࠩ࠾ࠕ
ࡑࡂ࡟ጊ႙࿖┙౏࿦
H24.2.13
H24.3.29
46
AJJ
MacIntosh
㔤㐳㘃⎇ⓥᚲ
ࡐࠬ࠼ࠢ
ࡈ࡜ࡦࠬ
ࠬ࠻࡜ࠬࡉ࡯࡞ᄢቇ
H23.5.7
H23.5.29
23
Ỉ↰᥏ሶ
㔤㐳㘃⎇ⓥᚲ
ඳ჻ቇ↢
ࠬࠗࠬ
࠴ࡘ࡯࡝࠶ࡅᄢቇ
H23.5.9
H23.6.5
28
⍫㊁⥶
㔤㐳㘃⎇ⓥᚲ
ࡐࠬ࠼ࠢ
ࠝ࡯ࠬ࠻࡝ࠕ
࠙ࠖ࡯ࡦᄢቇ
H23.8.19
H23.8.20
2
⍫㊁⥶
㔤㐳㘃⎇ⓥᚲ
ࡐࠬ࠼ࠢ
࠼ࠗ࠷
ࡑ࠶ࠢࠬࡊ࡜ࡦࠢੱ㘃ㅴൻ⎇ⓥᚲ
H23.8.21
H23.9.9
20
(਄Ბߣห৻ߩ⠪)
㔤㐳㘃⎇ⓥᚲ
ࡐࠬ࠼ࠢ
ࠗࠡ࡝ࠬ
࡛࡯ࠢᄢቇ
H23.9.10
H23.10.31
52
ᯅᧄ⵨ሶ
㔤㐳㘃⎇ⓥᚲ
ࡐࠬ࠼ࠢ
ࠗࠡ࡝ࠬ
ᄢ⧷ඳ‛㙚㧘ࠛ࠺ࠖࡦࡃ࡜ᄢቇ
H23.8.30
H23.9.20
22
⷏ጟ૓৻㇢
㔤㐳㘃⎇ⓥᚲ
ඳ჻ቇ↢
ࠗࠡ࡝ࠬ
ᄢ⧷ඳ‛㙚
H23.9.4
H23.9.17
14
⷏ጟ૓৻㇢
(਄Ბߣห৻ߩ⠪)
㔤㐳㘃⎇ⓥᚲ
ඳ჻ቇ↢
࠼ࠗ࠷
ࡒࡘࡦࡋࡦᄢቇ
H23.9.18
H23.9.30
13
㜞੗ᱜᚑ
㔤㐳㘃⎇ⓥᚲ
ᜂᒰᢎຬ
ࠗࠡ࡝ࠬ
ᄢ⧷ඳ‛㙚
H23.9.4
H23.9.17
14
㜞੗ᱜᚑ
(਄Ბߣห৻ߩ⠪)
㔤㐳㘃⎇ⓥᚲ
ᜂᒰᢎຬ
࠼ࠗ࠷
ࡒࡘࡦࡋࡦᄢቇඳ‛㙚
H23.9.18
H23.9.24
7
㒙ㇱ⑲᣿
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
࠼ࠗ࠷
ࡌ࡞࡝ࡦേ‛࿦‫ޔ‬
ࡑ࠶ࠢࠬࡊ࡜ࡦࠢ⎇ⓥᚲ
H23.9.11
H23.9.24
14
C Martin
㔤㐳㘃⎇ⓥᚲ
ඳ჻ቇ↢
ࠕࡔ࡝ࠞ
ࠞ࡝ࡈࠜ࡞࠾ࠕᎿ⑼ᄢቇ
࡝ࡦࠞ࡯ࡦࡄ࡯ࠢേ‛࿦
H23.7.5
H23.8.17
44
⁚㊁ᢥᶈ
㔤㐳㘃⎇ⓥᚲ
ඳ჻ቇ↢
ࠕࡔ࡝ࠞ
ࠗࡦ࠺ࠖࠕ࠽ᄢቇ‫ޔ‬
ࠗࡦ࠺ࠖࠕ࠽ࡐ࡝ࠬേ‛࿦
H23.7.25
H23.8.5
12
ዊୖආବ
㔤㐳㘃⎇ⓥᚲ
ࡐࠬ࠼ࠢ
ࠕࡔ࡝ࠞ
╙ 10 ࿁ࠛࡦ࡝࠶࠴ࡔࡦ࠻ળ⼏‫ޔ‬
ࠝ࡟ࠧࡦേ‛࿦
H23.8.12
H23.8.27
16
(਄Ბߣห৻ߩ⠪)
⍫㊁⥶
⑧භ㓷ผ
㔤㐳㘃⎇ⓥᚲ
ඳ჻ቇ↢
ࠕࡔ࡝ࠞ
ർ☨␹⚻ቇળ
H23.11.11
H23.11.18
8
ች࿾㊀ᒄ
㔤㐳㘃⎇ⓥᚲ
ᜂᒰᢎຬ
ࠕࡔ࡝ࠞ
ർ☨␹⚻ቇળ
H23.11.11
H23.11.18
8
㡨ᳰ⩿଻
㔤㐳㘃⎇ⓥᚲ
ඳ჻ቇ↢
ࠕࡔ࡝ࠞ
╙ 41 ࿁ർ☨␹⚻⑼ቇળ
H23.11.12
H23.11.18
7
૒‫ᧁޘ‬෹♿ሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
ࡉ࡜ࠫ࡞
࿖┙ࠕࡑ࠱ࡦ⎇ⓥᚲ‫ޔ‬
ࡑࡒ࡜࠙ࠕ⥄ὼ଻⼔඙
H24.1.21
H24.2.28
39
૒‫ᧁޘ‬෹♿ሶ
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ඳ჻ቇ↢
࠴࡝
࠴࡝࿖┙⑼ቇ⎇ⓥᚲ
H24.2.29
H24.3.30
31
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯
ഥᢎ
ࡉ࡜ࠫ࡞
ࡃࡆ࠻ࡦࠟḧ৻Ꮺ
H24.2.24
H24.3.10
16
(਄Ბߣห৻ߩ⠪)
᫪㒋ආㅢ
ᐔᚑ 23 ᐕᐲߩว⸘ߪએ਄ߩ 74 ઙߢ޽ࠆ‫ޕ‬ᵷ㆜ᣣᢙߪ 1,880 ᣣߦߥߞߚ‫✚⾌⚻ޕ‬㗵ߪ‫ޔ‬25,860,424 ౞ߛߞߚ‫ޕ‬
㧔ᢥ⽿㧦᧻ᴛື㇢࡮਄၂ᵏᶈ㧕
3. ᦨవ┵⎇ⓥၮ⋚੐ᬺ㧦ᔃߩవ┵⎇ⓥߩߚ߼ߩㅪ៤᜚ὐ㧔WISH㧕᭴▽
ᢥㇱ⑼ቇ⋭ߩᦨవ┵⎇ⓥၮ⋚੐ᬺߩ⹺ቯࠍ੩ㇺᄢቇ߇ฃߌߚ‫ޕ‬੐ᬺฬߪ‫ޟޔ‬ᔃߩవ┵⎇ⓥߩߚ߼ߩㅪ៤᜚ὐ
㧔WISH㧕᭴▽㧕
‫⇛ޕࠆ޽ߢޠ‬⒓ࠍ‫ޟ‬ᔃߩవ┵⎇ⓥ WISH ੐ᬺ‫ޕࠆߔߣޠ‬੐ᬺታᣉᯏ㑐ߪ੩ㇺᄢቇ㧔ᔃℂቇ࡮⹺⍮
⑼ቇ╬ࠍታᣉߔࠆᄢቇ߿⎇ⓥᯏ㑐ߣߩㅪ៤‫ޔ‬੐ᬺઍ⴫⠪㧦᧻ᴛື㇢㧕
‫ޕ‬੐ᬺ᭎ⷐߪ‫ߩ╬࡯ࠫࡦࡄࡦ࠴ޔ࠻ࡅޔ‬Ყセ
⹺⍮ታ㛎╬ࠍ߅ߎߥ߁ࡀ࠶࠻ࡢ࡯ࠢ⎇ⓥ᜚ὐࠍᢛ஻ߒ‫ޔ‬ᔃℂቇ‫⹺ޔ‬⍮⑼ቇ‫ޔ‬⣖⑼ቇ߿␠ળ⑼ቇߩಽ㊁ࠍ⿥߃ߚቇ
㓙⎇ⓥࠍⴕ޿‫ઁޔ‬⠪ߣߩ⋧੕૞↪ߦࠃࠆᔃߩߪߚࠄ߈ࠍ⸃᣿ߔࠆߚ߼ߩవ┵⎇ⓥࠍផㅴߔࠆ‫ޕ‬ታᣉᦼ㑆 3 ᐕ㑆‫ޔ‬
ᐔᚑ 22-24 ᐕᐲ‫⵬ޕ‬ഥ㊄㗵ߪ‫ޔ‬ᐔᚑ 22 ᐕᐲ 5 ం౞‫ޔ‬ᐔᚑ 23 ᐕᐲ 5 ం౞‫ޔ‬ᐔᚑ 24 ᐕᐲ 4 ం౞‫ޔ‬ว⸘ 14 ం౞‫ޕ‬
ᢥㇱ⑼ቇ⋭ߢߪ‫ᦨޔ‬వ┵⎇ⓥ㐿⊒ᚢ⇛⊛ᒝൻ੐ᬺㆇ↪ၮᧄᣇ㊎㧔ᐔᚑ 22 ᐕ 4 ᦬ 27 ᣣ ✚ว⑼ቇᛛⴚળ⼏᳿ቯ㧕
ߦၮߠ߈‫ޔ‬ᐔᚑ 22 ᐕᐲߦഃ⸳ߐࠇߚᦨవ┵⎇ⓥ㐿⊒ᚢ⇛⊛ᒝൻ⾌⵬ഥ㊄㧔400 ం౞㧕ߩ৻ㇱࠍᵴ↪ߒߚ‫ᦨޟ‬వ┵
⎇ⓥၮ⋚੐ᬺ‫ޔߡ޿ߟߦޠ‬ᐔᚑ 22 ᐕᐲߩ⵬ഥኻ⽎੐ᬺߣߒߡ 14 ઙߩ੐ᬺࠍㆬቯߒߚ‫ޕ‬ᔃߩవ┵⎇ⓥ WISH ੐ᬺ
ߪߘߩ߭ߣߟߢ޽ࠆ‫ޕ‬
ᐔᚑ 22 ᐕᐲߩ⚻⾌ߪ‫ޔ‬੤ઃ᳿ቯ߇ㆃࠇߚߚ߼‫ߡߴߔ߷߶ޔ‬ᐔᚑ 23 ᐕᐲߦ➅⿧ߒߚ‫ޕ‬ᐔᚑ 23 ᐕᐲߪ‫ޔ‬ታ⾰⊛ߦ
੐ᬺ߇ㅴዷߒߚᐕߢ޽ࠆ‫ᧄޕ‬੐ᬺߦࠃࠅ‫ޔ‬ԘᲧセ⹺⍮⑼ቇታ㛎⸳஻㧔ᄢဳࠤ࡯ࠫ㧕
‫ޔ‬ԙfMRI‫ ߩޔ‬2 ߟߩ⸳஻ࠍ੩ᄢ
ߦᢛ஻ߒߚ‫ౕޕ‬૕⊛ߦߪ‫ޔ‬Ყセ⹺⍮⑼ቇታ㛎ߩߚ߼ߩᄢဳࠤ࡯ࠫࠍ›ጊߩ㔤㐳㘃⎇ⓥᚲߦ 1 บ‫ޔ‬ᾢᧄࠨࡦࠢ࠴ࡘ
㧙㧙
ࠕ࡝ߦ 1 บᢛ஻ߒߚ‫ߦ↱⥄ޔߡߞࠃߦࠇߎޕ‬㔌ว㓸ᢔߔࠆ⟲ࠇో૕ࠍ⎇ⓥኻ⽎ߣߒߟߟ‫ޔ‬㗻⹺⸽ࠪࠬ࠹ࡓߩዉ౉
ߢ‫ޔ‬1 ੱ߭ߣࠅࠍ୘૕⼂೎ߒߚ਄ߢߩ␠ળ੤ᷤታ㛎߇น⢻ߦߥߞߚ‫ޕ‬෩ኒߥᔃℂ‛ℂቇ⊛ታ㛎ࠍน⢻ߦߒߟߟ‫ޔ‬2
୘૕႐㕙‫ⶄߪߦࠄߐޔ‬ᢙ୘૕หᤨߩ␠ળ੤ᷤߩታ㛎⊛⸃ᨆࠍน⢻ߦߒߚ‫ߦࠄߐޕ‬㆙㓒࿾߆ࠄߩᠲ૞ߢታ㛎߇ߢ߈
ࠆࠃ߁ߦḰ஻ࠍߔߔ߼ߚ‫ޕ‬
‫ ⑼࠻ࡅߩࡏࡁࡏ࡮࡯ࠫࡦࡄࡦ࠴࡮࠻ࡅޟ‬3 ⒳ߩᲧセ⹺⍮⑼ቇታ㛎‫ࠍޠ‬ታ⃻ߔࠆ╙৻ᱠࠍ
〯ߺ಴ߖߚߣ޿߃ࠆ‫ੱ߅ߥޕ‬㑆ࠍ⎇ⓥኻ⽎ߦߒߚ fMRI ⸳஻ߦߟ޿ߡߪ‫ޔ‬ᐔᚑ 23 ᐕᐲᧃߦ੩ᄢᧄㇱ᭴ౝ∛㒮⷏࿾
඙ߦ‫␠ࠬࡦࡔ࡯ࠪޔ‬㧔3 ࠹ࠬ࡜㧕ࠍዉ౉ߒ‫⎇᧪ᧂߩࠈߎߎޔ‬ⓥ࠮ࡦ࠲࡯ߩᚲ᝿ߣߒߚ‫ޔ߅ߥޕ‬ᔃߩవ┵⎇ⓥ WISH
੐ᬺࠍታᣉߔࠆਛᩭ⚵❱ߣߒߡ‫ޔ‬੩ㇺᄢቇߦ‫ޟ‬ᔃߩవ┵⎇ⓥ࡙࠾࠶࠻‫߇ޠ‬ᐔᚑ 22 ᐕ 12 ᦬ 12 ᣣߦᱜᑼߦ⊒⿷ߒߚ‫ޕ‬
ᔃℂቇ࡮⹺⍮⑼ቇࠍᮡ᭙ߔࠆ੩ᄢߩ 10 ㇱዪ 64 ฬߩᢎຬߩෳടߔࠆ࡙࠾࠶࠻ߢ޽ࠆ‫ޔߪ⚦⹦ޕ‬એਅߩ HP ࠍෳᾖߐ
ࠇߚ޿‫ޕ‬http://www.kokoro-kyoto.org ߥ߅ೋઍߩ࡙࠾࠶࠻੐ോขࠅᛒ޿ㇱዪ߇㔤㐳㘃⎇ⓥᚲߣߥࠅ‫ޔ‬ᐔᚑ 22-23 ᐕ
ᐲߩೋઍ࡙࠾࠶࠻㐳ࠍ᧻ᴛື㇢߇ߟߣ߼ߚ‫ޕ‬
㧔ᢥ⽿㧦᧻ᴛື㇢㧕
4. ࠕࠫࠕ࡮ࠕࡈ࡝ࠞቇⴚၮ⋚ᒻᚑ੐ᬺ㧦ࡅ࠻⑼㘃ੱ₎ߩⅣႺㆡᔕᯏ᭴ߩᲧセ⎇ⓥ
੐ᬺߩ⋡⊛
࠴ࡦࡄࡦࠫ࡯㧔Pan㧕ዻߩ࠴ࡦࡄࡦࠫ࡯ߣࡏࡁࡏߪ‫ߦ࠻ࡅ߽ߣߞ߽ߦ⊛⛔♽ޔ‬ㄭ޿㘃ੱ₎ߢ޽ࠅ‫ޔ‬ᚒ‫ߣߣ࠻ࡅޘ‬
߽ߦࡅ࠻⑼ࠍ᭴ᚑߔࠆ‫ޕ‬ᓐࠄߪࠕࡈ࡝ࠞߩ⿒㆏ࠍਛᔃߦ‫ޔ‬ᾲᏪᄙ㔎ᨋ߆ࠄࠨࡃࡦ࠽࠙࠶࠼࡜ࡦ࠼ߦ޿ߚࠆᄙ᭽ߥⅣ
Ⴚߦ↢ᕷߒߡ߅ࠅ‫ߩࠇߙࠇߘޔ‬࿾ၞߢ᭽‫␠ߥޘ‬ળ᭴ㅧ߿㆏ౕ૶↪ࠍ⊒㆐ߐߖߡ㘩‛ⅣႺߣߘߩᐕᄌേ࡮ቄ▵ᄌേߦ
ኻᔕߒߡ޿ࠆ‫ߩࠄࠇߎޕ‬⒳ߩⅣႺㆡᔕᚢ⇛ߩㅴൻࠍ࿾ၞ㑆ߩᲧセࠍㅢߓߡ⸃᣿ߔࠆߎߣߪ‫ޔ‬㘃ੱ₎ߩㅴൻߩ⸃᣿ߦ
ߣߤ߹ࠄߕ‫ޔ‬Pan ዻߣߩ౒ㅢ␲వ߆ࠄᵷ↢ߒߡࠃࠅੇ῎ߒߚ࿾ၞߢ↢߈ᱷࠅ‫ࠄ߆ߎߘޔ‬਎⇇ߩ޽ࠄࠁࠆⅣႺߦㅴ಴
ߒߚࡅ࠻ߩㅴൻߩ಴⊒ὐࠍតࠆ਄ߢ߽‫ߡ߼ࠊ߈ޔ‬㊀ⷐߢ޽ࠆ‫ޕ‬
੩ㇺᄢቇࠍਛᔃߣߒߡ⊒ዷߒߡ߈ߚ㔤㐳㘃ቇߪ‫ޔ‬㘃ੱ₎ߩㅴൻߩ⎇ⓥࠍㅢߒߡࡅ࠻ߩ࡞࡯࠷ࠍតࠆߎߣࠍ߭ߣߟ
ߩᄢ߈ߥ⋡ᮡߣߒߡ߈ߚ‫ޔ߼ߚߩߘޕ‬᭽‫ߥޘ‬㘃ੱ₎⒳ࠍ㐳ᦼߦࠊߚߞߡ⺞ᩏߔࠆ⺞ᩏ࿾ࠍࠕࡈ࡝ࠞߣࠕࠫࠕߦᄙᢙ
߽ߜ‫߇ࠇߎޔ‬ᣣᧄߩ㔤㐳㘃ቇߩ਎⇇ߦ⹶ࠇࠆ․⦡ߣߥߞߡ޿ࠆ‫ߦߊߣޕ‬੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲߪ‫ߩߘޔ‬ᢎຬ߇ઍ⴫
ࠍോ߼ࠆ Pan ዻߩ㐳ᦼ⺞ᩏ࿾ࠍࠡ࠾ࠕ౒๺࿖ߩࡏ࠶࠰࠙‫ࠧࡦࠦޔ‬᳃ਥ౒๺࿖ߩࡢࡦࡃ‫౒࠳ࡦࠟ࠙ޔ‬๺࿖ߩࠞ࡝ࡦ࠭
ߣ 3 ࠞᚲ߽߽ߟ‫⿒ߪࠄࠇߎޕ‬㆏ᴪ޿ߦ⷏ࠕࡈ࡝ࠞ‫ޔ‬ਛᄩࠕࡈ࡝ࠞ‫ࠆߥ⇣ߩࠞ࡝ࡈࠕ᧲ޔ‬ⅣႺࠍࠞࡃ࡯ߒߡ߅ࠅ‫⋧ޔ‬
ᚻ࿖ߩ᜚ὐᯏ㑐ߣߩ㐳ᐕߦࠊߚࠆ⎇ⓥදജࠍㅢߒߡ᭽‫⎇ߥޘ‬ⓥᚑᨐࠍ޽ߍߡ߈ߡ޿ࠆ‫ޕ‬
ߎߩ⎇ⓥ੤ᵹߩ⋡ᮡߪ‫ޔ‬㔤㐳㘃⎇ⓥᚲߩᢎຬߣ⋧ᚻ࿖᜚ὐᯏ㑐ߣߩ⎇ⓥදജࠍࠃࠅᒝ࿕ߥ߽ߩߦߔࠆߛߌߢߥߊ‫ޔ‬
3 ࿖ߩ᜚ὐᯏ㑐ห჻ߩ⎇ⓥ੤ᵹ߽⊒ዷߐߖ‫ޔ‬Pan ዻߩ↢ᘒቇ⊛࡮ㅴൻቇ⊛ߥ⎇ⓥߩ਎⇇⊛ߥᩭࠍᒻᚑߔࠆߎߣߦ޽
ࠆ‫ޕ‬
੐ᬺ⸘↹ߩ᭎ⷐ
ᧄ⸘↹ߪ‫ޔ‬ਥߣߒߡએਅߩ 2 ߟߩ੐ᬺ߆ࠄߥࠆ‫ޕ‬
౒ห⎇ⓥ࡮⎇ⓥ⠪੤ᵹ
㔤㐳㘃⎇ⓥᚲߣ⋧ᚻ࿖᜚ὐᯏ㑐ߩ㑆ߢߪ‫ࠧࡦࠦޔ‬᳃ਥ౒๺࿖ߢ 1973 ᐕ߆ࠄ‫౒ࠕ࠾ࠡޔ‬๺࿖ߢ 1975 ᐕ߆ࠄ‫࠙ޔ‬
ࠟࡦ࠳౒๺࿖ߢ 1996 ᐕ߆ࠄ౒ห⎇ⓥࠍⴕߞߡ߈ߡ޿ࠆ‫ߩࠇߙࠇߘޕ‬࿖ߩ⎇ⓥ⠪ߣ౒ห⎇ⓥࠍⴕ߁ߎߣߢ‫ޔ‬᡽ᴦ
ᖱ൓ߩਇ቟ቯߥߣ߈ߢ߽㐳ᦼߦࠊߚࠆ⛮⛯⺞ᩏ߇น⢻ߢ‫ߦߢ߹ࠇߎޔ‬ᄢ߈ߥ⎇ⓥᚑᨐࠍ਄ߍߡ߈ߚ‫ߩࠇߙࠇߘޕ‬
⎇ⓥߪ‫╬⾌⎇⑼ޔ‬୘೎ߩ⎇ⓥ⾌ߦࠃߞߡᡰ߃ࠄࠇߡ޿ࠆ߇‫ޔߪߢ↹⸘ᧄޔ‬㔤㐳㘃⎇ⓥᚲߣ⋧ᚻ࿖᜚ὐᯏ㑐ߣߩ⎇
ⓥ⠪ߩ⋧੕⸰໧ࠍታ⃻ߔࠆߎߣߦࠃߞߡ౒ห⎇ⓥࠍ౞Ṗߦㅴ߼‫ߟ߆ޔ‬ᯏ㑐㑆ߩ㑐ଥࠍᒝൻߔࠆߎߣࠍ⋡ᜰߔ‫ౕޕ‬
૕⊛ߦߪ‫ޔ‬ᣣᧄ஥⎇ⓥ⠪߇ฦ᜚ὐᯏ㑐ࠍ⸰໧ߒߡ⎇ⓥᣇᴺ߿࠺࡯࠲ߩಣℂᴺߩᜰዉࠍⴕ߁ߣߣ߽ߦ‫ޔ‬᜚ὐᯏ㑐ߩ
⧯ᚻ⎇ⓥ⠪ࠍ㔤㐳㘃⎇ⓥᚲߦ᜗⡜ߒ‫⎇ޔ‬ⓥᣇᴺ╬ߦߟ޿ߡߩ࠻࡟࡯࠾ࡦࠣࠍⴕ߁‫ߩࠄࠇߎߚ߹ޕ‬੤ᵹߦၮߠ޿ߡ‫ޔ‬
ᣣᧄࠍ฽߼ߚ 4 ࿖ߩ⎇ⓥ⠪ߢ౒ห⎇ⓥ࡮Ყセ⎇ⓥࠍ┙᩺ߒ‫⑼ޔ‬ቇ⎇ⓥ⾌⵬ഥ㊄ߥߤࠍ↪޿ߡታᣉߔࠆ‫ޕ‬
ԙ ࠮ࡒ࠽࡯╬ߩቇⴚળว
ᐔᚑ 21 ᐕᐲߦߪ‫⋧ޔ‬ᚻ࿖ 3 ࡩ࿖ߢ‫ޔ‬Pan ዻߩ⎇ⓥߦ㑐ߔࠆߎࠇ߹ߢߩᚑᨐߩ⊒⴫ߣ੹ᓟߩ⺖㗴ߦ㑐ߔࠆ⼏⺰
ࠍⴕ߁࠮ࡒ࠽࡯ࠍ㐿௅ߔࠆ‫ޕ‬ᣣᧄ஥߆ࠄߪ⧯ᚻ⎇ⓥ⠪ࠍᵷ㆜ߒ‫⋧ޔ‬ᚻ࿖ᯏ㑐ߩ⧯ᚻ⎇ⓥ⠪ߣߩ੤ᵹࠍᷓ߼ࠆ‫ޕ‬
ᐔᚑ 22 ᐕᐲߦߪ‫ޔ‬੩ㇺᄢቇ߇ਥ௅⠪ߣߥߞߡ‫ޔ‬ᣣᧄߢ࿖㓙㔤㐳㘃ቇળߩᄢળ߇㐿௅ߐࠇࠆ‫ߩߎޕ‬ቇળߢ‫ࠕޔ‬
ࡈ࡝ࠞߩ᧲ㇱ࡮ਛㇱ࡮⷏ㇱߩ Pan ዻߩⅣႺㆡᔕᚢ⇛ߩᲧセࠍ࠹࡯ࡑߣߒߚࠪࡦࡐࠫ࠙ࡓࠍ㐿௅ߔࠆ‫ߩࠇߙࠇߘޕ‬
⺞ᩏ࿾ߢߩ⎇ⓥታ❣ࠍᣣᧄ஥ෳട⎇ⓥ⠪ߣ⋧ᚻᯏ㑐ߩ⎇ⓥ⠪߇⊒⴫ߔࠆߎߣߦࠃࠅ‫ޔ‬㔤㐳㘃⎇ⓥᚲߣࠕࡈ࡝ࠞߩ
᜚ὐᯏ㑐ߩ⎇ⓥࡀ࠶࠻ࡢ࡯ࠢߩ߽ߟน⢻ᕈࠍ਎⇇ߦ߻ߌߡࠕࡇ࡯࡞ߔࠆ‫ޕ‬
ᐔᚑ 23 ᐕᐲߦߪ‫ޔ‬ฦ⺞ᩏ࿾ߦ߅ߌࠆ 3 ᐕ㑆ߩ౒ห⎇ⓥߣ‫⺞ޔ‬ᩏ࿾㑆ߩᲧセ⎇ⓥߩᚑᨐࠍ⊒⴫ߔࠆࠪࡦࡐࠫ࠙
ࡓࠍ‫ࠧࡦࠦޔ‬᳃ਥ౒๺࿖ߢ㐿௅ߔࠆ‫⎇ߩࠞ࡝ࡈࠕޕ‬ⓥ⠪߇࿖ߩᨒࠍ⿥߃ߡ㓸߹ࠆᯏળߪ‫ߡ߼ࠊ߈ޔ‬㒢ࠄࠇߡ޿ࠆ‫ޕ‬
ߎߩࠃ߁ߥ႐ࠍᜬߟߎߣߦࠃࠅ‫ޔ‬ฦᯏ㑐ߩ㑆ߩ㑐ଥࠍᒝ߼‫ޔ‬Ყセ⎇ⓥߩ⊒ዷߩ␆ࠍ▽ߊ‫ޕ‬
Ԙ
ᐔᚑ 23 ᐕᐲߩ⎇ⓥ੤ᵹᚑᨐ
⎇ⓥදജ૕೙ߩ᭴▽⁁ᴫ
㧙㧙
ᧄᐕᐲߪ‫ࠧࡦࠦޔߡߒߣࠅߊߊ߼✦ߩ࠻ࠢࠚࠫࡠࡊޔ‬᳃ਥ౒๺࿖ߩ᜚ὐᯏ㑐ߢ޽ࠆ↢ᘒ᫪ᨋ⎇ⓥᚲߦ‫ࠞ࡝ࡈࠕޔ‬
3 ࿖ߣᣣᧄߩ⎇ⓥ⠪߇㓸߹ߞߡ࠮ࡒ࠽࡯ࠍ㐿௅ߒߚ‫ࠆࠃߦ⾌ઁޕ‬ෳട߽฽߼‫ޔ‬ᒰೋ੍ቯߒߡ޿ߚⷙᮨࠍᄢ߈ߊ਄࿁
ࠆ 26 ฬߩෳട⎇ⓥ⠪߇㓸߹ࠆߎߣߦߥࠅ‫ޔ‬࿖㓙ࠪࡦࡐࠫ࠙ࡓߣߒߡߩ㐿௅ߣߥߞߚ‫ᧄޔߪߢࡓ࠙ࠫࡐࡦࠪߩߎޕ‬
ࡊࡠࠫࠚࠢ࠻ߢㅴ߼ߡ߈ߚ⎇ⓥ߿ߘࠇߙࠇߩ᜚ὐᯏ㑐ߢㅴ߼ߡ߈ߚ㑐ㅪ⎇ⓥߩ⎇ⓥ⊒⴫ࠍⴕߞߚ‫ޕ‬ᣣᧄ߆ࠄᄙᢙߩ
⎇ⓥ⠪߇ෳടߒߚߎߣ߽᱑ㄫߐࠇߚ߇‫⎇ߩࠞ࡝ࡈࠕޔ‬ⓥ⠪߇ઁߩ⎇ⓥᚲߦ㓸߹ߞߡࡢ࡯࡚ࠢࠪ࠶ࡊࠍ㐿ߊߣ޿߁ߎ
ߣ߇ߎࠇ߹ߢߦߪ޽߹ࠅߥߊ‫ޔ‬ᄢ߈ߥ⋓ࠅ਄߇ࠅࠍ⷗ߖߚ‫ޕ‬
ࠪࡦࡐࠫ࠙ࡓߩ✦߼ߊߊࠅߩ⸛⼏ߢߪ‫ࠞ࡝ࡈࠕޔ‬஥ෳട⠪߆ࠄ‫ޔ‬㔤㐳㘃⎇ⓥߩಽ㊁ߢᣣᧄߣߩㅪ៤ࠍ੹ᓟߐࠄߦ
ᒝൻߔࠆߎߣ‫ࠞ࡝ࡈࠕޔ‬㔤㐳㘃ቇળߩ⸳┙ߦะߌߡߐࠄߦදജ૕೙ࠍᒝൻߔࠆߎߣ߇ឭ⸒ߐࠇߚ‫ߥ߁ࠃߩߎޕ‬ឭ⸒
߇ߥߐࠇߚߎߣ߆ࠄ߽‫ޔ‬ᣣᧄߣࠕࡈ࡝ࠞ 3 ࿖ࠍ⚿߱ࡀ࠶࠻ࡢ࡯ࠢဳߩ⎇ⓥදജ૕೙ߩ᭴▽ߣ޿߁ೋᦼߩ⋡⊛߇චಽ
ߦ㆐ᚑߐࠇߚߎߣ߇ࠊ߆ࠆ‫ޕ‬
ቇⴚ㕙ߩᚑᨐ
ቇⴚ㕙ߢߪ‫ޔ‬3 ᐕ㑆ߩ⎇ⓥߢⴕߥߞߡ߈ߚ㘃ੱ₎ߩⅣႺㆡᔕᯏ᭴ߦߟ޿ߡߩᲧセ⎇ⓥߩᚑᨐߩ߶߆‫ᧄޔ‬੐ᬺߦෳ
ടߔࠆ⎇ⓥ⠪ߩⴕߞߡ޿ࠆߐ߹ߑ߹ߥ⎇ⓥߩᚑᨐ߇਄⸥ߩࡢ࡯࡚ࠢࠪ࠶ࡊߢ⊒⴫ߐࠇ‫ޔ‬ᾲᔃߥ⸛⺰߇੤ࠊߐࠇߚ‫ޕ‬
ߣߊߦ‫ࠞ࡝ࡈࠕޔ‬஥⎇ⓥ⠪ߩ૗ੱ߆߆ࠄ‫⊒ߦ⥄⁛ޔ‬᩺ߒߚ⎇ⓥߩ⊒⴫߇ߥߐࠇߚߎߣߪ‫ᧄޔ‬੐ᬺߩᄢ߈ߥᚑᨐߛߞ
ߚߣ޿߃ࠆ‫ᧄޔߦߊߣޕ‬੐ᬺߢ 2010 ᐕᐲߦ੩ㇺᄢቇඳ჻ภࠍขᓧߒߚ Mulavwa ඳ჻ߪ‫↢ޔ‬ᘒ᫪ᨋ⎇ⓥᚲߩ⎇ⓥ⠪
ߩ⁛┙ߒߚ⎇ⓥ⠪ߣߒߡߩ⢒ᚑߦജࠍ౉ࠇߡ߅ࠅ‫ᧄޔ‬੐ᬺߩᚑഞߦᄢ߈ߊ⽸₂ߒߡ޿ࠆ‫ޕ‬
ᬺ❣㕙ߢ߽‫ޔ‬ᩏ⺒ߩ޽ࠆቇⴚ㔀⹹߅ࠃ߮ᦠ☋ߦ 6 ᧄߩ⺰ᢥࠍ⊒⴫ߔࠆߎߣ߇ߢ߈ߚ‫ޔࠅࠃߦࠇߎޕ‬㘃ੱ₎ߩⅣႺ
ㆡᔕᯏ᭴ߦ㑐ߔࠆℂ⸃ࠍᄢ߈ߊㅴ߼ࠆߎߣ߇ߢ߈ߚ‫ޕ‬
⧯ᚻ⎇ⓥ⠪㙃ᚑ
਄⸥ߩࠪࡦࡐࠫ࠙ࡓߦෳടߒߚෳട⎇ⓥ⠪ߩ߁ߜ‫⚂ޔ‬ඨᢙ߇⧯ᚻߩ⎇ⓥ⠪ߢ޽ߞߚ‫ޔߚ߹ޕ‬150 ੱߦ߽ߩ߷ࠆ৻
⥸ෳട⠪ߦ߽ᄙᢙߩ⧯ᚻ⎇ⓥ⠪߇฽߹ࠇߡ߅ࠅ‫ޔ‬ᣣᧄ߅ࠃ߮ࠕࡈ࡝ࠞߩ⧯ᚻ⎇ⓥ⠪ߩⷐ⺧ߦߪᄢ߈ߊ⽸₂ߒߚ‫߹ޕ‬
ߚ‫ᧄޔ‬ᐕᐲ߽ᒁ߈⛯߈ⴕߞߚࠦࡦࠧ᳃ਥ౒๺࿖‫౒ࠕ࠾ࠡޔ‬๺࿖‫౒࠳ࡦࠟ࠙ޔ‬๺࿖ߢߩදห⎇ⓥߦ߽ᄙߊߩ⧯ᚻ⎇ⓥ
⠪߇ෳടߒ‫⎇ޔ‬ⓥ㕙ߢߩᛛⴚ߿⊒ᗐജࠍ㙃ߞߚ‫ޕ‬
␠ળ⽸₂
ࠕࡈ࡝ࠞᄢ㒽ߪ‫ޔ‬㔤㐳㘃ߩ↢ᕷߩਛᔃ࿾ߢ޽ࠅ‫ߩ↢⺀ߩ࠻ࡅޔ‬႐ߢ߽޽ࠆ‫ޔߪߢࠞ࡝ࡈࠕߩߘߒ߆ߒޕ‬㔤㐳㘃ቇ
߇චಽߦ⊒㆐ߒߡ޿ࠆߣߪ޿߃ߕ‫ޔ‬2006 ᐕߦ࠙ࠟࡦ࠳ߢ㐿௅ߐࠇߚ࿖㓙㔤㐳㘃ቇળߩቇⴚᄢળ߇ࠕࡈ࡝ࠞᄢ㒽ߢ
㐿௅ߐࠇߚߪߓ߼ߡߩᄢળߢ޽ߞߚ‫☨ޕ‬࿖‫ޔ‬᰷Ꮊߩᄙߊߩ࿖‫߮ࠃ߅ޔ‬ᣣᧄߦ޽ࠆࠃ߁ߥ࿖ߏߣߩ㔤㐳㘃ቇળߪࠕࡈ
࡝ࠞߦߪߥߊ‫ߩੱࠞ࡝ࡈࠕޔ‬㔤㐳㘃⎇ⓥ⠪ߪ‫ޔ‬ᄙߊߩ႐ว᰷☨߅ࠃ߮ᣣᧄߩ⎇ⓥ⠪ߩ౒ห⎇ⓥ⠪ߣߒߡᚻવ޿ࠍߔ
ࠆߣ޿߁┙႐ߦ↞ࠎߓߡ߈ߚ‫ޕ‬
ߘ߁޿ߞߚࠕࡈ࡝ࠞ߆ࠄ‫ࠞ࡝ࡈࠕޔ‬㔤㐳㘃ቇળࠍ⸳┙ߒߚ޿ߣ޿߁ᯏㆇ߇㜞߹ߞߡ߈ߚߩߪ‫ᧄޔ‬੐ᬺߩᄢ߈ߥᚑ
ᨐߢ޽ࠅ‫␠ߩࠞ࡝ࡈࠕޔ‬ળߩቇⴚ㕙ߢߩᚑᾫߦᄢ߈ߊ⽸₂ߔࠆ߽ߩߢ޽ࠆ‫ޔߚ߹ޕ‬ᣣᧄߩࠗ࠾ࠪࠕ࠹ࠖࡉߢࠕࡈ࡝
ࠞ㔤㐳㘃ቇળ߇⸳┙ߐࠇࠆߎߣߦߥࠇ߫‫ޔ‬਎⇇ߦ߅ߌࠆᣣᧄߩቇⴚ⊛࿾૏ߩะ਄ߦ߽ᄢ߈ߊ⽸₂ߔࠆ‫ޕ‬
ߥ߅‫ᦨߩࡓ࠙ࠫࡐࡦࠪޔ‬ᓟߦ‫ࠞ࡝ࡈࠕޔ‬㔤㐳㘃ቇળߩ⸳┙ߦ㆏╭ࠍߟߌࠆߎߣࠍ⋡ᮡߣߒߡࠕࠫࠕ࡮ࠕࡈ࡝ࠞቇ
ⴚၮ⋚ᒻᚑ੐ᬺߦౣᐲᔕ൐ߒߡ߶ߒ޿ߣ޿߁Ꮧᦸ߇ࠕࡈ࡝ࠞ஥᜚ὐᯏ㑐߆ࠄ⥄ਥ⊛ߦ಴ߐࠇߚ‫࠙ࠫࡐࡦࠪߢߎߘޕ‬
ࡓߦᒁ߈⛯޿ߡᰴᦼ⸘↹ߩౕ૕᩺ߦߟ޿ߡ⋧⺣ߒߡ↳⺧ߒߚߣߎࠈ‫ޔ‬ណᛯߒߡ޿ߚߛߊߎߣ߇ߢ߈ߚ‫ޕ‬ᰴᦼߩ 3 ᐕ
㑆ߢߪ‫ᧄޔ‬੐ᬺߩᚑᨐࠍ⏕ታߦᚑᾫߐߖ‫ޔ‬ᄢ⋡ᮡߦ߻ߌߚᄢ߈ߥᚑᨐࠍ਄ߍߚ޿ߣ⠨߃ߡ޿ࠆ‫ޕ‬
㧔ᢥ⽿㧦ฎᏒ೰ผ㧕
5. ⅣႺ⋭ ⅣႺ⎇ⓥ✚วផㅴ⾌㧦㜞ੱญኒᐲ࿾ၞߦ߅ߌࠆቅ┙ߒߚ㔤㐳㘃୘૕⟲ߩᜬ⛯⊛
଻⼔▤ℂߦ㑐ߔࠆ⎇ⓥ
ᧄ⎇ⓥߪ‫ޔ‬20 ᐕᓟߩ਎⇇ߦᄙߊߩ㔤㐳㘃⒳߇዁᧪ߦࠊߚߞߡሽ⛯น⢻ߥ⁁ᘒߢᱷߞߡ޿ࠆߎߣࠍⓥᭂߩ⋡⊛ߣ
ߒ‫ޔ‬ቅ┙୘૕⟲ߩሽ⛯ߩ࡝ࠬࠢⷐ࿃ߦ㑐ߔࠆቇⴚ⊛ߥ⎇ⓥߣ଻⼔᡽╷߳ߩឭ⸒ࠍ‫ߢ߹ࠇߎޔ‬ᣣᧄੱ⎇ⓥ⠪߇ᷓߊ㑐
ࠊߞߡ߈ߚࠕࡈ࡝ࠞ‫ޔࠕࠫࠕޔ‬ᣣᧄߩࡈࠖ࡯࡞࠼ߢታᣉߔࠆ‫⎇ᧄޕ‬ⓥࠍ᭴ᚑߔࠆ 3 ߟߩࠨࡉ㧙㑆ߩ੹ᐕᐲߩᚑᨐߪ
એਅߩㅢࠅߢ޽ࠆ‫ޕ‬
1) ᦨዊሽ⛯น⢻㓸࿅ߩቯ⟵ߦ߻ߌߚቅ┙୘૕⟲ߩ↢ᘒቇ⊛࡮㓸࿅ㆮવቇ⊛⎇ⓥ
♮⹜ᢱߩㆮવሶಽᨆᴺߩᡷ⦟
2 Ბ㓏ࡑ࡞࠴ࡊ࡟࠶ࠢࠬ PCR ᴺߩ᦭ലᕈࠍᬌ⸛ߒߚ⚿ᨐ‫ ╙ޔ‬1 Ბ㓏ߩ PCR ߦ߅ߌࠆࡊ࡜ࠗࡑ࡯┹วߦࠃࠆ PCR
ߩല₸ૐਅߪᦨ⚳್ቯߦᄢ߈ߊᓇ㗀ߒߥ޿ߎߣ‫ ╙ޔ‬2 Ბ㓏ߩ PCR ߢߪࡊ࡜ࠗࡑ࡯┹วࠍ⠨ᘦߒ⋧ᕈߩࠃ޿ࡊ࡜ࠗ
ࡑ࡯ࠍ࠮࠶࠻ߦߔࠇ߫‫ ࠬࠢ࠶࡟ࡊ࠴࡞ࡑޔ‬PCR ߢหᤨߦⶄᢙߩᐳ૏ߦ㑐ߔࠆᄙဳᖱႎ߇ᓧࠄࠇࠆߎߣ‫߇ޔ‬᣿ࠄ߆
ߦߥߞߚ‫⚿ߩߎޕ‬ᨐ‫ޔ‬೨ᐕᐲߦ⺞ᩏߢ߈ߚᮡ⼂ᐳ૏ߣᲧߴࠆߣᬌᩏߢ߈ࠆㆮવᮡ⼂ߪ‫ޔ‬Ᏹᨴ⦡૕ STR ᮡ⼂ߢߪ 6
ᐳ૏߆ࠄ 17 ᐳ૏‫ޔ‬Y ᨴ⦡૕ STR ᮡ⼂ߢߪ 6 ᐳ૏߆ࠄ 12 ᐳ૏‫ߣޔ‬㘧べ⊛ߦჇടߒߚ‫ޕ‬ᓥߞߡ‫ߩߎޔ‬ᡷ⦟ߦࠃࠅ‫ޔ‬
㧙㧙
DNA ෼㊂߇ૐ޿♮⹜ᢱߢ߽‫್ޔ‬ቯ⚿ᨐߩౣ⃻ᕈ⏕⹺ߩߚ߼ߦ⹜ᢱࠍᶖ⠻ߔࠆ࡝ࠬࠢ߇✭๺ߐࠇ‫⹜ޔ‬ᢱࠍ▵⚂⊛ߦ
೑↪ߢ߈ࠆߎߣߢએ೨ߦߪಽᨆߒߚߊߡ߽ኻ⽎ߦߢ߈ߥ߆ߞߚᮡ⼂ᐳ૏߇⺞ߴࠄࠇࠆࠃ߁ߦߥࠅ‫ޔ‬ᄙߊߩࡑ࡯ࠞ࡯
ߢ୘૕⟲ߩᄙ᭽ᕈࠍߐࠄߦቴⷰ⊛ߦ⹏ଔߢ߈ࠆࠃ߁ߦߥߞߚ‫ޕ‬
㘃ੱ₎ࠍኻ⽎ߣߒߚ࿾ၞ୘૕⟲ߩ᭴ㅧߩ⎇ⓥߣ♮⹜ᢱߦࠃࠆㆮવሶ᭴ㅧߩಽᨆ
ࡏࡁࡏߦߟ޿ߡߪ‫ޔ‬Wamba/Iyondji ࿾඙‫ޔ‬Lomako ࿾඙‫ޔ‬Salonga ࿾඙‫ޔ‬TL2 ࿾඙‫ޔ‬Lac Tumba ࿾඙‫ޔ‬Malebo ࿾඙
߆ࠄߩࠨࡦࡊ࡞෼㓸ߦᚑഞߒߚ‫ࠍ࡞ࡊࡦࠨߩࠄࠇߎޔߚ߹ޕ‬਄⸥ߩᣇᴺߢಽᨆߔࠆߎߣߦࠃࠅ‫↢ߩࡏࡁࡏޔ‬ᕷၞߩ
߶߷ోၞߦࠊߚࠆࡂࡊࡠ࠲ࠗࡊߩಽᏓ᭴ㅧࠍ᣿ࠄ߆ߦߔࠆߎߣ߇ߢ߈ߚ‫ޔߚ߹ޕ‬TL2 ߢߪ‫ઁޔ‬࿾ၞߣಽൻߒߚࡂࡊ
ࡠ࠲ࠗࡊߢ᭴ᚑߐࠇࠆ࡙࠾࡯ࠢߥࠢ࡜ࠬ࠲࡯߇หቯߐࠇߚ‫⚿ߩࠄࠇߎޕ‬ᨐߪ‫ߩࡓ࡜ࠣࡠࡊో଻ߩࡏࡁࡏޔ‬૞ᚑߦ޽
ߚߞߡߩࡊ࡜ࠗࠝ࡝࠹ࠖ࡯㓸࿅ߩㆬቯ╬ߦᄢ߈ߊ⽸₂ߔࠆ‫ޕ‬
࠾ࡎࡦࠩ࡞ߦߟ޿ߡߪ‫↢ޔ‬ᕷ⁁ᴫ߿୘૕⟲േᘒߩ⇣ߥࠆ 3 ߟߩ୘૕⟲߆ࠄᓧߚ DNA ⹜ᢱࠍಽᨆߒ‫ޔ‬MHC-STR6
ᐳ૏ߣ non MHC-STR 20 ᐳ૏ߦߺࠄࠇࠆᄙ᭽ᕈࠍቯ㊂ߒߚ‫੹ߦߊߣޕ‬࿁⠨᩺ߒߚ MHC ㆮવሶߩ㓞ធ㗔ၞߩᄙ᭽ᕈ
ߦ㑐ߔࠆಽᨆߢߪ‫ޔ‬ᓥ᧪ߩ MHC ߩ࠲ࠗࡇࡦࠣߣߪ⇣ߥߞߡ↢ሽଔߦ㑐ଥߔࠆㆮવሶߩᄙ᭽ᕈࠍ⹏ଔߔࠆߎߣ߇ߢ
߈‫ޔ‬㊁↢୘૕⟲ߩ⛘Ṍ࡝ࠬࠢࠍㆮવቇ⊛ߦ᣿ࠄ߆ߦߔࠆߎߣߦᓎ┙ߟน⢻ᕈ߇޽ࠆ‫ޕ‬
㜞ੱญኒᐲ࿾ၞߦ↢ᕷߔࠆࠕࠫࠕߩࡑࠞࠢ㘃ߦߟ޿ߡߪ‫࡯ࠠࡦࡕࠢ࠻ߩࠞࡦ࡜࡝ࠬޔ‬3 ੝⒳ߣࡃࡦࠣ࡜࠺ࠪࡘߩ
ࠕࠞࠥࠩ࡞ߦߟ޿ߡ‫ޔ‬ᄙߊߩ㓸࿅߆ࠄ♮⹜ᢱࠍ෼㓸ߒߡ DNA ࠍ᛽಴ߒ‫ޔ‬ㆮવሶᄙဳߣߘߩಽᏓߦ㑐ߔࠆ੍஻⊛ಽ
ᨆࠍⴕߞߚ‫ޕ‬
2) ቅ┙୘૕⟲ߦ߅ߌࠆੱ₞౒ㅢᗵᨴ∝ߩ࡝ࠬࠢࠕ࠮ࠬࡔࡦ࠻ߣࠨ࡯ࡌࠗ࡜ࡦࠬ
ࠕࡈ࡝ࠞ㘃ੱ₎ߩੱ₞౒ㅢᗵᨴ∝ࠨ࡯ࡌࠗ࡜ࡦࠬ㧦
੹ᐕᐲߪࠕࡈ࡝ࠞ߆ࠄߩ㊁↢࠴ࡦࡄࡦࠫ࡯㧛ࡏࡁࡏߩ♮ଢࠨࡦࡊ࡞߆ࠄߩ∛ේᓸ↢‛․⇣᛫૕ߦࠃࠆࠬࠢ࡝࡯
࠾ࡦࠣࠍⴕߥ߁ߚ߼‫ޔ‬㊁↢࠴ࡦࡄࡦࠫ࡯㧛ࡏࡁࡏ♮ଢߩ᛽಴ᶧ߆ࠄౣ⃻ᕈ⦟ߊ EBV IgA ᛫૕ࠍᬌ಴ߔࠆߎߣߦᚑ
ഞߒߚ‫⚿ߩࠄࠇߎޕ‬ᨐ߆ࠄ‫ߢࠞ࡝ࡈࠕޔ‬෼ᜪߒੇ῎଻ሽߐࠇߚ♮ଢࠨࡦࡊ࡞߆ࠄߩ᛫࠙ࠗ࡞ࠬ․⇣᛫૕ߩ᛽಴࡮ᬌ
಴ᣇᴺ߇⏕┙ߢ߈ߚ߽ߩߣ⠨߃ࠄࠇߚ‫ߩࠄࠇߎޔߢߎߘޕ‬᛫࠙ࠗ࡞ࠬ․⇣᛫૕ߩᬌ಴♽ࠍ↪޿ߡ‫ޔ‬㊁↢ࡏࡁࡏ 65
㗡ߦ߅ߌࠆ EBV ߦኻߔࠆ․⇣᛫૕ߩ଻᦭₸ࠍ⺞ߴߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬65 㗡ਛ 15 㗡ߦ߅޿ߡ EBV ߦኻߔࠆ․⇣᛫૕ࠍ
଻᦭ߒߡ޿ࠆߎߣ߇᣿ࠄ߆ߦߥߞߚ‫ޕ‬㔤㐳㘃⎇ⓥᚲߢ㘺⢒ߐࠇߡ޿ࠆ 14 㗡ߩ࠴ࡦࡄࡦࠫ࡯߆ࠄᓧߚⴊᶧ↱᧪⹜ᢱ
ࠍ↪޿ߡฦ⒳∛ේ૕ߦኻߔࠆ․⇣⊛ IgG ᛫૕ߩ᦭ήࠍᬌ⸛ߒߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬15 ⒳ߩࡅ࠻∛ේᓸ↢‛ߦኻߔࠆ᛫૕
߇ᬌ಴ߐࠇߚ‫ߩࠄࠇߎޕ‬ᄙߊߪࡅ࠻↱᧪ߛߣ⠨߃ࠄࠇ‫ੱߩ߳࡯ࠫࡦࡄࡦ࠴ࠄ߆࠻ࡅޔ‬₞౒ㅢᗵᨴ∝ߩᗵᨴ߇੍ᗐએ
਄ߦᐢ߇ߞߡ޿ࠆߎߣ߇ࠊ߆ߞߚ‫ޕ‬
ᰴߦ‫ ߦ․ޔ‬IgG ᛫૕ଔ߇㜞߆ߞߚ᛫ EBV ᛫૕ߦߟ޿ߡ‫♮ޔ‬ଢ⹜ᢱߩ᛽಴ᶧౝߩ IgA ᛫૕ߩ᦭ήߦߟ޿ߡᬌ⸛ࠍ
ⴕߥߞߚ‫⚿ߩߘޕ‬ᨐ‫ࠬ࡞ࠗ࠙ޔ‬ᗵᨴೝỗߦࠃࠅኋਥ఺∉ᔕ╵ߣߒߡ IgG ဳߣ IgA ဳ߇หᤨߦ⺃ዉߐࠇ߁ࠆߎߣ‫⺃ޔ‬
ዉߐࠇߚ IgA ߇♮⹜ᢱ߆ࠄᬌ಴น⢻ߢ޽ࠆߎߣ߇␜ߐࠇߚ‫⹜♮ޔࠄ߆ߣߎߩߎޕ‬ᢱ߆ࠄߩ IgA ᛫૕ߩᬌ಴߇‫ޔ‬㊁↢
㘃ੱ₎ߩੱ₞౒ㅢᗵᨴ∝ߩᗵᨴ⁁ᴫߩࠨ࡯ࡌࠗ࡜ࡦࠬߦ᦭ലߢ޽ࠆߎߣ߇⏕߆߼ࠄࠇߚ‫ޕ‬
ᄖ᧪ࡑࠞࠢࠩ࡞↱᧪∛ේ૕ߦࠃࠆ࠾ࡎࡦࠩ࡞ߩᗵᨴ∝࡝ࠬࠢ⎇ⓥ㧦
ᤓᐕᐲߦᒁ߈⛯߈‫ޔ‬੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲߢ⊒↢ߒߚ‫ⴊޔ‬ዊ᧼ߩᕆỗߥᷫዋߢ࠾ࡎࡦࠩ࡞߇ᱫ੢ߔࠆ∛᳇ߩේ࿃
⸃᣿ߦ޽ߚߞߚ‫⚿ߩߘޕ‬ᨐ‫∛ߩߎޔ‬᳇ߩේ࿃߇⥄ὼኋਥߢ޽ࠆࠞ࠾ࠢࠗࠩ࡞߆ࠄߩ SRV-4 ߩᗵᨴߢ޽ࠆߎߣ߇߶߷
⏕⹺ߐࠇߚ‫ޔߦࠄߐޕ‬SRV-4 ߩ∉ቇ⺞ᩏ߆ࠄ‫ޔ‬SRV-4 ߪỚෘធ⸅ߦࠃࠅᗵᨴ߇ᚑ┙ߒߡ޿ࠆߎߣ߇‫ޔ‬㘺⢒ᣉ⸳ߏߣ
ߩ∉ቇ⺞ᩏߦࠃࠅ߆᣿ࠄ߆ߦߥߞߚ‫ޕ‬એ਄ߩߎߣ߆ࠄ‫ޔ‬ᄖ᧪ࡑࠞࠢࠩ࡞ߦ⥄ὼᗵᨴߒߡ޿ࠆ∛ේᓸ↢‛߇࠾ࡎࡦࠩ
࡞ߦ‫ޟ‬⒳ߩო‫ߡ߃⿧ࠍޠ‬વ᠞ߔࠆߎߣߦࠃࠆෂ㒾ᕈߦߟ޿ߡలಽᵈᗧߔߴ߈ߎߣ‫੹ޔ‬ᓟߪᧂߛᧂ⸃᣿ㇱಽߢ޽ࠆ࠾
ࡎࡦࠩ࡞ߦ߅ߌࠆⴊዊ᧼ᷫዋ∝ߩ⊒∝ࡔࠞ࠾࠭ࡓߩ⸃ᨆ߇ᔅⷐߢ޽ࠆߣ⠨߃ࠄࠇߚ‫ޕ‬
3) ቅ┙୘૕⟲ߩ⃻⁁ಽᨆߣ↢ᕷ࿾ߩ⛽ᜬ࡮࿁ᓳߩߚ߼ߩ↢ᘒቇ⊛࡮␠ળቇ⊛⎇ⓥ
࠺࡯࠲ࡌ࡯ࠬߩ᜛ల
ኻ⽎ߣߒߚᄢඨߩ୘૕⟲ߦ߅޿ߡ‫ࠍࠬ࡯ࡌ࠲࡯࠺ޔ‬᭴▽ߢ߈ߚ‫੹ޕ‬ᓟ‫↹ޔ‬௝ߩ૶↪⸵⻌ߥߤߩᚻ⛯߈߇ቢੌߔࠇ
߫‫ࠫ࡯ࡍࡓ࡯ࡎޔ‬਄ߢ౏㐿ߔࠆ੍ቯߢ޽ࠆ‫ޕ‬ሽ⛯น⢻ᕈಽᨆߦᔅⷐߥࡄ࡜ࡔ࡯࠲ߪ‫ޔ‬㐳ᦼ⎇ⓥ߇ⴕࠊࠇߡ޿ࠆ࠴ࡦ
ࡄࡦࠫ࡯ߩቅ┙୘૕⟲㧔ࠞ࡝ࡦ࠭࡮ࡏ࠶࠰࠙࡮ࡑࡂ࡟㧕ߣࡏࡁࡏߩቅ┙୘૕⟲ߦߟ޿ߡ෼㓸࡮ᢛℂߢ߈ߚ‫ࠄࠇߎޕ‬
ߩ୘૕⟲ߢߪ‫ޔ‬ሽ⛯ߦᓇ㗀ࠍ෸߷ߔੱ㑆ᵴേߦߟ޿ߡߩ⹦⚦ߥ࠺࡯࠲߽෼㓸ߢ߈ߡ޿ࠆ‫ࠍ࠲࡯࠺ߩࠄࠇߎޕ‬ᵴ↪ߔ
ࠆߎߣߢ‫ޔ‬⠨߃ࠄࠇࠆ଻ోኻ╷ߦࠃࠆലᨐ߿ੱ㑆␠ળߦ߆߆ࠆࠦࠬ࠻ࠍ੍᷹ߒߚ߁߃ߢ‫ౕޔ‬૕⊛ߥ଻ోኻ╷ࠍ⸘↹
ߒ‫ޔ‬ታ⸽⊛ߥ⎇ⓥࠍ㐿ᆎߔࠆߎߣ߇ߢ߈ࠆ‫ޕ‬
ሽ⛯น⢻ᕈಽᨆߦࠃࠆ੍᷹ߩᅷᒰᕈߩᬌ⸛
㊄⪇ጊፉ୘૕⟲ߩ୘૕ᢙჇട₸ߣߘࠇߦᓇ㗀ࠍ෸߷ߔ 3 ߟߩേᘒࡄ࡜ࡔ࡯࠲ߪ‫ⷐߥ߹ߑ߹ߐޔ‬࿃ߩⶄว⊛ߥᓇ㗀
ߦࠃߞߡᄢ߈ߊᄌേߔࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬಴↥₸ߪ೨ᐕߩ಴↥₸ߣ⑺ߩ㘩‛᧦ઙ‫ޔ‬಴↥⋥೨ߩ౻ߩ᳇⽎᧦ઙߩᓇ
㗀ࠍฃߌߡ޿ߚ‫ޕ‬
ߎࠇࠄߩ࠺࡯࠲ߦ߽ߣߠ޿ߚሽ⛯น⢻ᕈಽᨆߢߪ‫ޔ‬2 ㅢࠅߩࠪ࠽࡝ࠝߦߟ޿ߡ୘૕ᢙᄌേࠍࠪࡒࡘ࡟࡯࡚ࠪࡦߒ‫ޔ‬
੍᷹ߩᅷᒰᕈࠍᬌ⸛ߒߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬േᘒࡄ࡜ࡔ࡯࠲ߩᐕᰴᄌേߩᄢ߈޿୘૕⟲ࠍኻ⽎ߣߔࠆ႐วߦߪ‫࡯ࡔ࡜ࡄޔ‬
࠲ߩ㐳ᦼ㑆ᐔဋ߿ಽᢔߥߤߩઍ⴫୯ߛߌߢߪ‫ޔ‬ᅷᒰߥ੍᷹ࠍᓧࠄࠇߥ޿น⢻ᕈ߇㜞ߊ‫ޔ‬േᘒࡄ࡜ࡔ࡯࠲ߩઍ⴫୯ߛ
㧙㧙
ߌߢߥߊᄌേ․ᕈࠍචಽߦᬌ⸛ߒߚ߁߃ߢ‫ޔ‬ሽ⛯น⢻ᕈಽᨆࠍⴕ߁ᔅⷐ߇޽ࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬
ᦨᓟߦ‫ޔ‬㊄⪇ጊፉ୘૕⟲ߩ 100 ᐕᓟ߹ߢߩ୘૕ᢙᄌേࠍࠪࡒࡘ࡟࡯࡚ࠪࡦߒߚ⚿ᨐ‫ޔ‬୘૕ᢙߪߐࠄߦᷫዋߒߡ޿
ߊน⢻ᕈ߇㜞޿ߎߣ߇␜ߐࠇߚ‫ޕ‬୘૕⟲ߩ቟ቯߒߚ⛽ᜬߩߚ߼ߦߪ‫↢ޔ‬ᕷⅣႺߩᄌൻߣ୘૕⟲Ⴧട₸ߩૐਅ߹ߚߪ
ᱫ੢₸ߩჇടߣߩ࿃ᨐ㑐ଥࠍಽᨆߒ‫ޔ‬ኻ╷ࠍ⻠ߓࠆᔅⷐ߇޽ࠆ‫ޕ‬
ੱὑ⊛ߥ୘૕ᢙỗᷫ߇ፉᎩߩቅ┙୘૕⟲ߩㆮવ⊛ᄙ᭽ᕈߦ෸߷ߔᓇ㗀ߩᬌ⸛
ፉᎩ㧔㊄⪇ጊፉ㧕ߣᧄ࿯㧔ችၔ⋵ᧄ࿯㧕ߩ୘૕⟲ߦ㑐ߔࠆ㓸࿅ㆮવቇ⊛ಽᨆࠍⴕߞߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬ፉᎩ࡮ᧄ࿯ߦ
߆߆ࠊࠄߕ‫ੱޔ‬ὑ⊛ߥ୘૕ᢙỗᷫߪ‫ࠢ࠶ࡀ࡞࠻ࡏޔ‬ലᨐߣਇචಽߥ᦭ല㓸࿅ࠨࠗ࠭ߦࠃࠆᒝ޿ㆮવ⊛ᶋേࠍᒁ߈⿠
ߎߒ‫ޔ‬ㆮવ⊛ᄙ᭽ᕈߩૐਅࠍടㅦߐߖࠆߎߣ߇␜ໂߐࠇߚ‫ߦߊߣޕ‬ፉᎩ୘૕⟲ߢߪ‫ޔ‬ⅣႺ෼ኈജߦࠃߞߡ᦭ല㓸࿅
ࠨࠗ࠭ߩ࿁ᓳ߇⪺ߒߊᛥ೙ߐࠇߚߎߣߢ‫ޔ‬ㆮવ⊛ᶋേ߇ࠃࠅᒝജߦ௛޿ߚน⢻ᕈ߇޽ࠆ‫ߩࠄࠇߎޕ‬୘૕⟲ߢߪ‫᦭ޔ‬
ല㓸࿅ࠨࠗ࠭ߩㅦ߿߆ߥ࿁ᓳ߇⷗ࠄࠇߥߌࠇ߫‫ޔ‬ㆮવ⊛ᄙ᭽ᕈ߇ᰴ╙ߦᄬࠊࠇߡ޿ߊ߅ߘࠇ߇޽ࠆ‫߈ⴕޕ‬ㆊ߉ߚ᦭
ኂ㚟㒰߿୘૕ᢙ▤ℂ߽‫ޔ‬዁᧪ߩㆮવ⊛ᄙ᭽ᕈߦห᭽ߩᓇ㗀ࠍ෸߷ߔߎߣ߇ᗐ௝ߢ߈ࠆ‫ޕ‬ㆊ෰ߦᄢⷙᮨߥ❗ዊࠍ⚻㛎
ߒߚ୘૕⟲‫߮ࠃ߅ޔ‬ⅣႺ෼ኈജߩዊߐߥፉᎩ୘૕⟲ߩ଻ో▤ℂߦߪ‫ࠅࠃޔ‬ᘕ㊀ߥᆫ൓߇᳞߼ࠄࠇࠆߎߣ߇␜ߐࠇߚ‫ޕ‬
㧔ᢥ⽿㧦ฎᏒ೰ผ㧕
6. 㗡⣖ᓴⅣࡊࡠࠣ࡜ࡓ
‫ੱޟ‬㑆ࠄߒߐߩ㔤㐳㘃⊛⿠Ḯࠍߐߋࠆᚢ⇛⊛࿖㓙౒ห⎇ⓥ‫ޠ‬
ᦨవ┵⎇ⓥ㐿⊒ᚢ⇛⊛ᒝൻ⾌⵬ഥ㊄ߦࠃࠆ‫⧯ޔ‬ᚻ⎇ⓥ⠪ߩੱ᧚⢒ᚑߣ࿖㓙౒ห⎇ⓥߩ᦭ᯏ⊛ㅪ៤ߦࠃࠆ੐ᬺߩ
ផㅴߣߒߡታᣉߐࠇࠆ‫ޔ‬㗡⣖ᓴⅣࠍᵴᕈൻߔࠆ⧯ᚻ⎇ⓥ⠪ᶏᄖᵷ㆜ࡊࡠࠣ࡜ࡓߢ޽ࠆ‫ޕ‬ᐔᚑ22ᐕᐲ10᦬߆ࠄታᣉߐ
ࠇߚᣂߚߥ10ࡩ᦬એ਄ߩ㐳ᦼᵷ㆜੐ᬺߢ‫ޔ‬ᐔᚑ25ᐕ3᦬31ᣣ߹ߢߩ3ᐕᐲ⛮⛯ߔࠆ߽ߩߢ޽ࠆ‫ޕ‬ᐔᚑ23ᐕᐲߪ
14,330,000౞ភ⟎ߐࠇߚ‫ޕ‬
ᧄࡊࡠࠣ࡜ࡓߪ‫ޔ‬ᣢߦዷ㐿ߒߡ޿ࠆ࿖㓙౒ห⎇ⓥࡀ࠶࠻ࡢ࡯ࠢߦࠝ࡜ࡦ࠳ࠍട߃‫ޔ‬੐ᬺߩ᜛ల‫ޔ‬ᒝൻ‫ޔ‬ᷓൻߐ
ߖ㗡⣖ᓴⅣࡊࡠࠣ࡜ࡓࠍ౞Ṗߦផㅴߔࠆߎߣࠍ⋡⊛ߣߔࠆ‫ޕ‬
‫ੱޟ‬㑆ࠄߒߐ‫ࠍޠ‬ⓥ᣿ߔࠆ┵✜ߣߒߡਅ⸥ߩ3ઙߩ⎇ⓥ
㗄⋡ࠍឝߍߚ‫(ޕ‬1)ᗧ⼂ߩࡔࠞ࠾࠭ࡓ㧦ࡅ࠻ߣࠨ࡞㘃ߩ㤗㈮૞↪ᯏᐨࠍ⑼ቇ⊛ߦᲧセ⸃ᨆߒ‫ޔ‬ᗧ⼂ߩᶖᄬᯏᐨߩⷞὐ
߆ࠄ㜞ᰴߩ♖␹ᵴേߩࡔࠞ࠾࠭ࡓࠍⓥ᣿ߔࠆ┵✜ߣߔࠆ‫ޕ‬㧔ᵷ㆜ᯏ㑐㧦⧷࿖࡮࡝ࡦࠞ࡯ࡦᄢቇ↢‛⑼ቇ⻠ᐳ‫࡜ࠝޔ‬
ࡦ࠳࡮ࡈࡠ࡯࠾ࡦࠥࡦᄢቇක≮࠮ࡦ࠲࡯㧕(2)ࡅ࠻ߩ㖸ჿ⸒⺆ߩ⿠Ḯ㧦㔤㐳㘃ߩࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߩㅴൻߣࡅ࠻ߩ
㖸ჿ⸒⺆ߩ⿠Ḯߦ㑐ߔࠆ⎇ⓥࠍᲧセ⹺⍮⑼ቇߩⷰὐ߆ࠄផኤߒ‫ߩ࠻ࡅޔ‬㖸ჿ⸒⺆ᯏ᭴ߩ⸃᣿ࠍ೨ឭߣߒߡ‫ޔ‬㔤㐳㘃
ߩࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߩᄙ᭽ᕈߣߘߩㅴൻᯏ᭴ࠍ᣿ࠄ߆ߦߔࠆ‫ޕ‬㧔ᵷ㆜ᯏ㑐㧦⧷࿖࠮ࡦ࠻ࠕࡦ࠼࡝ࡘ࡯࠭ᄢቇᔃℂ
ቇㇱ࡮ࠛࠫࡦࡃ࡜േ‛࿦㧕(3)Უ♽࡮ῳ♽␠ળߩ⊒↢ᯏᐨ㧦㔤㐳㘃ߩේὐߣ߽޿߃ࠆේ₎㘃ߩ␠ળࠍ‫ޔ‬㊁ᄖⷰኤߦࠃ
ࠆ࠺ࡕࠣ࡜ࡈࠖ⾗ᢱߣಽሶ㓸࿅ㆮવቇ⊛⾗ᢱࠍ⛔วߒߡ⸃ᨆߒ‫ߩઁߩߘ߿࠻ࡅޔ‬㔤㐳㘃ߣߩᲧセࠍㅢߒߡ‫ࠠࠝࡢޔ‬
࠷ࡀࠩ࡞ߩᲣ♽␠ળߩ⊒↢ᯏᐨࠍ᣿ࠄ߆ߦߔࠆ‫ߩߘޕ‬ኻ⽎␠ળߣߒߡ࠴ࡦࡄࡦࠫ࡯߅ࠃ߮ࡏࡁࡏߩῳ♽␠ળ߽⸃ᨆ
ߔࠆ‫ޕ‬㧔ᵷ㆜ᯏ㑐㧦࠼ࠗ࠷㔤㐳㘃࠮ࡦ࠲࡯‫ޔ‬ᵷ㆜ࡈࠖ࡯࡞࠼㧦ࡑ࠳ࠟࠬࠞ࡞㧕ߎࠇࠄߩ⎇ⓥ㗄⋡ߪ⣖㧙⸒⺆㧙␠ળ
ߣ޿߁ᔅὼ⊛ߦ⋧੕ߦ㑐ଥࠍ᦭ߔࠆ߽ߩߢ޽ࠅ‫੹ޔ‬ᓟߩቇ㓙⊛⎇ⓥߩ⊒ዷࠍⷞ㊁ߦ޿ࠇߡ޿ࠆ‫ߦߎߎޕ‬ឝߍߚ⎇ⓥ
ᯏ㑐ߦട߃ߡ‫ޔ‬ᓥ᧪߆ࠄ♖ജ⊛ߦ⛮⛯ߒߡ޿ࠆࡈࠖ࡯࡞࠼⎇ⓥ᜚ὐ߽฽߼ߡ✚ว⊛ߦㆇ↪ߔࠆ‫ޕ‬
ᐔᚑ23ᐕᐲߪ(1)‫ޟ‬ᗧ⼂ߩࡔࠞ࠾࠭ࡓ‫⧷ߪࡓ࡜ࠣࡠࡊޠ‬࿖࡮࡝ࡦࠞ࡯ࡦᄢቇߦችㇱ⾆ሶഥᢎࠍᵷ㆜ߒ‫੹ޔ‬ᓟᗧ⼂
ߩ⸃ᨆߦ㊀ⷐߦߥࠆ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ㗻ߩ⴫ᖱ⹏ଔ㧔․ߦ∩ߺߦߟ޿ߡ㧕ߩࠦࡦࡇࡘ࡯࠲⸃ᨆࠍታᣉߒߚ‫(ޕ‬2)
‫ߩ࠻ࡅޟ‬㖸ჿ⸒⺆ߩ⿠Ḯ‫⧷ߪࡓ࡜ࠣࡠࡊޠ‬࿖࡮࠮ࡦ࠻ࠕࡦ࠼࡝ࡘ࡯࠭ᄢቇ߅ࠃ߮ࠛ࠺ࠖࡦࡃ࡜േ‛࿦ߦ‫ޔ‬㚅↰໪⾆
ഥᢎࠍᵷ㆜ߒ‫⇣ߩ࡞ࠩࠬ࡝ߣ࡞ࠩࠠࡑࠝࠨࡈޔ‬⒳㑆㖸ჿࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦߩ⸃ᨆࠍ߅ߎߥ߁⸘↹ࠍታᣉߒߚ‫ޕ‬
(1)(2)ߩਔࡊࡠࠣ࡜ࡓߪ⸃ᨆߩ✜ߦ⌕޿ߚ߫߆ࠅߢ‫ߛ߹ޔ‬ᚑᨐߪ਄߇ߞߡ޿ߥ޿߇‫ޔ‬ਥᜂᒰ⎇ⓥ⠪ᐔ੗߇ਔ⎇ⓥᯏ
㑐ࠍⷞኤߒ‫ޔ‬ኻᔕᢎຬߣ⹤ߒว޿ࠍ߅ߎߥߞߚߣߎࠈ‫ޔ‬㕖Ᏹߦදജ⊛ߢ‫ޔ‬ᣉ⸳ⅣႺ߽⸘↹ࠍ㆐ᚑߔࠆߚ߼ߦචಽߥ
߽ߩߢ޽ߞߚ‫ޕ‬ᚑᨐߩ㆐ᚑࠍචಽߦᦼᓙߢ߈ࠆߣ⏕ାߒߚ‫(ޕ‬3)‫ޟ‬Უ♽࡮ῳ♽␠ળߩ⊒↢ᯏᐨ‫࠷ࠗ࠼ߪࡓ࡜ࠣࡠࡊޠ‬
㔤㐳㘃࠮ࡦ࠲࡯ߥࠄ߮ߦࡑ࠳ࠟࠬࠞ࡞߳Ꮢ㊁ㅴ৻㇢ᢎോ⵬૒ຬࠍᵷ㆜ߒ‫ߢ߹ࠇߎޔ‬෼㓸ߒߡ߈ߚࡢࠝࠠ࠷ࡀࠩ࡞
㧔Უ♽␠ળࡕ࠺࡞㧕ߩⴊᶧࠨࡦࡊ࡞ࠍ૶ߞߡಽሶ㓸࿅ㆮવቇ⊛⸃ᨆࠍ߅ߎߥߞߚ‫ߪߢ࡞ࠞࠬࠟ࠳ࡑޕ‬ᓥ᧪ߩⷰኤ࠺
࡯࠲ߦᣂߒߊ↢ߓߚ㓸࿅ߩᄌൻ࠺࡯࠲ࠍㅊടߒߚ‫ޔߚ߹ޕ‬ᜂᒰ⎇ⓥ⠪ߩᯅᧄߪ࠙ࠟࡦ࠳ߦ಴ᒛߒ‫⫾ߢ߹ࠇߎޔ‬Ⓧߒ
ߡ߈ߚ࠴ࡦࡄࡦࠫ࡯߅ࠃ߮ࡏࡁࡏ㧔ῳ♽␠ળࡕ࠺࡞㧕ߩ␠ળⴕേ࠺࡯࠲ࠍౣᬌ⸛ߔࠆߣߣ߽ߦ‫ޔ‬ᣂߚߥⷰኤ࠺࡯࠲
ࠍㅊടߒߚ‫(ޕ‬3)ߩࡊࡠࠣ࡜ࡓߪ޿ߕࠇ߽ߎࠇ߹ߢ㐳ᐕ⸃ᨆߒߡ߈ߚ࠺࡯࠲ߦၮߠ޿ߚ⸘↹ߥߩߢ‫ޔ‬Უ♽࡮ῳ♽ߩ࠺
࡯࠲ࠍᲧセ⸃ᨆߔࠆߎߣߢ‫ࠆߥࠄߐޔ‬චోߥᚑᨐ߇ᦸ߼ࠆ‫ޕ‬
㧔ᢥ⽿㧦ᐔ੗໪ਭ㧕
7. ੱ㑆ߩㅴൻ
੩ㇺᄢቇࡉ࡯࠲ࡦ෹ᅢࡊࡠࠣ࡜ࡓ
ࡉ࡯࠲ࡦ࿖ࠍ⥰บߦ‫ޔ‬੩ㇺᄢቇ࿕᦭ߩ㊁ᄖ⎇ⓥߩવ⛔ࠍ〯߹߃ߚోቇ⊛ߥ࿖㓙੤ᵹ੐ᬺࠍ߅ߎߥ߁੐ᬺߢ޽ࠆ‫ޕ‬
ࡉ࡯࠲ࡦߪ‫ੱޔ‬ญ 70 ਁੱߩࡅࡑ࡜ࡗߩዊ࿖ߢ޽ࠆ‫ᦨޕ‬ೋߩ✼ߪ‫ޔ‬1957 ᐕ᥅⑺ߩ᪀ේᱞᄦᢎ᝼ࠄߦࠃࠆ╙ 3 ઍ₺ᅥ
㧙㧙
ߩ᱑ᓙߦ޽ߞߚ‫ޕ‬ਗሽߔࠆฦㇱዪ⁛⥄ߩขࠅ⚵ߺࠍ᧤ߨ‫ޔ‬ᣣᧄߢᦨ߽ࡉ࡯࠲ࡦߣ✼ߩᷓ޿ᄢቇߣߒߡ‫₺ࡦ࠲࡯ࡉޔ‬
┙ᄢቇߥࠄ߮ߦ଻ஜ⋭╬ߣදജߒߡ‫ߩࡦ࠲࡯ࡉޔ‬࿖ᤚߢ޽ࠆ࿖᳃✚ᐘ⑔㊂㧔GNH㧕ࠍߪߓ߼‫ޔ‬ஜᐽ‫ޔ‬ᢥൻ‫ޔ‬቟ో‫ޔ‬
↢ᘒ♽‫⋧ޔ‬੕⽸₂ߩ 5 ߟߩ஥㕙߆ࠄ✚ว⊛ߥ੤ᵹࠍ߅ߎߥ߁‫ߦ߼ߚߩߘޕ‬ᔅ㗇ߥ⃻࿾⺞ᩏ‫ޔ‬੤ᵹ࠺࡯࠲ࡌ࡯ࠬᢛ஻‫ޔ‬
ᤋ௝࡮ᢥᦠࠕ࡯ࠞࠗࡉ૞⵾‫ޔ‬HP ߆ࠄߩ⊒ା‫ޔ‬ᰴ਎ઍࠍᜂ߁⧯ᚻ⎇ⓥ⠪ߩ੤ᵹࡊࡠࠣ࡜ࡓ╬ࠍផㅴߔࠆ‫ޕ‬HP ෳᾖ
http://www.jp.kubhutan.org/
⧷ᢥߩᱜᑼฬ⒓ࠍ‫ޔ‬
‫ޟ‬Kyoto University Bhutan Friendship Program‫ߣޠ‬⒓ߔࠆ‫ޕ‬ᐔᚑ 22 ᐕ 10 ᦬ߦ╙㧝⸰໧࿅㧔㓌㐳㧦
᧻ᴛື㇢߶߆ 6 ฬ㧕ࠍᵷ㆜ߒ‫ޔ‬
GNH ߩឭ໒⠪ߢ޽ࠆ╙ 4 ઍ࿖₺ߩࠫࠣࡒ࡮ࠪࡦࠡ࡮ࡢࡦ࠴ࡘࠢ㧔Jigme Singye Wangchuk㧕
Ლਅߦ߅ળ޿ߒߚ‫ޕ‬એᓟ‫ޔ‬ᐔᚑ 22 ᐕᐲౝߦ 4 ⸰໧࿅ 25 ฬ‫ޔ‬ᐔᚑ 23 ᐕᐲ߽ 4 ⸰໧࿅ 25 ฬࠍᵷ㆜ߒߚ‫ޕ‬ᢎຬߛߌߢ
ߥߊ‫ޔ‬੐ോ⡯ຬ‫⎇ޔ‬ⓥຬ‫ޔ‬ᄢቇ㒮↢‫ޔ‬1 ᐕ↢ࠍ฽߻ቇㇱቇ↢ߢ޽ࠆ‫ޕ‬
ᧄ੐ᬺߪ‫ޔ‬ᐔᚑ 22 ᐕ 3 ᦬ߦ㐿௅ߐࠇߚ੩ㇺᄢቇ㒝⟎⎇ⓥᚲ࡮⎇ⓥ࠮ࡦ࠲࡯ߩᐕᰴࠪࡦࡐࠫ࠙ࡓߦ߅޿ߡోቇ⊛ࡊ
ࡠࠣ࡜ࡓߣߒߡ᭴ᗐߐࠇ‫ޔ‬੩ᄢߩ⎇ⓥᚲ⟲ 22 ㇱዪ߆ࠄߩ౒หឭ᩺ߣߒߡ⊒⿷ߒߚ‫ޕ‬ᐘ޿‫✚⚔ᧄ᧻ޔ‬㐳߿‫ޔ‬ศᎹẖ
⎇ⓥᜂᒰℂ੐ࠄߩℂ⸃ߣᡰេ߇޽ࠅ‫ޔ‬ᐔᚑ 22 ᐕᐲߩ਎⹤ᓎㇱዪߢ޽ࠆౣ↢ක⑼ቇ⎇ⓥᚲ‫ޔ‬ᐔᚑ 23 ᐕᐲߩ↢ሽ࿤⎇
ⓥᚲ߆ࠄోቇ⚻⾌ࠍ↳⺧ߒណᛯߐࠇߚ‫ޕ‬㔤㐳㘃⎇ⓥᚲߪ‫ߩࡓ࡜ࠣࡠࡊߩ߽ߘ߽ߘޔ‬ឭ᩺ㇱዪߣߒߡ‫ޔ‬੩ㇺᄢቇࡉ࡯
࠲ࡦ෹ᅢࡊࡠࠣ࡜ࡓߩታⴕߣ੐ോ૞ᬺࠍᜂᒰߒߚ‫ޕ‬
ᐔᚑ 23 ᐕᐲߩ 4 ⸰໧࿅ 25 ฬߩᷰ⥶ߦ߅޿ߡ‫ޔ‬㔤㐳㘃⎇ⓥᚲߪ‫ߩߘޔ‬ᄢඨߩ⚻⾌ࠍ⽶ᜂߒߚ‫․ޕ‬೎⚻⾌ࡊࡠࠣ࡜
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⋭࿖㓙ኾ㐷ቭ‫ޔ‬ዊఽ⑼ක㧕ࠍ㓹↪ߒ‫ޔ‬5 ᦬߆ࠄㅢᐕߦࠊߚߞߡ‫∛࡯ࡊࡦࠖ࠹┙₺ޔ‬㒮ߦᵷ㆜Ᏹ㚢ߐߖߚ‫ޔ߅ߥޕ‬ᐔ
ᚑ 24 ᐕᐲએ㒠ߪ‫ޔ‬ᒁ߈⛯߈㔤㐳㘃⎇ⓥᚲࠍਥോㇱዪߣߔࠆㇱዪ㑆ㅪ៤੐ᬺߣߒߡ⛮⛯ߔࠆ‫ޕ‬੩ᄢᢎ⢒⎇ⓥ⽷࿅ߩ
ഥᚑ߇᳿߹ߞߡ޿ࠆ‫ޕ‬
㧔ᢥ⽿㧦᧻ᴛື㇢㧕
㧙㧙
Ψ. ᐢႎᵴേ
㔤㐳㘃⎇ⓥᚲߢߪᐢႎᆔຬળ߇ᜂᒰߒߡ‫౏ޔ‬㐿⻠ᐳ‫⎇ޔ‬ⓥᚲ౏㐿‫ࠬࡄࡦࡖࠠࡦࡊ࡯ࠝޔ‬㧔ᄢቇ㒮ࠟࠗ࠳ࡦࠬ㧕ߥߤ
ࠍ㐿௅ߒ‫⎇ޔ‬ⓥᚲߩᵴേࠍᚲᄖߩᣇ‫࡮੺⚫߽ߢߤߥࠫ࡯ࡍࡓ࡯ࡎޔ࠻࠶࡟ࡈ࡯࡝ޔߚ߹ޕࠆ޿ߡߒ੺⚫ߦޘ‬ᐢႎᵴ
േࠍ߅ߎߥߞߡ޿ࠆ‫ޕ‬
1. ౏㐿⻠ᐳ
›ጊ౏㐿⻠ᐳ‫ޟ‬㔤㐳㘃ቇߩᗢߒߺ‫ޠ‬
2011 ᐕ 7 ᦬ 30 ᣣ㧔࿯㧕㨪31 ᣣ㧔ᣣ㧕‫ޔ‬੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲߦߡ㐿⻠ߒߚ‫ޕ‬ෳട⠪ߪ 47 ฬ‫ޕ‬
㧨ࡊࡠࠣ࡜ࡓ㧪
⻠⟵㧦㜞੗ᱜᚑ‫ߚ᧪ߩ࡞ࠨޟ‬㆏‫ޠ‬
⻠⟵㧦ࡑࠗࠤ࡞࡮ࡂࡈࡑࡦ‫ޟ‬㔤㐳㘃ߩ⮎⤝ᢱℂ㧦੍㒐ߣᴦ≮ߩ਎⇇‫ޠ‬
⻠⟵㧦ᨋ⟤㉿‫⊒ߩߘߣࠈߎߎߩ࡯ࠫࡦࡄࡦ࠴ޟ‬㆐‫ޠ‬
⻠⟵㧦ጟᧄቬ⵨‫ߩ࡞ࠨޟ‬ነ↢⯻ߣࡅ࠻ߩነ↢⯻㧦౒ㅴൻߣኋਥォ឵‫ޠ‬
ታ⠌㧦ᒻᘒ‫ޟ‬㛽ቇ࡮ൻ⍹‫ޠ‬㧔Ძ೑ବ㓶㧕
ᔃℂ‫ⷰ࡯ࠫࡦࡄࡦ࠴ޟ‬ኤ‫ޠ‬㧔ᨋ⟤㉿㧕
↢ᘒ‫ⴕ࡞ࠩࡦࡎ࠾ޟ‬േⷰኤ‫ޠ‬㧔ᷰㆻ㇌ᄦ㧕
ㆮવ‫ޟ‬ㆮવሶಽᨆ‫ޠ‬㧔Ꮉᧄ⧐㧕
᧲੩౏㐿⻠ᐳ‫ࠍ࡞ࠨޟ‬ㅢߒߡߺߚ਎⇇‫ޠ‬
2011 ᐕ 9 ᦬ 25 ᣣ㧔ᣣ㧕‫ޔ‬ᣣᧄ⑼ቇᧂ᧪㙚 7 㓏ߺࠄ޿ CAN ࡎ࡯࡞ߦߡታᣉߒߚ‫ޕ‬ෳട⠪ߪ 206 ฬ‫ޕ‬
㧨ࡊࡠࠣ࡜ࡓ㧪
ᚲ㐳᜿ᜦ㧦᧻ᴛື㇢
⻠Ṷ 1㧦ㄞᄢ๺‫ࠍ࡞ࠨޟ‬ㅢߒߡ↢߈‛ߩߟߥ߇ࠅࠍ⠨߃ࠆ㨪㊄⪇ጊፉߩ࠾ࡎࡦࠩ࡞ߩⷰኤ߆ࠄ㨪‫ޠ‬
⻠Ṷ 2㧦㚅↰໪⾆‫ޟ‬᱌߁ࠨ࡞ߩਇᕁ⼏㧦࠹࠽ࠟࠩ࡞ߩࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦ‫ޠ‬
⻠Ṷ 3㧦㜞↰᣽ᒾ࡮᧻ᧄᱜᐘ‫ߣࡦࡒࡄ࡯࠼ޟ‬⣖ᯏ⢻㧦ࡄ࡯ࠠࡦ࠰ࡦ∛߆ࠄႎ㈽ⴕേ߹ߢ‫ޠ‬
⻠Ṷ 4㧦ฎ⾐┨ᒾ‫ ߩࡓࡁࠥ࠻ࡅޟ‬45㧑ࠍභ߼ࠆേߊㆮવሶ‫ޠ‬
⾰⇼ᔕ╵
2. ╙ 21 ࿁Ꮢ᳃౏㐿ᣣ
2011 ᐕ 10 ᦬ 30 ᣣ㧔ᣣ㧕13㧦00 ߆ࠄ 15㧦30 ߹ߢᏒ᳃౏㐿߇߅ߎߥࠊࠇߚ‫ޕ‬ౝኈߪ‫੹ޔ‬੗໪㓶ߦࠃࠆ⻠Ṷ‫ޟ‬๧ⷡߪ
ㆮવߔࠆ㧫̆୘ੱᏅ࡮୘૕Ꮕߣㅴൻ‫ߣޠ‬᡼㘺႐࡮ዷ␜ቶߩ⷗ቇߛߞߚ‫ޕ‬ෳട⠪ߪ 43 ฬ‫ޕ‬
3. ࠝ࡯ࡊࡦࠠࡖࡦࡄࠬ࡮ᄢቇ㒮ࠟࠗ࠳ࡦࠬ
ᄢቇߩቇㇱ↢ࠍ߅߽ߥኻ⽎ߣߒ‫ޔ‬ᄢቇ㒮ࠟࠗ࠳ࡦࠬࠍ౗ߨߚ 2011 ᐕᐲߩࠝ࡯ࡊࡦࠠࡖࡦࡄࠬࠍ‫ޔ‬2012 ᐕ 2 ᦬ 21
ᣣ‫ޔ‬22 ᣣߦ㐿௅ߒߚ‫ޕ‬ฦಽ⑼ߩᢎຬߦࠃࠆ⻠⟵‫ޔ‬ᚲౝ⷗ቇ‫ޔ‬ฦಽ⑼ᢎຬߣߩᙣ⺣ળ‫ߦࠄߐޔ‬ᄢቇ㒮↢࡮⎇ⓥຬ╬
߽ෳടߒߚᙣⷫળ߇߅ߎߥࠊࠇߚ‫ޕ‬ෳട⠪ߪ 26 ฬ‫ޕ‬
㧨ࡊࡠࠣ࡜ࡓ㧪
2 ᦬ 21 ᣣ㧔Ἣ㧕
ᚲ㐳⺑᣿
ฦಽ⑼ߩ⚫੺ 1
⹺⍮ቇ⠌ಽ⑼‫↢⊒⛔♽ޔ‬ಽ⑼‫␠ޔ‬ળ↢ᘒಽ⑼㧔↢ᘒ଻ో㧕‫␠ޔ‬ળ↢ᘒಽ⑼㧔␠ળㅴൻ㧕‫ࡓࡁࠥޔ‬ᄙ᭽ᕈಽ⑼
౉ቇ⹜㛎⺑᣿
ฦಽ⑼ߩ⚫੺ 2
ታ㛎േ‛⑼ቇಽ⑼‫ޔ‬ᕁ⠨⸒⺆ಽ⑼‫ޔ‬㜞ᰴ⣖ᯏ⢻ಽ⑼‫⛔ޔ‬ว⣖ࠪࠬ࠹ࡓಽ⑼‫ࡓࡁࠥޔ‬ㅴൻಽ⑼‫ޔ‬ㅴൻᒻᘒಽ⑼
ᙣⷫળ㧔ᄕ㘩ࠍ౗ߨߚ┙㘩ᒻᑼߩᙣⷫળߢ‫ޔ‬ᢎຬ߿ᄢቇ㒮↢ߣߩࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦࠍ࿑ߞߚ㧕
2 ᦬ 22 ᣣ㧔᳓㧕
ᚲౝ⷗ቇ
ಽ⑼ߏߣߩᙣ⺣ 1
ಽ⑼ߏߣߩᙣ⺣ 2
㧔ᢥ⽿㧦ᣧᎹᢅਯ㧕
㧙㧙
Ω. ࠽࡚ࠪ࠽࡞ࡃࠗࠝ࡝࠰࡯ࠬࡊࡠࠫࠚࠢ࠻㧔NBR㧕
1. ࠽࡚ࠪ࠽࡞ࡃࠗࠝ࡝࠰࡯ࠬࡊࡠࠫࠚࠢ࠻㧔࠾ࡎࡦࠩ࡞㧕ߩᵴേ
ᐔᚑ 19 ᐕᐲࠃࠅ‫ޔ‬5 ᐕ⸘↹ߢ╙ 2 ᦼ࠽࡚ࠪ࠽࡞ࡃࠗࠝ࡝࠰࡯ࠬࡊࡠࠫࠚࠢ࠻㧔NBRP㧕߇㐿ᆎߐࠇߚ‫ޕ‬NBRP ߪ
ታ㛎⎇ⓥߩࡕ࠺࡞ߣߥࠆࡑ࠙ࠬ߿࡜࠶࠻߿࡚࡚ࠪ࠙ࠫ࠙ࡃࠛߥࠄ߮ߦࠗࡀ߿ࠪࡠࠗ࠽࠭࠽╬ߩേᬀ‛‫ߦ߮ࠄߥޔ‬
ES ⚦⢩ߥߤߩ⚦⢩ᩣࠍ฽߼ߚࡃࠗࠝ࡝࠰࡯ࠬߩ߁ߜ‫ޔ‬࿖߇ᚢ⇛⊛ߦᢛ஻ࠍߔߔ߼ࠆ 22 ઙ߇ਛᩭ⊛᜚ὐᢛ஻ࡊࡠࠣ
࡜ࡓߣߒߡផㅴߐࠇߡ޿ࠆ‫⎇ࠬࡦࠛࠗࠨࡈࠗ࡜ޟߦߟ৻ߩ߆ߥߩߘޕ‬ⓥ↪࠾ࡎࡦࠩ࡞ߩ㘺⢒࡮❥ᱺ࡮ଏ⛎‫ޕࠆ޽߇ޠ‬
⥄ὼ⑼ቇ⎇ⓥᯏ᭴↢ℂቇ⎇ⓥᚲ߇ਛᩭᯏ㑐‫ޔ‬㔤㐳㘃⎇ⓥᚲ߇ಽᜂᯏ㑐ߣߒߡ‫࠰࡝ࠝࠗࡃ࡮࡞ࠩࡦࡎ࠾ޟࠆࠁࠊ޿ޔ‬
࡯ࠬ㧔NBR㧕‫ࠍޠ‬ផㅴߒߡ޿ࠆ‫ޕ‬NBR ߪ∛ේᓸ↢‛ቇ⊛ߦ቟ోߢ‫ޔ‬ஜᐽߥ⎇ⓥ↪࠾ࡎࡦࠩ࡞ࠍᐕ㑆 200 㗡⒟ᐲଏ⛎
ߔࠆ૕೙ߩ⏕┙ࠍ⋡ᮡߣߒߡ޿ࠆ‫ޕ‬
㔤㐳㘃⎇ⓥᚲߢߪቭᨋࠠࡖࡦࡄࠬ㧔╙ 1 ࠠࡖࡦࡄࠬ㧕߆ࠄ⋥✢〒㔌ߢ⚂ 1 ࠠࡠ᧲ߦ૏⟎ߔࠆༀᏧ㊁࿾඙ߦ╙ 2
ࠠࡖࡦࡄࠬ㧔⚂ 76ha㧕ߣߒߡ‫ޔ‬ᄢဳࡊࡠࠫࠚࠢ࠻‫ࡦ࡚ࠪ࡯࠹ࠬࠬ࡯࠰࡝࠴࡯ࠨ࡝ޟ‬㧔RRS㧕
‫ࠍޠ‬ᐔᚑ 19 ᐕᐲߦ㐿⸳
ߒߚ‫ᧄޕ‬੐ᬺߪⅣႺ౒ሽဳᄢဳ᡼㘺႐ࠍ⸳⟎ߒ‫␠ߩ࡞ࠩࡦࡎ࠾ޔ‬ળⴕേ╬ߩⷰኤ⎇ⓥߦ⾗ߔࠆߣߣ߽ߦ‫ޔ‬ᣂߚߥ⎇
ⓥ↪࠾ࡎࡦࠩ࡞ࠍഃ಴࡮❥ᱺ࡮⢒ᚑߒ‫ోޔ‬࿖ߩ⎇ⓥᯏ㑐ߦଏ⛎ߔࠆߎߣࠍ⋡⊛ߣߔࠆ‫ޕ‬ⅣႺߣߩ౒ሽࠍ㊀ⷞߔࠆᧄ
ࠠࡖࡦࡄࠬߢߪ‫ޔ‬ᢝ࿾ౝߩᬀ↢ߩ଻ోߣ‫ޔ‬ឃ᳓ߩಣℂߦਁోࠍᦼߒߡ޿ࠆ‫ߦߊߣޕ‬ᳪ᳓ߪ BOD ߢ 5ppm એਅߦಣ
ℂᓟ‫ޔ‬᡼㘺႐߳ㆶరᢔᏓߒ‫ޔ‬ᢝ࿾ᄖߦߪ಴ߐߥ޿ࠪࠬ࠹ࡓࠍ᭴▽ߒߡ޿ࠆ‫ޕ‬ᄢဳ᡼㘺႐ߪ࠾ࡎࡦࠩ࡞ߩ㊁↢ߢߩ↢
ᕷⅣႺࠍౣ⃻ߔࠆ߽ߩߢ‫੹ޔ‬ᓟߩᄙ᭽ߥ㔤㐳㘃⎇ⓥߩផㅴߩᩭߣߥࠆ߽ߩߣᦼᓙߐࠇࠆ‫ޕ‬
RRS ߩᢛ஻ߦߣ߽ߥߞߡ NBR ߩಽᜂᯏ㑐ߣߒߡ‫ޟ‬ᄢဳ㘺⢒ᣉ⸳ߢߩ࠾ࡎࡦࠩ࡞ߩ❥ᱺ࡮⢒ᚑ੐ᬺ‫⺖ߩޠ‬㗴ࠍផ
ㅴߒ‫ోޔ‬࿖߳ߩ࠾ࡎࡦࠩ࡞ଏ⛎ࠍታ⃻ߔࠆߚ߼ߩ‫❥ޔ‬ᱺ⢒ᚑࠍㆀⴕߒߡ޿ࠆ‫ޕ‬ᐔᚑ 21 ᐕᐲߦᦨᓟߩᄢဳࠣ࡞࡯ࡊ
ࠤ࡯ࠫࠍ⸳⟎ߒ‫੍ޔ‬ቯߒߚᣉ⸳ᢛ஻ߪቢੌߒߚ‫ ߢࠇߎޕ‬1 ᑼ 2 ᧦ߩ᡼㘺႐߇ 3 ᑼ‫ޔ‬⢒ᚑ⥡ 1 ᫟‫ ߦࠄߐޔ‬3 ᫟ߩࠣ࡞
࡯ࡊࠤ࡯ࠫ߇ቢᚑߒ‫ޔ‬Უ⟲✚ᢙ 350 㗡ߩ࠾ࡎࡦࠩ࡞ߩ㘺⢒߇น⢻ߣߥߞߚ‫ޕ‬ᐔᚑ 21 ᐕᐲ߆ࠄ NBR ߩ⚻⾌߇⵬ഥ㊄
ൻߐࠇ቟ቯଏ⛎ߦะߌߚᬺോ߇ᧄᩰ⊛ߦታᣉߐࠇߡ޿ࠆ‫ ߢ߹ࠇߎޕ‬NBR ੐ᬺߪ․೎⚻⾌ࡊࡠࠫࠚࠢ࠻㧔RRS㧕ߣ
⋧੕ᛔഥ੐ᬺߣߒߡ߅ߎߥࠊࠇߡ߈ߚ߇‫ޔ‬RRS ࡊࡠࠣ࡜ࡓߪ 22 ᐕᐲߢ⚳ੌߒߚ‫ޕ‬21 ᐕᐲߣ 22 ᐕᐲߩ⚻⾌ߢ⎇ⓥ
ၮ⋚ߩផㅴߣ NBR ߩⷙᮨ᜛ᄢߩ૏⟎ߠߌߣߒߡ‫ޔ‬ቭᨋ࿾඙ߦߘࠇߙࠇ 1 ᫟ߩࠣ࡞࡯ࡊࠤ࡯ࠫࠍ⸳⟎ߒߚ‫ޕ‬
2010 ᐕᐲߣห᭽‫ޔ‬
‫ⴊޟ‬ዊ᧼ᷫዋ∝‫ߦޠ‬㑐ㅪߒߡಣಽߐࠇߚേ‛߇⋧ᒰᢙ޿ߚߚ߼‫ޔ‬Უ⟲✚ᢙߪ 228 㗡߹ߢᷫዋߒ
ߚ‫৻ޕ‬ᣇߢ‫⧯ޔ‬㦂ߛߞߚᲣ⟲ߩᚑᾫ߇ㅴࠎߛߚ߼❥ᱺߪ㗅⺞ߦㅴࠎߢ߅ࠅ‫ޔ‬2011 ᐕᐲߩ಴↢࡮⢒ᚑ୘૕ߪ 48 㗡ߣ
ߥߞߡ޿ࠆ‫ޕ‬2010 ᐕᐲߪ‫ⴊޔ‬ዊ᧼ᷫዋ∝ߩᓇ㗀ߢ㔤㐳㘃⎇ⓥᚲ߆ࠄߩଏ⛎ࠍ஗ᱛߒߚ߇‫ޔ‬2011 ᐕᐲߩଏ⛎ᢙߪ 41
㗡ߦ㆐ߒ‫⥄ߡ߼ߓߪޔ‬ὼ⑼ቇ⎇ⓥᯏ᭴߆ࠄߩଏ⛎ಽࠍ฽߼ߚ NBR ࠾ࡎࡦࠩ࡞✚ଏ⛎ᢙߩඨᢙࠍභ߼ࠆߦ⥋ߞߚ‫ޕ‬
ᐕᐲᧃߩ⢒ᚑ⟲✚ᢙߪ 122 㗡ߣߥߞߡ߅ࠅ‫⃻ޔ‬࿷ว⸘ 350 㗡ߩ NBR ࡊࡠࠫࠚࠢ࠻↪࠾ࡎࡦࠩ࡞ࠍ㘺㙃ߒߡ޿ࠆ‫ޕ‬
2010 ᐕᐲߦታᣉߒߚੱ㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ࠲࡯ࠍਛᔃߣߒߚ⎇ⓥߢ‫ޔ‬㔤㐳㘃⎇ⓥᚲߦ߅ߌࠆ‫ⴊޟ‬ዊ᧼ᷫዋ∝‫ޠ‬
ߪࠨ࡞࡟࠻ࡠ࠙ࠗ࡞ࠬ 4 ဳ߇‫⥄ޔ‬ὼ⑼ቇ⎇ⓥᯏ᭴ߦ߅ߌࠆ‫ⴊޟ‬ዊ᧼ᷫዋ∝‫ ࠬ࡞ࠗ࠙ࡠ࠻࡟࡞ࠨߪޠ‬5 ဳ߇ේ࿃ߢ޽
ࠆߎߣ߇᣿ࠄ߆ߣߥߞߚ‫ޕ‬2011 ᐕ 8 ᦬ࠃࠅ NBR ࡊࡠࠫࠚࠢ࠻ߣߒߡ․ቯഥᢎࠍ㓹↪ߒ‫ੱޔ‬㘃ㅴൻࡕ࠺࡞⎇ⓥ࠮ࡦ
࠲࡯‫ޔ‬੩ㇺᄢቇ࠙ࠗ࡞ࠬ⎇ⓥᚲߣㅪ៤ߒߡ‫ⴊޟ‬ዊ᧼ᷫዋ∝‫∝⊒ߩޠ‬ᯏᐨߦߟ޿ߡߩ⎇ⓥࠍㅴ߼ࠆߣߣ߽ߦ‫ޔ‬ᬌᩏᴺ࡮
⸻ᢿᴺߩᡷ⦟ࠍⴕߞߡ޿ࠆ‫ޕ‬
㧔ᢥ⽿㧦ጟᧄቬ⵨㧕
2. ࠽࡚ࠪ࠽࡞ࡃࠗࠝ࡝࠰࡯ࠬࡊࡠࠫࠚࠢ࠻㧔GAIN㧕ߩᵴേ
GAIN㧦ᄢဳ㘃ੱ₎ᖱႎࡀ࠶࠻ࡢ࡯ࠢߩዷ㐿
ᧄ੐ᬺߪ‫ޔ‬ᐔᚑ 14 ᐕᐲߦᢥㇱ⑼ቇ⋭ߩਥዉߢ⊒⿷ߒߚ࠽࡚ࠪ࠽࡞ࡃࠗࠝ࡝࠰࡯ࠬࡊࡠࠫࠚࠢ࠻ߩ৻Ⅳߢ޽ࠆ‫ޕ‬
⎇ⓥ㐿⊒ᣉ⸳౒↪╬ଦㅴ⾌⵬ഥ㊄㧔࠽࡚ࠪ࠽࡞ࡃࠗࠝ࡝࠰࡯ࠬࡊࡠࠫࠚࠢ࠻㧕੐ᬺߦࠃࠆ߽ߩߢ‫⵬ޔ‬ഥ੐ᬺߩᱜᑼ
ฬ⒓ߪ‫ޔ‬
‫ޟ‬ᖱႎ⊒ା૕೙ߩᢛ஻ߣࡊࡠࠫࠚࠢ࠻ߩ✚ว⊛ផㅴ‫ޠ‬
㧔ᄢဳ㘃ੱ₎ᖱႎࡀ࠶࠻ࡢ࡯ࠢߩዷ㐿㧕ߢ޽ࠆ‫⧷ޕ‬ᢥ
ฬ⒓ࠍ Great Ape Information Network‫⧷ޔ‬ᢥ⇛⒓ࠍ GAIN ߣߔࠆ‫ޕ‬
GAIN ߪ‫ޔ‬ᐔᚑ 14 ᐕᐲߩࡈࠖ࡯ࠫࡆ࡝࠹ࠖ࡯⎇ⓥߦᆎ߹ߞߡ‫ޔ‬࿖ౝߩᄢဳ㘃ੱ₎ߩᖱႎࠍᢛ஻ߒߡ߈ߚ‫ࠇߎޕ‬
߹ߢߦᣣᧄ࿖ౝߩ㘃ੱ₎ 3 ⒳㧔࠴ࡦࡄࡦࠫ࡯࡮ࠧ࡝࡜࡮ࠝ࡜ࡦ࠙࡯࠲ࡦ㧕ߩో୘૕ߩ⊓㍳ࠍߒߚ‫ޕ‬Ფᣣ㓐ᤨ‫ޔ‬಴↢࡮
ᱫ੢࡮⒖☋╬ߩᖱႎߦߟ޿ߡ‫ᦨޔ‬ᣂ ߦᦝᣂߒߡ޿ࠆ‫ޔߪࠇߎޕ‬࿖┙ㆮવቇ⎇ⓥᚲߩࡃࠗࠝ࡝࠰࡯ࠬߦ㑐ߔࠆᖱႎ
ߩ࠙ࠚࡉࠨࠗ࠻ߢ৻⥸ߦ౏㐿ߐࠇߡ޿ࠆ‫ޕ‬HP ෳᾖ㧔http://www.shigen.nig.ac.jp/gain/index.jsp㧕
GAIN ੐ᬺߪ‫ޔ‬ᐔᚑ 23 ᐕᐲ߽‫ޔ‬ᓥ᧪ߤ߅ࠅ‫ޔ‬੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲߣ㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯ߩਔㇱዪߩ౒หㆇ
༡੐ᬺߣ૏⟎ߠߌߚ‫ޕ‬ᐔᚑ 23 ᐕᐲߦភ⟎ߐࠇߚ੤ઃ㊄㗵ߪᐕ㗵 960 ਁ౞ߢ೨ᐕᐲߣห㗵ߢ޽ࠆ‫⪭ޕ‬ว⍮⟤‫ޔ‬ᛂ⿧
ਁ༑ሶߩ 2 ฬߩ⎇ⓥຬࠍ㓹↪ߒߡታോߦ޽ߚߞߡ޿ߚߛ޿ߚ‫ⷫޔߚ߹ޕ‬੐ᬺߢ޽ࠆ‫ޟ‬ᖱႎ‫⛔ࠍޠ‬᜝ߔࠆ࿖┙ㆮવቇ
⎇ⓥᚲ㧔ዊේ㓶ᴦᚲ㐳㧕ߥࠄ߮ߦ‫⽿ߩߘޔ‬છ⠪ߢ޽ࠆ♽⛔ᖱႎ⎇ⓥቶߩጊፒ↱♿ሶవ↢߆ࠄߪ‫ߦߨߟޔ‬ෘ޿ߏᡰេ
ࠍ޿ߚߛ޿ߚ‫ߡߒ⸥ޕ‬ᗵ⻢ߒߚ޿‫ޕ‬GAIN ੐ᬺߩౝኈߪએਅߩߣ߅ࠅߢ޽ࠆ‫ޕ‬੐ᬺઍ⴫⠪㧔⺖㗴▤ℂ⠪㧕ߪ᧻ᴛື
㇢‫⚻ޔ‬ℂᜂᒰ⠪ߪ਄၂ᵏᶈ‫ޔ‬੐ᬺෳട⠪ߪ㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯ߩદ⼱ේ৻߿ᐘፉม㇢߶߆⚂ 20 ฬ‫ޔ‬੐ᬺදജ⠪
ߪർ㉿ᄢቇߩศᎹᵏᒄ߿᧲੩ᄢቇߩ㐳⼱Ꮉኼ৻߶߆⚂ 15 ฬߢ޽ࠆ‫ޕ‬
㧙㧙
1) ᖱႎߩឭଏ
㘺⢒ᣉ⸳ߩ୘૕⟲ࡕ࠾࠲࡝ࡦࠣߦࠃࠅ‫ޔ‬ᄢဳ㘃ੱ₎ߩᱫ੢߿಴↢ߦᔕߓߡㆡቱ࠺࡯࠲ࡌ࡯ࠬࠍᦝᣂߒߚ‫ޕ‬ᐔᚑ
23 ᐕᐲᧃߩᐔᚑ 24 ᐕ 2 ᦬ 20 ᣣ⃻࿷ߢ‫࡯ࠫࡦࡄࡦ࠴ޔ‬330 ୘૕㧔53 ᣉ⸳㧕
‫ ࡜࡝ࠧޔ‬23 ୘૕㧔10 ᣉ⸳㧕
‫࡯࠙ࡦ࡜ࠝޔ‬
࠲ࡦ 53 ୘૕㧔22 ᣉ⸳㧕߇࿖ౝߢ㘺㙃ߐࠇߡ޿ࠆ‫ޕ‬
ᐔᚑ 23 ᐕᐲߪ࠴ࡦࡄࡦࠫ࡯ߩᱫ੢ 4 ઙ‫ߩࡦ࠲࡯࠙ࡦ࡜ࠝޔ‬ᱫ੢ 2 ઙ‫ߩ࡯ࠫࡦࡄࡦ࠴ޔ‬಴↥ 4 ઙ‫ߩ࡜࡝ࠧޔ‬಴↥
1 ઙ‫ߩ࡯ࠫࡦࡄࡦ࠴ޔ‬⒖േ 1 ઙ߇޽ߞߚ‫ߚ߹ޕ‬ᣂߚߦዊဳ㘃ੱ₎࠹࠽ࠟࠩ࡞ߦߟ޿ߡ߽ᖱႎ෼㓸ࠍߒߡ‫ࡌ࠲࡯࠺ޔ‬
࡯ࠬ߳ߩ⊓㍳૞ᬺࠍߔߔ߼ߡ޿ࠆ‫ޕ‬
2) ฦ୘૕ߩࠥࡁࡓ࡮ⴕേᖱႎ࡮ᒻᘒᖱႎߩ෼㓸
୘૕ߏߣߩ↢ᐕ᦬ᣣ߿ኅ♽ᖱႎߦട߃ߡ‫ޔ‬DNA ᖱႎ࡮ⴕേᖱႎߦߟ޿ߡ߽ᢛ஻ࠍߔߔ߼ߡ޿ࠆ‫ ࡞ࡃ࡯ࡠࠣޕ‬COE
ࡊࡠࠣ࡜ࡓ㧔↢‛ߩᄙ᭽ᕈߣㅴൻ⎇ⓥߩߚ߼ߩ᜚ὐᒻᚑࠥࡁࡓ߆ࠄ↢ᘒ♽߹ߢ㧦੩ㇺᄢቇ࡮㒙ᒻᷡ๺࡝࡯࠳࡯㧕
ߣදജߒߡ‫ޔ‬
‫ޟ‬㔤㐳㘃ࠥࡁࡓ࠺࡯࠲ࡌ࡯ࠬ‫ࠍޠ‬૞ᚑ࡮౏㐿ߒߡ޿ࠆ‫( ޕ‬http://www.gcoe.biol.sci.kyoto-u.ac.jp/pgdb/) ߹
ߚ 3 ᰴర㛽ᩰᮡᧄ㧔CT ↹௝㧕ࠍ GAIN ↱᧪ߩ㘃ੱ₎ 12 ୘૕ಽߦߟ޿ߡ‫࡞࠲ࠫ࠺ޟ‬ᒻᘒඳ‛㙚㧔㔤㐳㘃⎇ⓥᚲ⾗ᢱ
ᆔຬળ㧕
‫౏ߢޠ‬㐿ߒߡ޿ࠆ‫(ޕ‬http://www.pri.kyoto-u.ac.jp/dmm/WebGallery/)
3) 㘺⢒ᣉ⸳⸰໧ߦࠃࠆᖱႎ෼㓸
㘺⢒ᣉ⸳ࠍ⸰໧ߒߡ‫⋥ޔ‬ធߩ⋡ⷞߣᜂᒰ⠪ߣߩ㕙⺣ߢ‫ޔ‬ᖱႎ෼㓸ࠍ߅ߎߥߞߚ‫ޕ‬ᐔᚑ 23 ᐕᐲߪ‫ޔ‬ᕲ⾦਄㊁േ‛
࿦‫ޔ‬㣮ఽፉ⋵ᐔᎹേ‛౏࿦‫ޔ‬੩ㇺᏒേ‛࿦‫ޔ‬㊖〝Ꮢേ‛࿦‫ޔ‬ᾢᧄᏒേᬀ‛࿦‫ᧅ ޔ‬ᏻᏒ౞ጊേ‛࿦‫ޔ‬ᄤ₺ኹേ‛࿦‫ޔ‬
᧲ᱞേ‛౏࿦‫ޔ‬ฬฎደᏒ᧲ጊേᬀ‛࿦‫ޔ‬㊁Ძጊേ‛࿦‫ޔ‬ᮮᵿᏒ┙㊄ᴛേ‛࿦ࠍ⸰໧ߒߚ‫ޕ‬ᓟㅀߔࠆᾢᧄࠨࡦࠢ࠴ࡘ
ࠕ࡝߇ᐔᚑ 23 ᐕᐲߦ⊒⿷ߒߚ‫੹ޕ‬ᓟߪ‫࡯ࠫࡦࡄࡦ࠴ޔ‬㘺⢒ߩ࿖ౝ᜚ὐߢ޽ࠆหᣉ⸳ࠍ GAIN ੐ᬺߩၮゲߦߒߡ‫ޔ‬
࿖ౝᄖߩേ‛࿦╬ߣߩㅪ៤ࠍߐࠄߦផㅴߔࠆ‫ޕ‬
4) ᱫ૕↱᧪࡮↢૕↱᧪㧔㕖ଚⷅ㧕⾗ᢱߩ㈩Ꮣ
ᐔᚑ 23 ᐕᐲߪ‫ޔ‬ᄢဳ㘃ੱ₎ 9 ୘૕߆ࠄ㕖ଚⷅ⊛ߥᣇᴺߢߩ⹜࡮⾗ᢱߩ෼㓸ࠍ߅ߎߥ޿‫⎇ޔ‬ⓥ೑↪ࠍࠨࡐ࡯࠻ߒ
ߚ‫ޕ‬ᒰ⹥㘺⢒ᣉ⸳߆ࠄᱫ੢߿಴↥ᖱႎߩឭଏࠍฃߌ‫ࠍࠇߘޔ‬㔤㐳㘃⎇ⓥᚲ⾗ᢱᆔຬળߥࠄ߮ߦ౒ห೑↪ታⴕᆔຬળ
ߦᖱႎឭଏߔࠆࠪࠬ࠹ࡓ߇⏕┙ߒߚ‫ߡߞ߇ߚߒޕ‬ᖱႎឭଏߪ GAIN ੐ᬺߢ޽ࠅ‫ޔ‬ታ㓙ߩࠨࡦࡊ࡞෼㓸ߣ㈩ઃߪ‫ޔ‬ᐔ
ᚑ 22 ᐕᐲ⊒⿷ߩ‫౒ޟ‬ห೑↪࡮౒ห⎇ⓥ᜚ὐ೙ᐲ‫ߩߡߒߣޠ‬㔤㐳㘃⎇ⓥᚲߢㆇ↪ߔࠆߎߣߣߒߚ‫ޕ‬ᱫ੢୘૕ߩ೑↪
ࠍ౒ห೑↪࡮౒ห⎇ⓥ೙ᐲߣㅪ៤ߒߡଦㅴߔࠆ૕೙߇⏕┙ߒߟߟ޽ࠆ‫ޕ‬੐ᬺෳട⠪ߢ޽ࠆ‫᧛⷏ޔ‬೰‫ޔ‬ㇹᐽᐢ‫੹ޔ‬੗
໪㓶ߩ 3 ฬߩᢎຬࠄ߇ਛᔃߣߥߞߡ‫ޔ‬㕖ଚⷅ⾗ᢱߩ౒ห೑↪⎇ⓥࠍផㅴߒߚ‫ޕ‬
5) ᐢႎᵴേߣᚑᨐ౏㐿
╙ 27 ࿁㔤㐳㘃ቇળᄢળ‫ ╙ޔ‬34 ࿁ᣣᧄಽሶ↢‛ቇળᐕળ‫ ╙ޔ‬14 ࿁ SAGA ࠪࡦࡐࠫ࠙ࡓߢ‫⎇ޔ‬ⓥ⠪߿േ‛࿦㑐ଥ
⠪ߦ GAIN ੐ᬺߩᐢႎࠍߒߚ‫ޕ‬ᐔᚑ 22 ᐕᐲ߆ࠄߪ GAIN ߩ࠙ࠚࡉࠨࠗ࠻ߦ⧷⺆ ࠍㅊടߒߚ‫ޕ‬ᐔᚑ 23 ᐕᐲ߽ᒁ߈
⛯߈‫ޔ‬਎⇇ߦะߌߡᣣᧄߩᄢဳ㘃ੱ₎ߩᖱႎࠍ⊒ାߒߚ‫☨ޔ߅ߥޕ‬࿖ࠝ࡟ࠧࡦߢ㐿௅ߐࠇߚ࿖㓙ⅣႺࠛࡦ࡝࠶࠴ࡔ
ࡦ࠻ળ⼏ߦ⪭ว⎇ⓥຬ߇ෳടߒߡ‫ޔ‬ᶏᄖߩേ‛࿦㑐ଥ⠪ߣᗧ⷗੤឵ࠍ߅ߎߥߞߚ‫߇ࠊޔߚ߹ޕ‬࿖ߩ࠽࡚ࠪ࠽࡞ࡃࠗ
ࠝ࡝࠰࡯ࠬߩ޽ࠅᣇߘߩ߽ߩࠍᄢᚲ㜞ᚲ߆ࠄ⼏⺰ߔࠆ႐ߣߒߡ‫‛↢ޟ‬ㆮવ⾗Ḯᆔຬળ‫ޔࠅ޽߇ޠ‬ᐕᐲᧃߩᐔᚑ 24
ᐕ 3 ᦬ 27 ᣣߦ╙ 14 ࿁ߩળ⼏߇᧲੩ߢ㐿௅ߐࠇߚ‫ޕ‬ઍℂᆔຬߣߒߡ‫ޔ‬ᐔᚑ 24 ᐕᐲߩ㔤㐳㘃⎇ⓥᚲ㐳ࠍߟߣ߼ࠆߎ
ߣߦߥߞߚᐔ੗໪ਭ߇಴Ꮸߒߚ‫ޕ‬
6) ᾢᧄࠨࡦࠢ࠴ࡘࠕ࡝ߩ⊒⿷
࿖ౝߢ࠴ࡦࡄࡦࠫ࡯ࠍᦨ߽ᄙߊ㘺⢒ߒߡ޿ࠆᾢᧄߩ‫ޟ‬ਃ๺ൻቇ⎇ⓥᚲ࠴ࡦࡄࡦࠫ࡯࡮ࠨࡦࠢ࠴ࡘࠕ࡝࡮ቝ࿯
㧔CSU㧕‫ޔߪޠ‬ᐔᚑ 23 ᐕ 8 ᦬ 1 ᣣࠍ߽ߞߡ‫ޟ‬੩ㇺᄢቇ㊁↢േ‛࠮ࡦ࠲࡯ᾢᧄࠨࡦࠢ࠴ࡘࠕ࡝㧔KS㧕
‫ߦޠ‬⒖ⴕߒᡷฬ
ߐࠇߚ‫ޕ‬᳃㑆ߩකቇ⎇ⓥᣉ⸳ߛߞߚ߽ߩ߇‫ޔ‬੩ᄢ߇ㆇ༡ߔࠆࠨࡦࠢ࠴ࡘࠕ࡝ᣉ⸳ߣߥߞߡ޿ࠆ‫߅ߥޕ‬ਃ๺ൻቇ⎇ⓥ
ᚲߩෘᗧߢᐔᚑ 19 ᐕ 8 ᦬ 1 ᣣߦ⊒⿷ߒߚነ㒝⻠ᐳ‫␩⑔ޟ‬㐳ኼ⎇ⓥㇱ㐷‫߇ޠ‬ᐔᚑ 23 ᐕᐲ߽⛮⛯ߒߡ߅ࠅ‫ޔ‬ะߎ߁ߐ
ࠄߦ 21 ᐕ㑆ߩ⛮⛯߇⚂᧤ߐࠇߡ⽷᡽⊛ၮ⋚ߩ৻ㇱߣߥߞߡ޿ࠆ‫ޕ‬ᐔᚑ 23 ᐕᐲᧃߩ KS ߩ࠴ࡦࡄࡦࠫ࡯଻᦭୘૕ᢙ
ߪ 50 ୘૕ߢ޽ࠆ‫ޔ߅ߥޕ‬ᐔᚑ 24 ᐕ 5 ᦬ 15 ᣣߦ᳃㑆කቇ⎇ⓥᣉ⸳߆ࠄ 3 ୘૕߇ KS ߦዉ౉ߐࠇߚ‫ޔߡߞ߽ࠍࠇߎޕ‬
ᣣᧄߩකቇᗵᨴታ㛎࠴ࡦࡄࡦࠫ࡯ߪ࠯ࡠߦߥߞߚ‫ޕ‬1998 ᐕߩᤨὐߢ᳃㑆 3 ᣉ⸳ߦว⸘ 136 ୘૕ߩකቇᗵᨴታ㛎࠴
ࡦࡄࡦࠫ࡯߇޿ߚ‫ޕ‬14 ᐕ㑆߆߆ߞߡ‫ޔ‬ᣣᧄߩ࠴ࡦࡄࡦࠫ࡯ߦኻߔࠆකቇᗵᨴታ㛎ߪᑄ⛘ߐࠇߚ‫ߩߘޕ‬ㆊ⒟ߦ߅޿
ߡ‫ޔ‬GAIN ߪ‫ޔ‬୘૕ᖱႎࠍᢛ஻ߒ‫ޔ‬ᱫᓟ೑↪ࠍଦㅴߔࠆߎߣߢ↢૕೑↪ࠍᛥ೙ߒ‫ޔ‬කቇᗵᨴታ㛎ࠍߪߓ߼ߣߔࠆߔ
ߴߡߩଚⷅታ㛎ߩᑄ⛘ߦะߌߚਥⷐߥේേജߣߒߡᯏ⢻ߒߚߣ޿߃ࠆ‫ޕ‬
7) ╙ 3 ᦼߩ੍ቯ
࠽࡚ࠪ࠽࡞ࡃࠗࠝ࡝࠰࡯ࠬ੐ᬺߩ╙ 1 ᦼ㧔ᐔᚑ 14-18 ᐕᐲ㧕ߣ╙ 2 ᦼ㧔ᐔᚑ 19-23 ᐕᐲ㧕ࠍㅢߓߡቢᚑߒߚ㘺⢒
ᣉ⸳ߣ⎇ⓥ⠪ࠍ⚿߱ࡀ࠶࠻ࡢ࡯ࠢ߿୘૕ᖱႎ࠺࡯࠲ࡌ࡯ࠬࠍ⏕଻ߒߥ߇ࠄ‫ࠆߥࠄߐޔ‬లታࠍ߼ߑߔ‫ޔߚ߹ޕ‬ᖱႎࠍ
ၮߦߒߚ୘૕⟲▤ℂߩឭ⸒‫ޔ‬࿖㓙⊛ߥᖱႎ⊒ାࠍ߅ߎߥ߁ߎߣࠍ⋡ᮡߣߔࠆ‫ߩߢ߹ࠇߎޔߕ߹ޕ‬ദജࠍ⛮⛯ߒߡ‫ޔ‬
ᣣᧄ߇଻᦭ߒߡ߈ߚ㘃ੱ₎ 1000 ୘૕㧔↢ሽ୘૕ߣᱫ੢୘૕ࠍ޽ࠊߖߚ✚ᢙ㧕ࠍ⿧߃ࠆ࿕᦭ߩ↢‛⾗Ḯᖱႎ࠺࡯࠲
ࡌ࡯ࠬߩలታࠍ࿑ࠆ‫ౕޕ‬૕⊛ߦߪ‫ޔ‬
‫ᦨޟ‬ᣂᖱႎߩㅊട࡮ᦝᣂ‫ޠ‬
‫ޟ‬୘૕ᖱႎߩలታ‫ޠ‬
‫ޟ‬୘૕⟲ߩ▤ℂߩឭ⸒‫ޠ‬
‫ޟ‬࿖㓙ൻߩ
ផㅴ‫ ߩޠ‬4 ߟߩ૞ᬺࠍ߅ߎߥ߁‫ޕ‬ᄢဳ㘃ੱ₎ߩ୘૕ᖱႎߩᢛ஻ߢߪᣣᧄ߇਎⇇⊛ߦ⷗ߡ߽৻ᱠㅴࠎߛ⁁ᴫߦ޽ࠆ‫ޕ‬
᰷☨ߣߩද⺞ࠍ࿑ࠅ‫ޔ‬㖧࿖߿ࡑ࡟࡯ࠪࠕߥߤㄭ㓞⻉࿖ߣߩ޽޿ߛߢᄢဳ㘃ੱ₎ᖱႎࠍ੤឵ߒ‫ࠍࠞ࡝ࡈࠕޔ‬ೋ߼ߣߔ
㧙㧙
ࠆ↢ᕷ࿾࿖ߣߩද⺞ࠍ࿑ࠆ‫ޕ‬ᖱႎߣ޿߁㕙ߢᢛ஻ߩㅴࠎߛ‫ޔ‬ᣣᧄ⁛⥄ߢ਎⇇⊛ߦ߽㘃଀ߩߥ޿࡙࠾࡯ࠢߥ↢‛⾗Ḯ
ᖱႎߩឭଏࠍ╙ 3 ᦼߩ GAIN ੐ᬺߣߒߡ⋡ᜰߔ‫ޕ‬
ᐔᚑ 23 ᐕᐲߩਥⷐߥ಴ ‛ߪᰴߩㅢࠅ‫ޕ‬
Morimura N, Idani G, Matsuzawa T (2011) The First chimpanzee sanctuary in Japan: An attempt to care for the ‘‘Surplus’’ of
biomedical research, American Journal of Primatology, 73: 226-232.
㧔ᢥ⽿㧦᧻ᴛື㇢㧕
㧙㧙
Ϊ. ౒ห೑↪⎇ⓥ
1. ᭎ⷐ
ᐔᚑ 23 ᐕᐲߩ౒ห೑↪⎇ⓥߩ⎇ⓥ⺖㗴ߪએਅߩ 4 ߟߩࠞ࠹ࠧ࡝࡯ߢታᣉߐࠇߡ޿ࠆ‫ޕ‬
A ⸘↹⎇ⓥ
B ৻⥸୘ੱ⎇ⓥ
C ৻⥸ࠣ࡞࡯ࡊ⎇ⓥ
D 㓐ᤨ൐㓸⎇ⓥ
౒ห೑↪⎇ⓥߪ‫ޔ‬ᤘ๺ 57 ᐕᐲߦ‫⎇↹⸘ޟ‬ⓥ‫⎇↱⥄ޟߣޠ‬ⓥ‫ ߩޠ‬2 ߟߩ⎇ⓥ⺖㗴ߢታᣉߐࠇ‫ޔ‬ᤘ๺ 62 ᐕᐲ߆ࠄߪ
‫⾗ޟ‬ᢱឭଏ‫ޠ‬㧔ᐔᚑ 14 ᐕᐲ߆ࠄ‫ޟ‬ᣉ⸳೑↪‫ߣޠ‬ฬ⒓ࠍᄌᦝ‫ߦࠄߐޔ‬ᐔᚑ 20 ᐕᐲ߆ࠄ‫ޟ‬㓐ᤨ൐㓸⎇ⓥ‫ߣޠ‬ฬ⒓ࠍᄌ
ᦝ㧕ࠍ‫ޔ‬ᐔᚑ 6 ᐕᐲ߆ࠄߪ‫ޟ‬ᚲᄖଏ⛎‫ޠ‬㧔ᐔᚑ 14 ᐕᐲ߆ࠄ‫ޟ‬ᚲᄖ⾉ਈ‫ߣޠ‬ฬ⒓ࠍᄌᦝߒ‫ޔ‬ᐔᚑ 15 ᐕᐲߢ⚳ੌ㧕߇
ታᣉߐࠇߚ‫ߦࠄߐޕ‬ᐔᚑ 23 ᐕᐲ߆ࠄߪ‫⎇↱⥄ޟ‬ⓥ‫⥸৻ޟࠍޠ‬୘ੱ⎇ⓥ‫⎇ࡊ࡯࡞ࠣ⥸৻ޟߣޠ‬ⓥ‫ߦޠ‬ಽߌ‫ޔ‬4 ߟߩ
ࠞ࠹ࠧ࡝࡯ߣߥࠅ⃻࿷ߦ⥋ߞߡ޿ࠆ‫⎇ߩࠇߙࠇߘޕ‬ⓥ⺖㗴ߩ᭎⇛ߪએਅߩߣ߅ࠅߢ޽ࠆ‫ޕ‬
‫⎇↹⸘ޟ‬ⓥ‫⎇ᧄޔߪޠ‬ⓥᚲផㅴ⠪ߩડ↹ߦၮߠ޿ߡ౒ห೑↪⎇ⓥ⠪ࠍ౏൐ߔࠆ߽ߩߢ‫ޔ‬୘‫⎇↹⸘ޟߩޘ‬ⓥ‫ ߪޠ‬2
㨪3 ᐕߩᦼ㑆ౝߦ⚳ੌߒ‫ޔ‬ᚑᨐࠍ߹ߣ߼‫ޕ߁ⴕࠍ⴫౏ޔ‬
‫⥸৻ޟ‬୘ੱ⎇ⓥ‫⎇ࡊ࡯࡞ࠣ⥸৻ޟ߮ࠃ߅ޠ‬ⓥ‫⎇↹⸘ޟޔߪޠ‬ⓥ‫⹥ߦޠ‬ᒰߒߥ޿ࡊࡠࠫࠚࠢ࠻ߢ‫ޔ‬ᔕ൐⠪㧔⎇ⓥᚲ
ᄖߩⶄᢙߩ⎇ⓥቶ߆ࠄߩ౒หឭ᩺ߦࠃࠆ߽ߩߪ৻⥸ࠣ࡞࡯ࡊ⎇ⓥ㧕ߩ⥄↱ߥ⌕ᗐߣ⸘↹ߦၮߠ߈‫ޔ‬ᚲౝኻᔕ⠪ߩද
ജࠍᓧߡ‫ᦼ⛯⛮ޔ‬㑆 3 ᐕࠍ⋡ಣߦ౒ห⎇ⓥࠍታᣉߔࠆ‫ޕ‬
‫ޟ‬㓐ᤨ൐㓸⎇ⓥ‫⾗ޔߪޠ‬ᢱ㧔૕ᶧ‫╭ޔེ⤳ޔ‬⡺‫ޔ‬Ძ⊹‫ޔ‬ᱤ‎࡮㛽ᩰ‫ޔ‬ឃᴭ‛╬㧕ࠍឭଏߒߡⴕࠊࠇࠆ౒ห⎇ⓥߢ
޽ࠆ‫ޕ‬
ߥ߅‫ޔ‬ᐔᚑ 22 ᐕᐲ߆ࠄ‫ޔ‬㔤㐳㘃⎇ⓥᚲߪᓥ᧪ߩో࿖౒ห೑↪ߩ㒝⟎⎇ⓥᚲ߆ࠄ‫౒ޟ‬ห೑↪㨯౒ห⎇ⓥ᜚ὐ‫ߥߣޠ‬
ࠅ‫౒ޔ޿઻ߦࠇߎޔ‬ห೑↪࡮౒ห⎇ⓥ߽᜚ὐ੐ᬺߣߒߡㅴ߼ࠄࠇࠆߎߣߣߥߞߚ‫ޕ‬
ട߃ߡ‫᧲ޔ‬ർ࿾ᣇᄥᐔᵗᴒ࿾㔡߅ࠃ߮㑐ㅪ੐⽎ߦࠃࠅ⎇ⓥ╬ߩ⛮⛯߇࿎㔍ߦߥߞߚᣇࠍኻ⽎ߦ‫ޔ‬㔡ἴ㑐ㅪ൐㓸ᨒ
߽⥃ᤨߦ⸳ቯߒߚ‫ޕ‬
ᐔᚑ 23 ᐕᐲߩ⸘↹⺖㗴‫ޔ‬ᔕ൐ਗ߮ߦណᛯ⁁ᴫߪએਅߩߣ߅ࠅߢ޽ࠆ‫ޕ‬
(1) ⸘↹⺖㗴
1㧚ᣥ਎⇇ࠩ࡞ߩᄌ⇣ᕈߣㅴൻߦ㑐ߔࠆᄙ㕙⊛ࠕࡊࡠ࡯࠴
ታᣉ੍ቯᐕᐲ ᐔᚑ21ᐕᐲ㨪23ᐕᐲ
⺖㗴ផㅴ⠪㧦㜞੗ᱜᚑ㧘⷏᧛೰㧘ᳯᧁ⋥ሶ㧘ࡑࠗࠢ࡮ࡂࡈࡑࡦ
ᣥ਎⇇ࠩ࡞㘃㧔ࠝ࠽ࠟࠩ࡞⑼㧕ߪࠝ࠽ࠟࠩ࡞੝⑼ߣࠦࡠࡉࠬ੝⑼ߩੑߟߩࠣ࡞࡯ࡊ߆ࠄߥࠆ߇‫ߩߘޔ‬ᒻᘒ࡮
㘩ᕈ࡮ⴕേࡄ࠲࡯ࡦߦߪᄢ߈ߥᄌ⇣߇ሽ࿷ߔࠆ‫ߚߞ޿߁ߎޕ‬ᣥ਎⇇ࠩ࡞㘃ߩᄙ᭽ᕈߣߘߩㅴൻߦ㑐ߒߡᒻᘒቇ
߿ห૏૕ಽᨆߥߤߩᄙ㕙⊛ߥಽ㊁࡮ᚻᴺࠍ↪޿ߡࠕࡊࡠ࡯࠴ߔࠆ‫ޕ‬
2㧚ⴕേ․ᕈࠍᡰ㈩ߔࠆࠥࡁࡓၮ⋚ߣ⣖ᯏ⢻ߩ⸃᣿
ታᣉ੍ቯᐕᐲ ᐔᚑ 23 ᐕᐲ㨪24 ᐕᐲ
⺖㗴ផㅴ⠪㧦㜞↰᣽ᒾ㧘ਛ᧛స᮸㧘ᐔ੗໪ਭ㧘੹੗໪㓶㧘ㇹᐽᐢ
ࡅ࠻ߩ♖␹࡮␹⚻∔ᖚߦ㑐ㅪߔࠆ࡝ࠬࠢㆮવሶߩ⊒⃻ࠍࡑࠞࠢࠩ࡞╬ߢ✂⟜⊛ߦᬌ⚝ߒ‫⥄ޔ‬ὼ⊒↢⊛ㆮવሶ
ᄌ⇣ࡕ࠺࡞ߩត⚝ߣੱὑ⊛ㆮવሶᡷᄌࡕ࠺࡞ߩ㐿⊒ࠍߣ߅ߒߡ‫ⴕޔ‬േ․ᕈࠍ᳿ቯߔࠆࠥࡁࡓ‫߇ࡓࡁࠥޔ‬೙ᓮߔ
ࠆ⣖ᯏ⢻‫ޔ‬⣖ᯏ⢻߇ⷙቯߔࠆⴕേ․ᕈࠍ᣿ࠄ߆ߦߔࠆ‫ޕ‬
(2) ᔕ൐ਗ߮ߦណᛯ⁁ᴫ
ᐔᚑ 23 ᐕᐲߪߎࠇࠄߩ⎇ⓥ⺖㗴ߦߟ޿ߡ‫ޔ‬103 ઙ㧔246 ฬ㧕ߩᔕ൐߇޽ࠅ‫౒ޔ‬ห೑↪ታⴕᆔຬળ㧔੹੗໪㓶‫ޔ‬ฎ
⾐┨ᒾ‫↰ޔ‬ਛᵗਯ‫ޔ‬ㄞᄢ๺‫ޔ‬ች࿾㊀ᒄ㧕ߦ߅޿ߡណᛯේ᩺ࠍ૞ᚑߒ‫౒ޔ‬ห೑↪ኾ㐷ᆔຬળ㧔ᐔᚑ 23 ᐕ 2 ᦬ 25 ᣣ㧕
ߩክ⼏࡮᳿ቯࠍ⚻ߡ‫ޔ‬᜚ὐㆇ༡ද⼏ળ㧔ᐔᚑ 23 ᐕ 3 ᦬ 7 ᣣ㧕ߢੌᛚߐࠇߚ‫ޕ‬
ߘߩ⚿ᨐ‫ޔ‬102 ઙ㧔245 ฬ㧕߇ណᛯߐࠇߚ‫ޕ‬
ฦ⺖㗴ߦߟ޿ߡߩᔕ൐࡮ណᛯ⁁ᴫߪએਅߩߣ߅ࠅߢ޽ࠆ‫ ޕ‬
⺖ 㗴
⸘↹⎇ⓥ 1
⸘↹⎇ⓥ 2
৻⥸୘ੱ⎇ⓥ
৻⥸ࠣ࡞࡯ࡊ⎇ⓥ
㓐ᤨ൐㓸⎇ⓥ
㔡ἴ㑐ㅪ
ว ⸘
ᔕ ൐
5ઙ
( 7 ฬ)
5ઙ
( 12 ฬ)
63 ઙ
( 135 ฬ)
8ઙ
( 37 ฬ)
21 ઙ
( 48 ฬ)
1ઙ
( 7 ฬ)
103 ઙ
( 246 ฬ)
㧙㧙
ណ ᛯ
5ઙ
( 7 ฬ)
5ઙ
( 12 ฬ)
62 ઙ
( 134 ฬ)
8ઙ
( 37 ฬ)
21 ઙ
( 48 ฬ)
1ઙ
( 7 ฬ)
102 ઙ
( 245 ฬ)
2. ⎇ⓥᚑᨐ
(1) ⸘↹⎇ⓥ
A-1 ⴕേ․ᕈࠍᡰ㈩ߔࠆࠥࡁࡓၮ⋚ߣ⣖ᯏ⢻ߩ⸃᣿
ዊᨋ๺ੱ㧔⑔ፉකᄢ࡮ක㧕 ᚲౝኻᔕ⠪㧦㜞↰᣽ᒾ
ᚒ‫ޔߦߢ߹ࠇߎޔߪޘ‬㜞↰᣽ᒾᢎ᝼ߣߩ౒ห⎇ⓥߦࠃࠅ‫⾰ࠢࡄࡦ࠲♧ࠬ࡞ࠗ࠙∛›⁅ޔ‬㧔RV-G㧕ࠍ↪޿ߚ࡟ࡦ
࠴࠙ࠗ࡞ࠬࡌࠢ࠲࡯ࠍ㐿⊒ߒ‫ߞߍޔ‬ᱤ㘃߅ࠃ߮㔤㐳㘃ߩ⣖ౝߦ߅޿ߡㅒⴕᕈㆮવሶዉ౉߇น⢻ߥߎߣࠍ␜ߒߚ‫ߐޕ‬
ࠄߦ‫ޔ‬RV-G ߩ⚦⢩ᄖ࡮⤑⽾ㅢ࠼ࡔࠗࡦߣ᳓ᵃᕈญౝἻ࠙ࠗ࡞ࠬ㧔VSV-G㧕ߩ⚦⢩ౝ࠼ࡔࠗࡦ߆ࠄ᭴ᚑߐࠇࠆⲢว
♧࠲ࡦࡄࠢ⾰ B ဳ㧔FuG-B㧕ࠍ㐿⊒ߒ‫ޔ‬ㅒⴕᕈㆮવሶዉ౉ߩല₸ࠍᩰᲑߦะ਄ߐߖࠆߎߣߦᚑഞߒߚ‫⎇ᧄޕ‬ⓥߢߪ‫ޔ‬
RV-G ⚦⢩ᄖ࠼ࡔࠗࡦ N ᧃ┵㗔ၞߣ VSV-G ⚦⢩ᄖ࠼ࡔࠗࡦ C ᧃ┵㗔ၞࠍ฽ߺ‫ޔ‬VSV-G ⤑⽾ㅢ࡮⚦⢩ౝ࠼ࡔࠗࡦߦ
ㅪ⚿ߒߚⲢว♧࠲ࡦࡄࠢ⾰ C ဳ㧔FuG-C㧕ࠍ૞⵾ߒ‫ޔ‬ᣂߒ޿࡟ࡦ࠴࠙ࠗ࡞ࠬࡌࠢ࠲࡯ࠍ㐿⊒ߒߚ‫ޕ‬FuG-C ࡌࠢ࠲࡯
ࠍࠨ࡞✢᧦૕ߦᵈ౉ߒ‫ޔ‬⣖ౝߢߩㆮવሶዉ౉ߩ․ᕈߣല₸ࠍᬌ⸛ߒߚ‫ޔߪ࡯࠲ࠢࡌࡦࡎޕ‬㤥⾰✺ኒㇱ‫ޔ‬ᄢ⣖⊹⾰‫ޔ‬
ⷞᐥߥߤߩߐ߹ߑ߹ߥ⣖㗔ၞ߳ߩ㜞޿ㅒⴕᕈㆮવሶዉ౉ല₸ࠍ␜ߒߚ‫ޔߚ߹ޕ‬ᵈ౉ㇱ૏ߦ߅޿ߡ␹⚻⚦⢩ߣࠣ࡝ࠕ
⚦⢩߳ߩㆮવሶዉ౉ߩല₸ߦߟ޿ߡ߽⸃ᨆߒߚ‫ޕ‬FuG-C ࡌࠢ࠲࡯ߪ‫ޔ‬ᵈ౉ㇱ૏ߩዋᢙߩ␹⚻⚦⢩ߦዉ౉ߐࠇࠆ߇‫ޔ‬
ࠣ࡝ࠕ⚦⢩ߦߪ߶ߣࠎߤዉ౉ߐࠇߥ߆ߞߚ‫ޔߡߞ߇ߚߒޕ‬FuG-C ࡌࠢ࠲࡯ߪ‫ࠅࠃޔ‬㜞޿ㅒⴕᕈㆮવሶዉ౉ല₸ࠍ␜
ߔߣߣ߽ߦ‫ޔ‬ᵈ౉ㇱ૏ߩࠣ࡝ࠕ⚦⢩߳ߩㆮવሶዉ౉ࠍᷫዋߐߖ‫ࠍ்៊❱⚵ޔ‬シᷫߔࠆߚ߼ߦ᦭⋉ߥߎߣ߇್᣿ߒ
ߚ‫․ߥ߁ࠃߩߎޕ‬ᕈࠍᜬߟࡌࠢ࠲࡯ߩ㐿⊒ߪ‫〝⚻ޔ‬ㆬᛯ⊛ߦ⚦⢩޽ࠆ޿ߪㆮવሶᯏ⢻ࠍᠲ૞ߒ‫ⴕޔ‬േ․ᕈࠍ೙ᓮߔ
ࠆㆮવሶᯏ⢻߿㜞ᰴ⣖ᯏ⢻ࠍ⸃ᨆߔࠆߚ߼ߩࡕ࠺࡞૞⵾ߦ߅޿ߡ᦭⋉ߥታ㛎♽ࠍឭଏߔࠆ‫ޕ‬
A-2 ⃻↢߅ࠃ߮ൻ⍹ࠝ࠽ࠟࠩ࡞㘃ߦ߅ߌࠆㅴൻᒻᘒቇ⊛⎇ⓥ
ዊ⮐ᄢテ㧔੩ᄢ࡮ඳ‛㙚㧕 ᚲౝኻᔕ⠪㧦㜞੗ᱜᚑ
ࠝ࠽ࠟࠩ࡞㘃ߩ㗡ㇱߦ߅ߌࠆᒻᘒቇ⊛ᄙ᭽ᕈ߇ߤߩࠃ߁ߥⷐ࿃ߩ߽ߣߢㅴൻߒߡ߈ߚߩ߆ࠍℂ⸃ߔࠆߎߣࠍ⋡
ᜰߒ‫ޔ‬ᯏ⢻ᒻᘒቇ‫⛔♽ޔ‬ቇ‫ޔ‬ㅴൻ↢ᘒቇߩⷰὐ߆ࠄ✚ว⊛ߥ⎇ⓥࠍⴕߞߚ‫ޕ‬CT ↹௝ࠍਃᰴరౣ᭴▽ߒ‫࠽ࠝ↢⃻ޔ‬
ࠟࠩ࡞㘃㗡㛽ߩਃᰴరᐞ૗ቇ⊛ߥቯ㊂ൻࠍㅴ߼ߚ‫ޔߚ߹ޕ‬㗡ㇱᒻᘒߣ㘩ᕈߩ㑐ଥࠍ⸃ᨆߔࠆߚ߼‫࡝ࠞ࡮࠳ࡦࠟ࠙ޔ‬
ࡦ࠭᫪ᨋߢࠝ࠽ࠟࠩ࡞㘃 4 ⒳㧔ࠕࠞࠝࠩ࡞‫ࠬࡉࡠࠦࠕ࠾ࠪࡆࠕޔ࡯ࠠࡦࡕ࠻ࠬࠛࡠޔ࡞ࠩࠝࠕޔ‬㧕ߩណ㘩‛ߩ⎕უ
㕢ᕈߩቯ㊂ൻࠍⴕߞߚ‫ޕ‬෼㓸ߐࠇߚ࠺࡯࠲ࠍ߽ߣߦ㘩ᕈߣ㗡ㇱߩਃᰴరᒻᘒߩ⛔⸘⊛ߥኻᔕ㑐ଥߩ᛽಴ࠍㅴ߼ߡ޿
ࠆ‫ޕ‬
㧨⺰ᢥ㧪
ዊ⮐ᄢテ㧚㘩ߴࠆߎߣ߆ࠄߺߚູ੃㘃ߩ㗡㛽ㅴൻԟ̆㔤㐳㘃㧚THE BONE 25 (2): 167-170 (2011) ᩏ⺒᦭ࠅ.
㧨⪺ᦠ㧪
1) ዊ⮐ᄢテ. (in press) ࠾ࡎࡦࠩ࡞ߩᒻᘒ㧚
‫ޟ‬㊁↢േ‛කቇࠞ࡜࡯ࠕ࠻࡜ࠬ‫ޠ‬㧘ᣣᧄ㊁↢േ‛කቇળ ✬㧘ᢥ⧷ၴ
಴ .
2) ዊ⮐ᄢテ. (in press) 㗡ߩᒻߩㅴൻߣ㘩ߴ‛ߩၷߐߩ㑐ଥࠍ㊁ᄖߢតࠆ. ‫ޟ‬ᣣᧄߩࠨ࡞ቇ̆⧯ᚻ⎇ⓥ⠪ߩᦨ೨
✢㧔઒㧕‫ޠ‬㧘ਛᎹዏผ㧘෹᳗㓷Ꮖ㧘ጊᭂᄈ৻ ✬㧘੩ㇺㅢା␠.
A-3 ⃻↢ᣥ਎⇇ࠩ࡞ߦ߅ߌࠆ〒㛽ߩᄌ⇣ߣൻ⍹߳ߩᔕ↪
㎄ᧄᱞਭ㧔ᨋේ࡮↢ൻ⎇㧕 ᚲౝኻᔕ⠪㧦㜞੗ᱜᚑ
ൻ⍹㔤㐳㘃ߩ⎇ⓥ߳ᔕ↪ߔࠆߚ߼‫↢⃻ޔ‬㔤㐳㘃ߦ߅ߌࠆ〒㛽ࠨࠗ࠭ߣᱤࠨࠗ࠭࡮૕㊀ߣߩ㑐ଥࠍ⺞ߴߡ޿ࠆ‫ᧄޕ‬
ᐕᐲߪએਅߩੑߟߩߎߣࠍ⺞ߴߚ‫ޕ‬
(1) 〒㛽ൻ⍹߆ࠄߘߩൻ⍹㔤㐳㘃ߩ૕㊀ࠍផቯߔࠆߚ߼ߦ‫↢⃻ޔ‬㔤㐳㘃ߩ〒㛽ࠨࠗ࠭ߩ⸘᷹ࠍ߅ߎߥߞߡ‫〒ޔ‬㛽ࠨ
ࠗ࠭ߣ૕㊀ߣߩ㑐ଥࠍᬌ⸛ߒߚ‫⚿ߩߘޕ‬ᨐ‫〒ޟޔ‬㛽ߩో㐳‫߇ߩ߁૶ࠍޠ‬૕㊀ផቯߦᦨ߽ㆡߒߡ޿ࠆߎߣ߇ࠊ߆
ߞߚ‫⚿ߩߎޕ‬ᨐࠍᔕ↪ߒߡ‫⑼ࠬࠢ࠹ࡇࠖࡈࡦࠕߩ↥࡯ࡑࡦࡖࡒޔ‬㔤㐳㘃ߩ〒㛽ൻ⍹ NMMP-39 ߅ࠃ߮ NMMP-82
ࠍ૕㊀ផቯߒߚ‫⚿ޕ‬ᨐߪ‫ޔ‬NMMP-39 ߪ⚂ 2.7kg‫ޔ‬NMMP-82 ߪ⚂ 5.1kg ߣߥߞߚ‫ޕ‬
(2) ࠾ࡎࡦࠩ࡞ߩᄢੱߩ୘૕ࠍኻ⽎ߦ〒㛽࡮ਅ㗶╙৻⥓ᱤ㧔m1㧕ࠨࠗ࠭ߣ૕㊀ߣߩ㑐ଥࠍ⺞ߴߚ‫੹ޕ‬࿁૶↪ߒߚ
ో୘૕ࠍኻ⽎ߣߒߚ႐ว‫〒ޔ‬㛽ࠨࠗ࠭ߣ૕㊀ߣߩ㑆ߦᱜߩ⋧㑐߇޽ߞߚ‫ޔߒ߆ߒޕ‬㓶㓽ࠍ඙೎ߒߡ೎‫⸃ߦޘ‬ᨆ
ߒߚ႐วߪ‫ޔ‬㓶㓽ߣ߽ߦ〒㛽ࠨࠗ࠭ߣ૕㊀ߣߩ㑆ߦ⋧㑐㑐ଥߪ⹺߼ࠄࠇߥ߆ߞߚ‫ޕ‬m1 ࠨࠗ࠭ߣ૕㊀ߣߩ㑐ଥ‫ޔ‬
߅ࠃ߮〒㛽ࠨࠗ࠭ߣ m1 ࠨࠗ࠭ߣߩ㑐ଥ߽ห᭽ߢ޽ߞߚ‫੹ޕ‬ᓟߪ‫ޔ‬୘૕ᢙ߅ࠃ߮⸃ᨆߔࠆᱤ⒳ࠍჇ߿ߒߡౣᐲ
⸃ᨆߔࠆߣߣ߽ߦ‫ߦࠄߐޔ‬ሶߤ߽ߩ୘૕ߦ߅ߌࠆ〒㛽ࠨࠗ࠭ߣ૕㊀ߣߩ㑐ଥࠍ⺞ߴࠆ੍ቯߢ޽ࠆ‫ߦࠄߐޕ‬዁᧪
⊛ߦߪ‫⸃ޔ‬ᨆኻ⽎ࠍ࠾ࡎࡦࠩ࡞એᄖߩᣥ਎⇇ࠩ࡞ߦ߽ᐢߍߡ޿ߊ੍ቯߢ޽ࠆ‫ޕ‬
㧨ቇળ⊒⴫㧪
㎄ᧄᱞਭ㧘߶߆㧔2011㧕⃻↢㔤㐳㘃ߩ〒㛽ࠨࠗ࠭ߩ⸘᷹ߣࡐࡦ࠳࠙ࡦߩൻ⍹㔤㐳㘃ߩ૕㊀ផቯ߳ߩᔕ↪㧚㔤㐳㘃
⎇ⓥ, vol. 27, Supplement, p. 34㧚ᣣᧄ㔤㐳㘃ቇળ㧔2011/07/16-18㧘›ጊ㧕
A-4 㔤㐳㘃ߦ߅ߌࠆ᭎ᣣᤨ⸘ߣ⣖㜞ᰴᯏ⢻ߣߩㅪ㑐
ᷡ᳓⾆⟤ሶ㧔᧲ᄢ࡮㒮࡮ℂ㧕㧘ᷓ↰ศቁ㧔᧲ᄢ࡮㒮࡮ℂ㧕 ᚲౝኻᔕ⠪㧦੹੗໪㓶
ᚒ‫ޔߢ߹ࠇߎߪޘ‬㦅ᱤ㘃ࠍ߽ߜ޿ߡ᭎ᣣᤨ⸘ߣ⸥ᙘᒻᚑᯏ᭴ߣߩ㑐ଥߦߟ޿ߡ‫ޔ‬ಽሶ↢‛ቇ⊛‫ޔ‬ㆮવቇ⊛‫ⴕޔ‬േ
㧙㧙
ቇ⊛ⷞὐ߆ࠄ⎇ⓥࠍⴕߞߡ߈ߚ‫ޔߢ߹ࠇߎޕ‬㦅ᱤ㘃ࠍ↪޿ߚታ㛎ߦ߅޿ߡ‫ޔ‬ᶏ㚍ଐሽᕈߩ㐳ᦼ⸥ᙘᒻᚑല₸ߦ᭎ᣣ
࡝ ࠭ ࡓ ߇ ޽ ࠆ ߎ ߣ ࠍ ⏕ ⹺ ߒ ‫ ⺧ ↳ ޔ‬⠪ ߇ ⷗ ޿ ߛ ߒ ߚ K-Ras ೙ ᓮ ࠲ ࡦ ࡄ ࠢ ⾰ SCOP (SCN circadian oscillatory
protein)/PHLPP ߣ޿߁ಽሶ߇᭎ᣣᤨ⸘ߣ⣖㜞ᰴᯏ⢻ࠍ⚿߮ߟߌࠆ㎛࿃ሶߢ޽ࠆน⢻ᕈࠍ␜ߔ⚿ᨐࠍᓧߡ߈ߚ
㧔Shimizu et al.Cell 2007㧕‫⎇ᧄޕ‬ⓥߢߪ‫ࠅࠃߦ࠻ࡅޔ‬ㄭ޿⣖᭴ㅧ࡮࿁〝ࠍᜬߟࠨ࡞ࠍ↪޿ߡ‫ޔ‬SCOP ࠍਛᔃߦ᭎ᣣᤨ
⸘ߣ⣖㜞ᰴᯏ⢻ߣߩ㑐ଥࠍ᣿ࠄ߆ߦߔࠆߎߣࠍ⋡⊛ߣߔࠆ‫ޕ‬㦅ᱤ㘃ߢߪਇน⢻ߢ޽ߞߚᖱേ㕖ଐሽᕈߩ⸥ᙘല₸ߩ
ᣣౝ࡝࠭ࡓ࡮ࠨ࡯ࠞ࠺ࠖࠕࡦ࡝࠭ࡓߩ⸃ᨆߦขࠅ⚵߻ߚ߼ߦ‫ޔ‬SCOP shRNA ⊒⃻࠙ࠗ࡞ࠬࠍ૶ߞߡㇱ૏․⇣⊛ߦ
SCOP ߩ⊒⃻ࠍᛥ೙ߒ‫⸥ߣ⸘ᤨޔ‬ᙘࠍ⚿߮ߟߌࠆ㎛࿃ሶߣߒߡߩᯏ⢻ࠍ⹏ଔߔࠆߎߣߦ⌕ᚻߒߚ‫ޕ‬ല₸ࠃߊࠨ࡞ߩ
⣖ౝߢ shRNA ࠍ⊒⃻ߐߖࠆߚ߼‫ ߩ࡞ࠩࠥࠞࠕޔ‬U6 pol III promoter ࠍන㔌ߒ‫ޔ‬SCOP shRNA ⊒⃻࠙ࠗ࡞ࠬࡌࠢ࠲࡯
ߦ⚵ߺㄟࠎߛ‫ޕ‬૞ᚑߒߚ shRNA ⊒⃻ࡊ࡜ࠬࡒ࠼ߪ COS7 ⚦⢩ਛߩ SCOP ㊂ࠍᷫዋߔࠆ੐ࠍ⏕⹺ߒߚ‫ޕ‬ᰴᐕᐲߪ‫ޔ‬
ߎߩ shRNA ⊒⃻࡟ࡦ࠴࠙ࠗ࡞ࠬࠍࠨ࡞ߩᶏ㚍ߦᛩਈߒ‫ޔ‬SCOP ߩࡁ࠶ࠢ࠳࠙ࡦࠍ⏕⹺ߔࠆߣߣ߽ߦ⸥ᙘല₸ߩ᷹
ቯࠍⴕ߁੍ቯߢ޽ࠆ‫ޕ‬
A-5 ࠥࡁࡓߦࠃࠆ㔤㐳㘃ߦ߅ߌࠆ⣖ᯏ⢻ߩᄙ᭽ᕈߩ⸃᣿
ᯅᧄ੫ᄥ㧔ᄢ㒋ᄢ࡮㒮࡮ㅪวዊఽ⊒㆐ቇ⎇ⓥ⑼㧕㧘቟↰↱⪇㧔ᄢ㒋ᄢ࡮㒮࡮කቇ♽⎇ⓥ⑼࡮♖␹කቇᢎቶ㧕㧘ጊ᫪⧷
㐳㧔ᄢ㒋ᄢ࡮㒮࡮කቇ♽⎇ⓥ⑼࡮ಽሶ♖␹␹⚻ቇ㧕 ᚲౝኻᔕ⠪㧦੹੗໪㓶
⛔วᄬ⺞∝‫ޔ‬෺ᭂᕈ㓚ኂ‫∔␹♖ߩߤߥ∛ߟ߁ޔ‬ᖚߦ㑐ㅪߔࠆߎߣ߇ߔߢߦ⍮ࠄࠇߡ޿ࠆ࡝ࠬࠢㆮવሶߢ޽ࠆ
COMT‫ޔ‬BDNF‫ޔ‬DISC1 ߦߟ޿ߡ‫ߢ࡞ࠨߣ࠻ࡅޔ‬ᄙဳ⸃ᨆࠍⴕߞߚ‫ޕ‬COMT (catechol-o-methyltransferase gene) ߪ‫ޔ‬
࠼࡯ࡄࡒࡦߩઍ⻢㉂⚛ߢ޽ࠅ‫ޔ‬COMT ߦߪᯏ⢻⊛ㆮવሶᄙဳ㧔Val158Met㧕߇޽ࠆߎߣ߇⍮ࠄࠇߡ޿ࠆ‫ޕ‬Val ᄙဳߪ
Met ᄙဳߣᲧセߒߡ࠼࡯ࡄࡒࡦࠍઍ⻢ߔࠆ㉂⚛ᵴᕈ߇㜞޿ߎߣ߆ࠄ‫ߩ࠻ࡅޔ‬೨㗡⪲ߦ߅޿ߡ Val ᄙဳߢߪ Met ᄙဳ
ࠃࠅ࠼࡯ࡄࡒࡦ߇ᄙߊઍ⻢ߐࠇ‫ࡦࡒࡄ࡯࠼ޔ‬㊂߇ૐਅߔࠆߎߣ߇ᗐቯߐࠇࠆ‫⛔ޔߢߎߘޕ‬วᄬ⺞∝ߦ߅޿ߡ㓚ኂߐ
ࠇߡ޿ࠆߎߣ߇⍮ࠄࠇߡ޿ࠆ೨㗡⪲⺖㗴ߢ޽ࠆ WCST ࠍⴕ޿‫ޔ‬Val ᄙဳࠍᜬߟߣ Met ᄙဳࠍᜬߟ߽ߩࠃࠅ WCST
ߩᚑ❣߇ૐ޿ߎߣࠍ⷗಴ߒߚ‫ޔߦࠄߐޕ‬೨㗡⪲ᯏ⢻ല₸ࠍ fMRI ߦߡ᷹ቯߒ‫ޔ‬Val ᄙဳࠍᜬߟ߽ߩߢߪߘߩല₸߇
ᖡ޿ߎߣࠍ␜ߒߚ‫ᦨޕ‬ᓟߦ‫ޔ‬ㆮવሶ㑐ㅪ⸃ᨆߦࠃࠅ‫ޔ‬Val ㆮવሶᄙဳߪ⛔วᄬ⺞∝ߩ࡝ࠬࠢߣߥࠆߎߣࠍႎ๔ߒߡ
޿ࠆ‫ ߜࠊߥߔޕ‬COMT ㆮવሶߩ Val ᄙဳߪ Met ᄙဳߣᲧセߒߡ COMT ㉂⚛ᵴᕈ߇㜞ߊ‫⚿ߩߘޔ‬ᨐ‫ޔ‬೨㗡⪲ߩ࠼࡯
ࡄࡒࡦ㊂߇ૐਅߒ‫ޔ‬೨㗡⪲ᯏ⢻ല₸߇ᖡߊߥࠅ‫⛔ޔ‬วᄬ⺞∝ߩ࡝ࠬࠢߣߥࠆߣ޿߁ߎߣߢ޽ࠆ‫࠻ࡅޕ‬એᄖߢߪ࠾ࡎ
ࡦࠩ࡞ߣࠕࠞࠥࠩ࡞ߢᄙဳ⸃ᨆࠍⴕߞߚ⚿ᨐ‫ߥ⊛⇣․ߦ࡞ࠩࡦࡎ࠾ޔ‬ㆮવሶᄙဳ߇⊒⷗ߐࠇߚ‫ߩߎޕ‬ᄌ⇣ߪࠕࡒࡁ
㉄ߩᕈ⾰ࠍᄌ߃ࠆ⟎឵ࠍ઻߁ߚ߼‫ߩ⾰ࠢࡄࡦ࠲ޔ‬ᯏ⢻ߦᓇ㗀ࠍਈ߃ࠆߎߣ߇␜ໂߐࠇࠆ‫ޕ‬
BDNF (Brain-Derived Neurotrophic Factor) ߪ‫⊒⚻␹ޔ‬㆐࡮␹⚻น႟ᕈ࡮␹⚻଻⼔ߦ㊀ⷐߥᓎഀࠍᨐߚߔઍ⴫⊛ߥ
␹⚻ᩕ㙃࿃ሶߢ޽ࠆ‫ߩߎޕ‬ಽሶߪ‫∔␹♖ޔ‬ᖚߩਛߢ߽․ߦ෺ᭂᕈ㓚ኂ߿߁ߟ∛ߣߩ㑐ࠊࠅ߇ᷓ޿ߎߣ߇ࠃߊ⍮ࠄࠇ
ߡ޿ࠆ‫ޕ‬BDNF ߪ♖␹∔ᖚߣߩ㑐ㅪߛߌߢߪߥߊ‫⸥ޔ‬ᙘࠍਛᔃߣߔࠆ⹺⍮ᯏ⢻ߣ㑐ㅪߔࠆߎߣ߇‫ߣ࠻ࡅޔ‬േ‛ߩ෺
ᣇߩ⎇ⓥ߆ࠄ᣿ࠄ߆ߦߐࠇߡ߈ߚ‫ ߦ․ޕ‬BDNF ߩ Val66Met ᄙဳߩ Met ဳࠃࠅ߽ Val ဳ߇⸥ᙘജ߇㜞ߊ‫ޔ‬Met ဳߩ
Ⱞ⊕ߪࠪ࠽ࡊࠬ߳ߩಽᴲ⇣Ᏹ߇⹺߼ࠄࠇࠆ‫ߡ޿ߟߦ࡞ࠨޕ‬ᄙဳ⸃ᨆࠍⴕߞߚ⚿ᨐ‫ޔ‬ㆮવሶᄙဳߪోߊ⹺߼ࠄࠇߥ߆
ߞߚ‫ޕ‬
DISC1(Disrupted in schizophrenia 1) ߪࠬࠦ࠶࠻࡜ࡦ࠼ߩ⛔วᄬ⺞∝߿߁ߟ∛ߩᄙ⊒ኅ♽ߦ߅޿ߡ⷗಴ߐࠇߚㆮવ
ሶߢ޽ࠆ‫⊒⚻␹ޕ‬㆐ߦ߅ߌࠆ⊹⾰ߩᒻᚑ߿␹⚻⚦⢩ߩㆆ⿛ߥߤߦ㑐ࠊߞߡ޿ࠆߎߣ߇⍮ࠄࠇߡ޿ࠆ‫ ߩߎޕ‬Ser704Cys
ᄙဳ߇ ERK ࠪࠣ࠽࡞ࠍ⺞▵ߔࠆᯏ⢻ᄙဳߢ޽ࠅ‫ޔ‬⣖᭴ㅧߩ⣀ᒙᕈࠍ੺ߒߡ߁ߟ∛ߩ࡝ࠬࠢߦߥࠆߎߣ߇␜ߐࠇߡ
޿ࠆ‫ ࡦ࠰ࠢࠛߡ޿߅ߦ࡞ࠩࡦࡎ࠾ߪߢ࡞ࠨޕ‬1‫ޔ‬3‫ޔ‬4‫ޔ‬5‫ޔ‬9 ߦ․⇣⊛ߥㆮવሶᄙဳ߇⷗޿ߛߐࠇߚ‫ޕ‬
A-6 ⴕേ․ᕈࠍᡰ㈩ߔࠆࠥࡁࡓၮ⋚ߣ⣖ᯏ⢻ߩ⸃᣿
ᤊ⧷ม㧔᧲੩ㇺකቇ✚ว⎇ⓥᚲ࡮೨㗡⪲ᯏ⢻ࡊࡠࠫࠚࠢ࠻㧕 ᚲౝኻᔕ⠪㧦㜞↰᣽ᒾ
㔤㐳㘃ߢᄢ߈ߊ⊒㆐ߒߚ೨㗡ㅪว㊁߇⋡⊛ᜰะ⊛ⴕേߦ߅޿ߡਛᔃ⊛ߥᓎഀࠍᨐߚߔߎߣ߇␜ߐࠇߡ߈ߡ޿ࠆ‫ޕ‬
⋡⊛⹜ⴕ⊛ⴕേߩઍ⴫଀ߢ޽ࠆ᧦ઙઃ߈ⷞⷡㆇേᄌ឵⺖㗴ߢߪ‫ޔ‬๒␜ߐࠇߚⷞⷡ‛૕ࠍ⍮ⷡߒߡ‫ߦࠇߘޔ‬ㅪวߐࠇ
ߚㆇേࠍㆬᛯ࡮ታⴕߔࠆᔅⷐ߇޽ࠆ‫⷗৻ޕ‬න⚐ߥⴕേߢ޽ࠆ߇‫ޔ‬⍮ⷡߣㆇേࠍᨵエߦㅪวߔࠆᔅⷐ߇޽ࠆߚ߼‫ⶄޔ‬
ᢙߩ⣖ࠪࠬ࠹ࡓߦ߹ߚ߇ࠆⶄ㔀ߥᖱႎಣℂ߇ⷐ᳞ߐࠇࠆ‫⺖ߩߎޕ‬㗴ߦ߅޿ߡߪ‫ޔ‬೨㗡ㅪว㊁ߩ߁ߜ೨㗡೨㊁⣻ᄖ஥
ㇱ㧔vlPFC㧕ߣㆇേ೨㊁⢛஥ㇱ㧔PMd㧕߇ᔅ㗇ߢ޽ࠆߎߣ߇␜ߐࠇߡ޿ࠆ߇‫ߩࠄࠇߎޔ‬㑆ߦߪ⋥ធᛩ኿߇ሽ࿷ߒߥ
޿ߎߣ߇ಽ߆ߞߡ޿ࠆ‫ޔߢߎߘޕ‬vlPFC ߣ PMd ߩ㑆ߦߪⶄᢙߩࠪ࠽ࡊࠬࠍ੺ߒߚᛩ኿߇޽ࠆߣ޿߁઒⺑ߩ߽ߣߦ
⸃೬ቇ⊛⎇ⓥࠍታᣉߒߚ‫ޕ‬ห৻୘૕ߦ߅޿ߡ‫ޔ‬PMd ߳ㅒⴕᕈ࠻࡟࡯ࠨ࡯ࠍᵈ౉ߒ‫ޔ‬vlPFC ߳㗅ⴕᕈ࠻࡟࡯ࠨ࡯ࠍᵈ
౉ߒߚߣߎࠈ‫ޔ‬೨㗡ㅪว㊁ߩ⢛஥ㇱߣ⣻஥ㇱߦ߅ߌࠆⶄᢙߩ㗔㊁ߦ߅޿ߡ‫ޔ‬PMd ߳ᛩ኿ߔࠆ⚦⢩ߩಽᏓߣ vlPFC
⚦⢩ߩ␹⚻⚳ᧃߩಽᏓߦ㊀ߥࠅ߇޽ࠆߎߣ߇᣿ࠄ߆ߣߥߞߚ‫ޕ‬ᓥߞߡ‫੺ߚߒ߁ߎޔ‬࿷㗔ၞࠍ฽߼ߚ೨㗡ㅪว㊁ౝߩ
ࡀ࠶࠻ࡢ࡯ࠢ߇‫⊛⋡ޔ‬ᜰะ⊛ⴕേߦ߅޿ߡ㊀ⷐߥᓎഀࠍᨐߚߔߎߣ߇␜ໂߐࠇߚ‫ޕ‬
A-7 ⃻↢߅ࠃ߮⠨ฎㆮ〔಴࿯࠾ࡎࡦࠩ࡞ߩ㛽ᒻᘒᄌ⇣ߦ㑐ߔࠆ⎇ⓥ
ᆌፒᥓሶ㧔⟲㚍⋵┙⥄ὼผඳ‛㙚㧕 ᚲౝኻᔕ⠪㧦㜞੗ᱜᚑ
࠾ࡎࡦࠩ࡞㧔Macaca fuscata㧕ᒻᘒ⊛․ᓽߦߟ޿ߡᄢಽ⋵‫ޔ‬ፉᩮ⋵‫⑔ޔ‬੗⋵‫ޔ‬ᣂẟ⋵‫ޔ‬㐳㊁⋵‫ޔ‬ጊ᪸⋵‫␹ޔ‬ᄹᎹ
⋵‫ޔ‬ජ⪲⋵‫⾗↢⃻ߩ⋵↰⑺ޔ‬ᢱߣ‫ޔ‬㣮ఽፉ⋵‫ޔ‬ጊญ⋵‫ޔ‬ᗲᇫ⋵‫␹ޔ‬ᄹᎹ⋵‫⑔ޔ‬੗⋵‫ޔ‬㐳㊁⋵‫ޔ‬ජ⪲⋵ߩ⠨ฎ⾗ᢱࠍ
Ყセߒ‫ᤨޔ‬ⓨ㑆⊛ᄌ⇣ࠍᬌ⸛ߒߚ‫ޕ‬Ყセߦ↪޿ߚ㗄⋡ߪਅ㗶ዊ⥓ᱤ࡮ᄢ⥓ᱤߩ㗠⥠ᓘߢ޽ࠆ‫⚿ߩߘޕ‬ᨐ‫⾗↢⃻ޔ‬ᢱ
ߣ⠨ฎ⾗ᢱߦߺࠄࠇࠆ࿾ℂ⊛ᄌ⇣ߦᄢ߈ߥ㆑޿ߪ‫⑔ޔ‬੗⋵ࠍ㒰޿ߡ⹺߼ࠄࠇߥ޿ߎߣ߇␜ߐࠇߚ‫ޔߚ߹ޕ‬㛽ᩰߣ⥓
ᱤࠨࠗ࠭ߩ࿾ℂ⊛ᄌ⇣ߦߪ⇣ߥࠆ௑ะ߇⹺߼ࠄࠇࠆߎߣ߇ᜰ៰ߐࠇߡ޿ࠆߎߣ߆ࠄ‫ޔ‬り૕⸘᷹࠺࡯࠲ߣ㛽ᩰ‫⥓ޔ‬ᱤ
㧙㧙
ࠨࠗ࠭ߩ⋧㑐ࠍᬌ⸛ߔࠆᔅⷐ߇޽ࠆ‫ޕ‬2011 ᐕᐲߪ‫ޔ‬2010 ᐕᐲߦᒁ߈⛯߈‫⟲ޔ‬㚍⋵ߢ᝝Ვߐࠇߚࠨ࡞ 20 ୘૕ࠍ೬ᬌ
ߒ‫࠲࡯࠺ޔ‬ᢛ஻࡮㛽ᮡᧄൻ࡮⸘᷹ࠍⴕߞߚ‫ޕ‬
৻ㇱߩᚑᨐߦߟ޿ߡߪ‫ޔ‬ᣣᧄ㔤㐳㘃ቇળ㧔ᣈ ›ጊᏒ 2011.7.16-7.18㧕ߦ߅޿ߡ‫ޕߚߒ⴫⊒ޔ‬㧔㗴⋡㧦⠨ฎㆮ〔߆
ࠄ಴࿯ߔࠆ࠾ࡎࡦࠩ࡞ൻ⍹ߦߟ޿ߡ㧕
A-8 㔤㐳㘃ᬁ㛽ߩᄖㇱᒻ⁁ߣౝㇱ᭴ㅧߩ⛔ว⸃ᨆ
᧲⪇ጪ㧔ጘ㒂ᄢ࡮ක㧕 ᚲౝኻᔕ⠪㧦⷏᧛೰
ᬁ㛽ߩᄖㇱᒻᘒߣౝㇱ᭴ㅧ߇ജቇ⊛ߦⶄ㔀ߦ⛊ߺว߁ߎߣߦࠃࠅᬁ㛽ߩ㛽ᒝᐲ߇଻ᜬߐࠇߡ޿ࠆ‫⎇ᧄޕ‬ⓥߪ‫ޔ‬ᬁ
㛽ߩ᭴ㅧ․ᕈࠍ⸃᣿ߔࠆߚ߼‫ޔ‬㔤㐳㘃ᬁ㛽ߩᄖㇱᒻᘒ㧔㜞ߐ‫ޔ‬᏷‫ޔ‬ᷓߐ‫ޔ‬ᢿ㕙Ⓧ㧕ߣౝㇱ᭴ㅧ㧔ౝㇱߦ޽ࠆᶏ✎㛽
ߩਃᰴరᓸ⚦᭴ㅧ㧕ߩቯ㊂⸃ᨆࠍⴕߥߞߚ‫ޕ‬3 ᱦ߆ࠄ 26 ᱦ߹ߢߩ࠾ࡎࡦࠩ࡞ 81 ୘૕㧔߅ߔ 38 㗡‫ ߔ߼ޔ‬43 㗡㧕ߩ
╙ 3 ⣶ᬁߩੇ῎㛽ᮡᧄࠍࡑࠗࠢࡠ CT ߢⷰኤߒ‫↹ޔ‬௝⸃ᨆ࠰ࡈ࠻࠙ࠚࠕࠍ↪޿ߡ‫ޔ‬⣶ᬁᬁ૕ߩ㜞ߐ‫ޔ‬᏷‫ޔ‬ᷓߐ‫ޔ‬ᢿ
㕙Ⓧࠍ᷹ቯߒ‫ޔ‬ᬁ૕ߩౝㇱߦ޽ࠆᶏ✎㛽ߩ㛽㊂㧔BV/TV㧕ߣ㛽ኒᐲߩ⸘᷹ࠍⴕߥߞߚ‫⚿ߩߘޕ‬ᨐ‫࡞ࠩࡦࡎ࠾ޔ‬⣶ᬁ
ᬁ૕ߩ㜞ߐߪട㦂ߦ઻޿ૐߊߥࠅ‫ޔ‬ᬁ૕ߩ᏷ߪട㦂ߣߣ߽ߦჇടߒߚ‫ޕ‬ᬁ૕ߩ᏷㧛㜞ߐߩഀวߪട㦂ߣߣ߽ߦჇട
ߒߚ‫ޕ‬ട㦂ߦࠃࠆᬁ૕ᷓߐߩ᦭ᗧߥᄌൻߪ⹺߼ࠄࠇߥ߆ߞߚ‫ޕ‬ᬁ૕ߩᢿ㕙Ⓧߪ‫ޔ‬ᬁ૕ߩ᏷ߩห᭽ߦട㦂ߣߣ߽ߦჇ
ടߒߚ‫ޕ‬ᬁ૕ౝㇱߦ޽ࠆᶏ✎㛽ߩ㛽㊂ߣ㛽ኒᐲߪട㦂ߦߒߚ߇ߞߡૐਅߒߚ‫ޕ‬ᬁ૕ߩ᏷㧛㜞ߐߩഀวߣᬁ૕ౝㇱߦ
޽ࠆᶏ✎㛽ߩ㛽㊂‫ߪߚ߹ޔ‬㛽ኒᐲߣߩ㑆ߦ᦭ᗧߥ⽶ߩ⋧㑐㑐ଥࠍ␜ߒߚ‫⚿ߩࠄࠇߎޕ‬ᨐߪ‫ޔ‬ᬁ૕ߩ᏷㧛㜞ߐߩഀว
ߪᬁ૕ౝㇱߩ㛽㊂ࠍ⴫ߔᜰᮡߩᅷᒰᕈࠍ⷗಴ߒߚ‫⃻ޕ‬࿷‫੃ູߩઁޔ‬േ‛ߣᲧセᬌ⸛ߒ‫ࠆߌ߅ߦ࠻ࡅޔ‬ട㦂ᕈ㛽☻㝃
∝ߩ⊒∝ᯏᐨߩ⸃᣿ࠍ⋡ᜰߔ‫ޕ‬
A-9 ⴕേ․ᕈࠍᡰ㈩ߔࠆࠥࡁࡓၮ⋚ߣ⣖ᯏ⢻ߩ⸃᣿
ධㇱ◊㧔⥄ὼ⑼ቇ⎇ⓥᯏ᭴࡮↢ℂ⎇࡮↢૕ࠪࠬ࠹ࡓ㧕㧘⇌ਛિᒾ㧘⍮⷗⡡⟤㧘➚➓ᄢテ㧘㜞⦟ᴕᐘ㧔⥄ὼ⑼ቇ⎇ⓥ
ᯏ᭴࡮↢ℂ⎇㧕 ᚲౝኻᔕ⠪㧦㜞↰᣽ᒾ
ᄢ⣖ၮᐩᩭߣᄢ⣖⊹⾰ߪ࡞࡯ࡊ࿁〝ࠍᒻᚑߒ‫ޔ‬ㆇേࠍߪߓ߼ߣߒߡ‫ޔ‬ᕁ⠨ߥߤߩ㜞ᰴᯏ⢻߿ᖱേߥߤ߽ࠦࡦ࠻ࡠ
࡯࡞ߒߡ޿ࠆߣ⠨߃ࠄࠇߡ޿ࠆ‫ࠍࡓ࠭࠾ࠞࡔߩࠄࠇߎޕ‬᣿ࠄ߆ߦߔࠆߚ߼ߦߪ‫ޔ‬ᄢ⣖⊹⾰ߩฦ㗔㊁߆ࠄ಴⊒ߒߚᖱ
ႎ߇‫ޔ‬ᄢ⣖ၮᐩᩭߩߤߩ㗔ၞߢಣℂߐࠇߡ޿ࠆߩ߆ࠍਤᔨߦㅊ߁ᔅⷐ߇޽ࠆ‫੹ޕ‬࿁ߪ‫࡞ࠨߦߊߣޔ‬ᄢ⣖⊹⾰ߩㆇേ
㑐ㅪ㗔㊁㧔৻ᰴㆇേ㊁‫⿷⵬ޔ‬ㆇേ㊁‫ޔ‬ㆇേ೨㊁㧕߆ࠄᄢ⣖ၮᐩᩭߩ౉ജㇱ㧔✢᧦૕‫ⷞޔ‬ᐥਅᩭ㧕߳ߩᛩ኿᭽ᑼࠍ‫ޔ‬
⸃೬ቇ⊛ߥᣇᴺߦࠃࠅ⺞ߴߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬ᄢ⣖⊹⾰৻ᰴㆇേ㊁ߪ‫᧦✢ޔ‬૕ߩᄖ஥ㇱߦᛩ኿ߔࠆߩߦኻߒ‫⿷⵬ޔ‬ㆇേ
㊁߆ࠄߩᛩ኿ߪ‫ޔ‬ౝ஥ㇱߦ⚳ᱛߔࠆߎߣ߇ࠊ߆ߞߚ‫ޕ‬ᰴߦ‫᧦✢ߥ߁ࠃߩߎޔ‬૕ߩ㗔ၞ߆ࠄ࠾ࡘ࡯ࡠࡦᵴേࠍ⸥㍳ߔ
ࠆߣ‫ޔ‬ㆇേߦ㓙ߒ‫⊒ߚߞߥ⇣ࠇߙࠇߘޔ‬኿ࡄ࠲࡯ࡦࠍ␜ߔߎߣ߇ࠊ߆ߞߚ‫ߜࠊߥߔޕ‬㓐ᗧㆇേߪ‫ߞߥ⇣ࠇߙࠇߘޔ‬
ߚᖱႎࠍᜂ߁ⶄᢙߩᄢ⣖⊹⾰—ᄢ⣖ၮᐩᩭ࡞࡯ࡊߦࠃߞߡࠦࡦ࠻ࡠ࡯࡞ߐࠇߡ޿ࠆߎߣ߇␜ໂߐࠇߚ‫੹ޕ‬ᓟߪ‫ߎޔ‬
ࠇએᄖߩ㜞ᰴ㗔㊁߆ࠄᄢ⣖ၮᐩᩭ߳ߩᛩ኿᭽ᑼߦߟ޿ߡ߽⺞ߴߡ޿߈ߚ޿‫ޕ‬
㧨⺰ᢥ⊒⴫㧪
Takara S, et al. (2011) Differential activity patterns of putaminal neurons with inputs from the primary motor cortex and
supplementary motor area in behaving monkeys. J Neurophysiol 106: 1203-1217
㧨ቇળ⊒⴫㧪
⇌ਛિᒾ㧘߶߆㧔2011/09/15㧕ㆇേ⺖㗴ㆀⴕਛߩࠨ࡞ߦ߅ߌࠆ᷆⫷⃿࠾ࡘ࡯ࡠࡦᵴേߩࠣ࡞࠲ࡒࡦ㉄߅ࠃ߮ GABA
૞േᕈ⺞▵㧚╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ㧔ᮮᵿ㧕
A-10 ᣥ਎⇇ࠩ࡞ߩᄌ⇣ᕈߣㅴൻߦ㑐ߔࠆᄙ㕙⊛ࠕࡊࡠ࡯࠴㧦࠾ࡎࡦࠩ࡞ߩᵢ─೑↪ߣൻ⍹ൻㆊ⒟
ᨰᧁஜม㧔ንጊᄢ࡮㒮࡮⎇ⓥㇱ࡮ℂቇ㧕
㧘ἑਯญ␽ቁ㧔ንጊᄢ࡮㒮࡮ᢎ⢒ㇱ࡮ℂቇ㧕
㧘㒙ㇱാᴦ㧔ᄙ⾐↸ඳ㧕 ᚲౝ
ኻᔕ⠪㧦㜞੗ᱜᚑ
⸘↹⎇ⓥ‫ޟ‬ᣥ਎⇇ࠩ࡞ߩᄌ⇣ᕈߣㅴൻߦ㑐ߔࠆᄙ㕙⊛ࠕࡊࡠ࡯࠴‫৻ߩޠ‬Ⅳߣߒߡ‫ޔ‬ንጊ⋵᧲ㇱ㤥ㇱጽ⼱ߩࠨ࡞ⓣ
㧔㎠੃ᵢ㧕߆ࠄ↥ߔࠆ࠾ࡎࡦࠩ࡞ൻ⍹ࠍኻ⽎ߦ‫ߩߘޔ‬ൻ⍹ൻㆊ⒟㧔࠲ࡈࠜࡁࡒ࡯㧕ߩᬌ⸛ࠍ‫ޔ‬2009 ᐕ 11 ᦬߆ࠄㅴ
߼ߡ޿ࠆ‫࡞ࠩࡦࡎ࠾ޔߡߒߘޕ‬ൻ⍹ߩ࠲ࡈࠜࡁࡒ࡯ࠍౕ૕⊛ߦ⼏⺰ߔࠆ਄ߢ‫ޔ‬3 Ბ㓏ߩ୘೎⎇ⓥࠍ〯ߺ‫ࠍࠄࠇߘޔ‬
✚ว⼏⺰ߔࠆᔅⷐ߇޽ࠆ‫ޕ‬හߜ‫ޔ‬1) ࠨ࡞ⓣߩᵢౝᒻ⁁⸥タ‫ޔ‬2) ⃻↢࠾ࡎࡦࠩ࡞ߦࠃࠆ♮ߩ↥⁁⸥タ‫ ߡߒߘޔ‬3) ࠾
ࡎࡦࠩ࡞ൻ⍹ߩ↥⁁߅ࠃ߮ฎ↢‛ቇ⊛⸥タ‫ޕࠆ޽ߢޔ‬2012 ᐕᐲߩ౒ห⎇ⓥߢߪ‫ᦨޔ‬ೋߩੑᲑ㓏߹ߢߩ࠺࡯࠲ࠍ߹
ߣ߼‫⺰ޔ‬ᢥߣߒߡᛩⓂ㧔ฃℂ࡮ᩏ⺒ਛ㧕ߔࠆߦ⥋ߞߡ޿ࠆ‫ޕ‬
⎇ⓥ᭎ⷐߪ‫ޔ‬એਅߩࠃ߁ߦ߹ߣ߼ࠄࠇࠆ‫࡞ࠨޕ‬ⓣߪ‫ޔ‬᷹✢✚ᑧ㐳ߢ 100 m ࠍ⿥߃ࠆ┱ᮮⶄวဳᵢ─ߢ‫ޔ‬ᵢญ߆ࠄ
20 m ߩᮮⓣࠍ੺ߒߡ┱ⓣߦㅪ⚿ߔࠆ‫࡞ࠩࡦࡎ࠾ޕ‬ൻ⍹ߪ‫ޔ‬ᮮⓣਛߩᡰᵢᅏߣ┱ⓣߩ⋥೨‫┱ߡߒߘޔ‬ⓣਛߩ 4 ▎ᚲ
ߢߺࠄࠇ‫ ోࠄࠇߎޔ‬6 ୘૕ߪోߡቢోߥᥧ㤥ߩⓨ㑆ߢ↥ߒߚ‫ޕ‬ൻ⍹ߩ↥⁁ߦၮߠߊߣ‫ޔ‬ේ࿾ᕈߥ޿ߒḰේ࿾ᕈߩൻ
⍹ߢ޽ࠆߎߣߪ㑆㆑޿ߥߊ‫ߪ࡞ࠩࡦࡎ࠾ޔ‬ᮮⓣߦ⥄ࠄ౉ࠅㄟߺ‫ޔ‬૗ࠄ߆ߩℂ↱ߢ┱ⓣߦ⪭ߜߡൻ⍹ൻߒߚ߽ߩߣߩ
ផ⺰߇ዉ߈಴ߐࠇࠆ‫ߩߘޕ‬ᓟ‫ޔ‬ᮮⓣਛߦ⃻↢࠾ࡎࡦࠩ࡞ߦࠃࠆᄙ㊂ߩ♮ࠍ⷗಴ߒ‫ޔ‬2010 ᐕᐲ౻ቄߦ㒐ኙ⋡⊛ߣߒ
ߡᵢ─ࠍ೑↪ߒߚ㓙ߦឃᴭߐࠇߚ♮ߢ޽ࠆߣߩ⼏⺰ࠍዷ㐿ߒߚ‫ޔߚ߹ޕ‬Ⓧ㔐㊂ߩჇᷫ߇࠾ࡎࡦࠩ࡞ߩᵢ─೑↪ࠍⷙ
೙ߒߡ޿ࠆὐࠍ‫ߎߎޔ‬ᢙᐕߩⓍ㔐㊂ᄌൻߦၮߠ޿ߡ⼏⺰ߒߚ‫ޕ‬ᣢߦ᷹ቯᷣߺߢ޽ࠆ࠾ࡎࡦࠩ࡞ൻ⍹ߩ὇⚛ 14 ᐕઍ
୯ࠍ⠨ᘦߔࠆߣ‫ޔ‬ᒎ↢ᤨઍ೨ᦼߦߪ࠾ࡎࡦࠩ࡞ߦࠃࠆᵢ─೑↪ߪ↢ᘒߣߒߡ⏕┙ߒߡ޿ߚߣ್ᢿߐࠇࠆ‫ޔߡߒߘޕ‬
┱ⓣਛߩ࠾ࡎࡦࠩ࡞ൻ⍹ߪ‫ޔ‬౻ቄߦᵢ─ߦ౉ࠅㄟࠎߛ࠾ࡎࡦࠩ࡞߇‫ޔ‬૗ࠄ߆ߩℂ↱ߢ⪭ߜㄟࠎߢൻ⍹ൻߒߚ߽ߩߣ
ߩ⼏⺰߇น⢻ߢ޽ࠆ‫ޕ‬
㧙㧙
਄⸥ߣਗⴕߒߡ‫࡞ࠩࡦࡎ࠾ޔ‬ൻ⍹ߩฎ↢‛ቇ⊛⸥タ㧔⸘᷹࡮CT ᠟ᓇ㧕ࠍⴕߞߚ‫ޕ‬ਥߣߒߡ‫ޔ‬㔤㐳⎇ᚲ⬿ߩ⃻↢
ᮡᧄߩዊ⥓ᱤߣᄢ⥓ᱤߦ⸘᷹ߦട߃‫⊕ޔ‬ጊ⥄ὼ଻⼔࠮ࡦ࠲࡯ᚲ⬿ߩ⊕ጊ↥⃻↢ᮡᧄ߽⸘᷹ኻ⽎ߣߒߚ‫ޔߚ߹ޕ‬CT
᠟ᓇߦࠃࠆ㗡⬄㛽ౝㇱߩᬌ⸛߽ታᣉߒߚ‫੍ޕ‬ኤ⊛ߦߪ‫⸘ޔ‬᷹୯ߦၮߠߊ㤥ㇱጽ⼱↥࠾ࡎࡦࠩ࡞ൻ⍹ߩ࿾ၞᕈࠍ⹺߼
ࠆߦ⥋ߞߡ޿ࠆ‫┃⌒ޔߚ߹ޕ‬਄ಾ∥ߩ᣿⍎ߥ⊒㆐ߥߤ‫ޔ‬㗡⬄㛽ߦ߅ߌࠆ․ᓽ⊛ߥᒻ⾰߽ߺࠄࠇࠆ‫ࠄࠇߎޕ‬ฎ↢‛ቇ
⊛⸥タߦߟ޿ߡߪ‫⸘ޔ‬᷹୯╬ߩౣᬌ⸽ࠍ⚻ߡ‫⺰ޔ‬ᢥਛߦᢥ┨ߣߒߡ᣿⸥ߔߴ߈ߢ޽ࠆߣ⠨߃‫ߥ⚦⹦ߪߢߎߎޔ‬ౝኈ
ࠍㆱߌࠆ‫ޕ‬
㧨⺰ᢥ㧪㧔ోߡ㧘⻢ㄉߦ੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲߩ౒ห⎇ⓥഥᚑࠍ૶↪ߒߚᣦࠍ᣿⸥㧕
1) ᨰᧁஜม㧘߶߆ ශ೚ਛ㧘ንጊ⋵㤥ㇱጽ⼱ߩ㏹㊒࿾ၞߩࠨ࡞ⓣ㧔㎠੃ᵢ㧕㧚࿾⾰ቇ㔀⹹㧚
㧔ᣣᧄ࿾⾰ቇળ㧘ᩏ
⺒⹹㧕
2) ᨰᧁஜม ශ೚ਛ㧘ንጊ⋵㤥ㇱጽ⼱ߩ㏹㊒࿾ၞߩ⍹Ἧጤᵢ─⎇ⓥผ㧚㤥ㇱ㧔㤥ㇱቇળ㧘㕖ᩏ⺒⹹㧕
3) ᨰᧁஜม㧘߶߆ 2012㧘࠾ࡎࡦࠩ࡞ߩᵢ─೑↪ߣൻ⍹ൻㆊ⒟㧔੍ኤ㧕㧚ฬฎደᄢቇടㅦེ⾰㊂ಽᨆ⸘ᬺ❣ႎ
๔ᦠ㧘XXIII㧘156-159㧚㧔ฬฎደᄢቇᐕઍ᷹ቯ✚ว⎇ⓥ࠮ࡦ࠲࡯㧘㕖ᩏ⺒⹹㧕
4) ᨰᧁஜม㧘߶߆ ᩏ⺒ਛ㧘⽕㔐࿾ၞߩ࠾ࡎࡦࠩ࡞ߦࠃࠆᵢ─೑↪㧚㔤㐳㘃⎇ⓥ㧚
㧔ᣣᧄ㔤㐳㘃ቇળ㧘ᩏ⺒⹹㧕
㧨ቇળ⊒⴫㧪㧔Ṷᣦήߒ㧕
1) ᨰᧁஜม㧘߶߆ 2011㧘ንጊ⋵᧲ㇱߩ㤥ㇱጽ⼱㏹㊒࿾ၞߩ㎠੃ᵢ↥࠾ࡎࡦࠩ࡞ൻ⍹㧚╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇ
ળߩ⥄↱㓸ળ‫ߩ࡞ࠩࡦࡎ࠾ޟ‬ൻ⍹‫ޠ‬㧚
2) ᨰᧁஜม㧘߶߆ 2012㧘㤥ㇱጽ⼱ࠨ࡞ⓣ↥࠾ࡎࡦࠩ࡞ൻ⍹ߩ὇⚛ 14 ᐕઍߣൻ⍹ൻㆊ⒟㧚╙ 24 ࿁㧔2011 ᐕᐲ㧕
ฬฎደᄢቇᐕઍ᷹ቯ✚ว⎇ⓥ࠮ࡦ࠲࡯ࠪࡦࡐࠫ࠙ࡓ㧚
એ਄ߩ⊒⴫߅ࠃ߮ႎ๔᭎ⷐߪ‫ޔ‬એਅߩࠃ߁ߦ߹ߣ߼ࠄࠇࠆ‫࡞ࠨޕ‬ⓣߪ‫ޔ‬᷹✢✚ᑧ㐳ߢ 100 m ࠍ⿥߃ࠆ┱ᮮⶄวဳ
ᵢ─ߢ‫ޔ‬ᵢญ߆ࠄ 20 m ߩᮮⓣࠍ੺ߒߡ┱ⓣߦㅪ⚿ߔࠆ‫࡞ࠩࡦࡎ࠾ޕ‬ൻ⍹ߪ‫ޔ‬ᮮⓣਛߩᡰᵢᅏߣ┱ⓣߩ⋥೨‫ߒߘޔ‬
ߡ┱ⓣਛߩ 4 ▎ᚲߢߺࠄࠇ‫ ోࠄࠇߎޔ‬6 ୘૕ߪోߡቢోߥᥧ㤥ߩⓨ㑆ߢ↥ߒߚ‫ޕ‬ൻ⍹ߩ↥⁁ߦၮߠߊߣ‫ޔ‬ේ࿾ᕈߥ
޿ߒḰේ࿾ᕈߩൻ⍹ߢ޽ࠆߎߣߪ㑆㆑޿ߥߊ‫ߪ࡞ࠩࡦࡎ࠾ޔ‬ᮮⓣߦ⥄ࠄ౉ࠅㄟߺ‫ޔ‬૗ࠄ߆ߩℂ↱ߢ┱ⓣߦ⪭ߜߡൻ
⍹ൻߒߚ߽ߩߣߩផ⺰߇ዉ߈಴ߐࠇࠆ‫ߩߘޕ‬ᓟ‫ޔ‬ᮮⓣਛߦ⃻↢࠾ࡎࡦࠩ࡞ߦࠃࠆᄙ㊂ߩ♮ࠍ⷗಴ߒ‫ޔ‬2010 ᐕᐲ౻
ቄߦ㒐ኙ⋡⊛ߣߒߡᵢ─ࠍ೑↪ߒߚ㓙ߦឃᴭߐࠇߚ♮ߢ޽ࠆߣߩ⼏⺰ࠍዷ㐿ߒߚ‫ޔߚ߹ޕ‬Ⓧ㔐㊂ߩჇᷫ߇࠾ࡎࡦࠩ
࡞ߩᵢ─೑↪ࠍⷙ೙ߒߡ޿ࠆὐࠍ‫ߎߎޔ‬ᢙᐕߩⓍ㔐㊂ᄌൻߦၮߠ޿ߡ⼏⺰ߒߚ‫ޕ‬ᣢߦ᷹ቯᷣߺߢ޽ࠆ὇⚛ 14 ᐕઍ
୯ࠍ⠨ᘦߔࠆߣ‫ޔ‬ᒎ↢ᤨઍ೨ᦼߦߪ࠾ࡎࡦࠩ࡞ߦࠃࠆᵢ─೑↪ߪ↢ᘒߣߒߡ⏕┙ߒߡ޿ߚߣ್ᢿߐࠇࠆ‫ޔߡߒߘޕ‬
┱ⓣਛߩ࠾ࡎࡦࠩ࡞ൻ⍹ߪ‫ޔ‬౻ቄߦᵢ─ߦ౉ࠅㄟࠎߛ࠾ࡎࡦࠩ࡞߇‫ޔ‬૗ࠄ߆ߩℂ↱ߢ⪭ߜㄟࠎߢൻ⍹ൻߒߚ߽ߩߣ
ߩ⼏⺰߇น⢻ߢ޽ࠆ‫ޕ‬
(2)
৻⥸୘ੱ⎇ⓥ
B-1 Diet and the Host-parasite ecology of chacma baboons
PA Pebsworth (Wildcliff Nature Reserve) ᚲౝኻᔕ⠪㧦MA Huffman
Geophagy is widespread in animals and occurs in 21% of all nonhuman primates, but has not been described by age class,
sex, or reproductive state. Because soil has the ability to alleviate gastro-intestinal (GI) distress and upsets, geophagy is
considered a self-medicative behavior. I analyzed data collected from my field study, which continually monitored soil
consumption in a troop of Papio hamadras ursinus at four geophagy sites with video camera traps from August 2009 through
January 2011. Using 60 hours of video recordings, I evaluated soil consumption by age class, sex, and reproductive state.
Pregnant baboons spent more time consuming soil at monitored geophagy sites than other baboons. This pattern of soil
consumption is similar to what is observed in humans. In addition to analyzing geophagic soils for physical, chemical, and
mineral properties, I analyzed these soils for presence of soil-transmitted helminths (STH) to evaluate the risk of parasite
transmission through soil consumption. I analyzed 272 fecal samples to determine parasites infecting this troop. Six
nematodes: Trichuris sp., one unidentified species from the suborder Spirurina, Strongyloides fuelleborni, Oesophagostomum
sp., Trichostrongylus sp., and Streptopharagus sp. were found. 80 soil samples were then analyzed for parasite presence, 40
from geophagy sites and 40 from foraging sites. My preliminary findings indicate that more Trichuris sp. ova were recovered
from samples collected where soils were consumed, and both geophagy and foraging sites was a potential source of STH
infection for this troop. However, black wattle stands pose a greater risk of STH infection than geophagy sites.
B-2 ㄘ࿾߳ߩଐሽᕈߩ⇣ߥࠆ࠾ࡎࡦࠩ࡞ 2 ⟲ߩ⟲⪭೑↪ߩᲧセ
ᶏ⠧ේኡ㧔㤗Ꮣᄢ࡮㒮࡮₞ක㧕 ᚲౝኻᔕ⠪㧦ㄞᄢ๺
ㄭᐕ‫࡞ࠨޔ‬㧔࠾ࡎࡦࠩ࡞㧕ߦࠃࠆㄘᬺⵍኂ߇↢ߓߡ޿ࠆ‫ޕ‬ㄘ᧛ߩㆊ⇹ൻߪ੹ᓟߐࠄߦㅴ߻ߣᕁࠊࠇ‫ੱޔ‬㉿ࠍ೑↪
ߔࠆࠨ࡞߽ߐࠄߦჇ߃ߡ޿ߊߣ੍ᗐߐࠇࠆ‫⁁ߚߒ߁ߎޕ‬ᴫߪ‫߇࡞ࠨޔ‬ⅣႺߩᄌൻߦኻߒߡ‫↢ߦ߁ࠃߩߤޔ‬ᵴࠍᄌൻ
ߐߖߚ߆ࠍ⍮ࠆᅢᯏߣ޿߃ࠆ‫⎇ᧄޕ‬ⓥߢߪ‫␹ޔ‬ᄹᎹ⋵ਤᴛ᧲ㇱߩㄘ࿾ࠍ೑↪ߒߥ޿⟲ࠇ㧔⥄ὼ⟲㧕ߣㄘ࿾ࠍ೑↪ߔ
ࠆ⟲ࠇ㧔ടኂ⟲㧕ߩ⟲⪭೑↪ࠍᲧセߒߚ‫࡝࠻ࡔ࡟࠹ࠝࠫ࡜ޕ‬ᴺߦࠃࠅ⟲ࠇߩ૏⟎ࠍᛠីߒ‫ޔ‬GIS ਄ߢⅣႺ⋭ߩᬀ↢
࿑ࠍ↪޿ߡ⸃ᨆࠍⴕߞߚ‫⥄ޕ‬ὼ⟲ߢߪ‫ޔ‬ᐢ⪲᮸ᨋߩ೑↪߇⑺ࠍ਄㒢ߣߒߡጊဳࠍ␜ߒ‫ޔ‬㊎⪲᮸ᨋߣ⨲࿾ߩ೑↪ߪ⑺
ࠍਅ㒢ߦ⼱ဳࠍ␜ߒߚ‫৻ޕ‬ᣇ‫ޔ‬ടኂ⟲ߢߪ‫ޔ‬ㄘ࿾ߩ೑↪߇⑺ࠍ਄㒢ߣߒߡጊဳࠍ␜ߒ‫⪭⟲ߩઁߩߘޔ‬೑↪ߦ௑ะߪ
⷗ࠄࠇߥ߆ߞߚ‫ޔߪߣߎߩߎޕ‬㘩‛ଏ⛎㊂߇㑐ଥߒߡ޿ࠆߎߣ߇⠨߃ࠄࠇࠆ‫ޕ‬2 ⟲ߩ⟲⪭೑↪ࠍᲧセߔࠆߣ‫ߩߤޔ‬
㧙㧙
ቄ▵ߦ߅޿ߡ߽ടኂ⟲ߩᣇ߇ㄘ࿾ߩ೑↪߇ᄙ߆ߞߚ‫ޔߪࠇߎޕ‬ㄘ࿾ߦᩕ㙃ଔߩ㜞޿૞‛߇㓸ਛ⊛ߦ޽ࠆߚ߼ߣ⠨߃
ࠄࠇࠆ‫⑺ޔߚ߹ޕ‬એᄖߪടኂ⟲ߩᣇ߇ᐢ⪲᮸ᨋߩ೑↪߇ᄙ߆ߞߚ‫ߩࠄ߆ੱޕ‬࿶ജ߿ᐢ⪲᮸ᨋ߇ㄘ࿾ߦ㓞ធߒߡ޿ࠆ
ߎߣ߇ᓇ㗀ߒߡ‫ޔ‬ᐢ⪲᮸ᨋߩ೑↪ࠍ㜞߼ߚߚ߼ߣ⠨߃ࠄࠇࠆ‫ޕ‬ㄘ࿾ߩሽ࿷ߪ‫⪭⟲ߩ࡞ࠨޔ‬೑↪ߦᄢ߈ߥᓇ㗀ࠍ෸߷
ߒߡ޿ࠆߎߣ߇␜ߐࠇߚ‫ޕ‬
㧨⊒⴫᭎ⷐ㧪
࠾ࡎࡦࠩ࡞ 2 ⟲ߩ⟲⪭೑↪ࡄ࠲࡯ࡦ 㨪㓞ធߔࠆ⥄ὼ⟲ߣടኂ⟲ߩᲧセ㨪
ᶏ⠧ේኡ㧘㜞ᮎᚑ♿㧔㤗Ꮣᄢ࡮㒮࡮₞ක㧕 ᚲౝኻᔕ⠪㧦ㄞᄢ๺
ㄭᐕ‫࡞ࠨޔ‬㧔࠾ࡎࡦࠩ࡞㧕ߦࠃࠆㄘᬺⵍኂ߇↢ߓߡ޿ࠆ‫⁁ߚߒ߁ߎޕ‬ᴫߪ৻⒳ߩታ㛎ߣߺࠆߎߣ߇ߢ߈‫߇࡞ࠨޔ‬
ⅣႺߩᄌൻߦኻߒߡ‫↢ߦ߁ࠃߩߤޔ‬ᵴࠍᄌൻߐߖߚ߆ࠍ⍮ࠆᅢᯏߣ޿߃ࠆ‫⎇ᧄޕ‬ⓥߢߪ‫␹ޔ‬ᄹᎹ⋵ਤᴛ᧲ㇱߩㄘ࿾
ࠍ೑↪ߒߥ޿⟲ࠇ㧔⥄ὼ⟲㧕ߣㄘ࿾ࠍ೑↪ߔࠆ⟲ࠇ㧔ടኂ⟲㧕ߩ⟲⪭೑↪ࠍᲧセߒߚ‫࡝࠻ࡔ࡟࠹ࠝࠫ࡜ޕ‬ᴺߦࠃࠅ
⟲ࠇߩ૏⟎ࠍᛠីߒ‫ޔ‬GIS ਄ߢⅣႺ⋭ߩᬀ↢࿑ࠍ↪޿ߡ⸃ᨆࠍⴕߞߚ‫⺞ޕ‬ᩏߪ 2011 ᐕ 6 ᦬㨪11 ᦬ߦ߅ߎߥߞߚ‫ޕ‬
⥄ὼ⟲ߪ‫ޔ‬ೋᄐߦ㊎⪲᮸ᨋߩ೑↪߇ᄙߊᐢ⪲᮸ᨋߩ೑↪߇ዋߥ߆ߞߚ߇‫ޔ‬ቄ▵߇ㅴ߻ߦߟࠇߡߎࠇ߇ㅒォߒߚ‫ޕ‬ട
ኂ⟲ߦ߅޿ߡߪ‫ޔ‬ೋᄐߦ⨲࿾ߩ೑↪߇ᄙߊㄘ࿾ߩ೑↪߇ዋߥ߆ߞߚ߇‫ޔ‬ቄ▵߇ㅴ߻ߦߟࠇߡㅒォߒߚ‫ޔߪߣߎߩߎޕ‬
߅ߘࠄߊ㘩‛ଏ⛎ߩ㆑޿ߦࠃࠅ‫⥄ޔ‬ὼ⟲ߢߪ᫪ᨋ߇‫ޔ‬ടኂ⟲ߢߪㄘ࿾ߥߤ߇⟲⪭೑↪ߩቄ▵ᄌൻߩࠠ࡯ߣߥߞߡ޿
ࠆߎߣࠍ␜ໂߔࠆ‫ޕ‬2 ⟲ߩ⟲⪭೑↪ࠍᲧセߔࠆߣ‫ߩߤޔ‬ቄ▵ߦ߅޿ߡ߽ടኂ⟲ߩᣇ߇ㄘ࿾ߩ೑↪߇ᄙ߆ߞߚ‫ޔߚ߹ޕ‬
ೋᄐ߿᥅ᄐߦߪടኂ⟲ߩᣇ߇ᐢ⪲᮸ᨋߩ೑↪߇ᄙ߆ߞߚ‫ޔߪࠇߎޕ‬ㄘ࿾ߦᩕ㙃ଔߩ㜞޿૞‛߇㓸ਛ⊛ߦ޽ࠆߚ߼ߣ
⠨߃ࠄࠇࠆ‫ޔߚ߹ޕ‬ㄘ࿾ߣᐢ⪲᮸ᨋ߇㓞ធߒߡ޿ࠆߎߣ߇ᄙ޿ߚ߼‫ޔ‬ㄘ࿾ࠍ೑↪ߔࠆߎߣ߇⚿ᨐߣߒߡᐢ⪲᮸ᨋߩ
೑↪ࠍ㜞߼ߚߚ߼ߣ⠨߃ࠄࠇࠆ‫ޕ‬
B-3 ࠨ࡞⣄㜑៊்ࡕ࠺࡞ࠍ↪޿ߚゲ⚝ౣ↢㒖ኂ࿃ሶߣߘߩ᛫૕ߦࠃࠆ␹⚻࿁〝ୃᓳߦ㑐ߔࠆ⎇ⓥ
ጊਅବ⧷㧘ਛᎹᶈ㧘ਛ᧛↱㚅㧘૒⮮ᓆୃ㧔ᄢ㒋ᄢ࡮㒮࡮ක㧕 ᚲౝኻᔕ⠪㧦㜞↰᣽ᒾ
⣄㜑៊்ߦࠃࠅ‫ޔ‬ਛᨔㆇേ࿁〝ߩゲ⚝߇ಾᢿߐࠇ‫ޔ‬਄ਅ⢇ߩ㤗∽߇ᗖ⿠ߐࠇࠆ‫ޕ‬ᚑ૕ߦ߅޿ߡ⣄㜑߇៊்ߐࠇࠆ
ߣ‫ޔ‬ᯏ⢻࿁ᓳ߇ᦼᓙߢ߈ߥ޿ߎߣ߇ᄙ޿‫ߩߘޕ‬ℂ↱ߩ߭ߣߟߣߒߡ‫ޔ‬ゲ⚝ౣ↢ࠍ㒖ኂߔࠆ࿃ሶߩሽ࿷߇޽ߍࠄࠇࠆ‫ޕ‬
ᚒ‫ޔߪޘ‬ᣂⷙߩゲ⚝ౣ↢㒖ኂ࿃ሶߣߒߡ Repulsive guidance molecule-a (RGMa) ࠍหቯߒ‫ޔ‬⣄㜑៊்ࡕ࠺࡞࡜࠶࠻ߦ
RGMa ߩᯏ⢻ࠍਛ๺ߔࠆ᛫૕ࠍᛩਈߔࠆߎߣߦࠃࠅ‫ޔ‬ㆇേᯏ⢻ߩ࿁ᓳ߇ଦㅴߐࠇࠆߎߣࠍႎ๔ߒߚ‫ޕ‬㧔Hata, et al.,
2006㧕ᧄ⎇ⓥߪ‫ޔ‬㔤㐳㘃ߦ߅޿ߡ RGMa ߇ゲ⚝ౣ↢㒖ኂ࿃ሶߣߒߡ௛޿ߡ޿ࠆ߆ߤ߁߆ߦߟ޿ߡᬌ⸽ߔࠆߎߣࠍ
⋡⊛ߣߒߡ޿ࠆ‫ޔߡ޿↪ࠍ࡞ࠩࠥࠞࠕޕ‬ㆇേࠍ೙ᓮߔࠆ⊹⾰⣄㜑〝ࠍ㗅ⴕᕈ࠻࡟࡯ࠨ࡯ߢ࡜ࡌ࡞ߒ‫ ߩߘޔ‬3 ㅳ㑆ᓟ
ߦ⣄㜑៊்㧔hemisection㧕ࠍᣉߒߚ‫ ߩߘߦࠄߐޕ‬10 ᣣᓟߦ‫ޔ‬⣄㜑ࠍណขߒ‫∉఺ޔ‬ᨴ⦡ᴺߦߡ៊்ㇱ๟࿐ߩ RGMa
⊒⃻ࠍⷰኤߒߡ޿ࠆߣߎࠈߢ޽ࠆ‫ߚ߹ޕ‬ᄢ⣖ㆇേ㊁㗔ၞ‫⾰⊹ޔ‬⣄㜑〝ߦ߅ߌࠆ‫ޔ‬RGMa ฃኈ૕ߢ޽ࠆ neogenin ߩ⊒
⃻߽⏕⹺ߒߡ޿ࠆ‫ޕ‬᛫૕ᴦ≮ࠍᣉߔ੍ቯߩࠕࠞࠥࠩ࡞ 2 㗡ߦߟ޿ߡߪ‫ޔ‬ㆇേᯏ⢻⹏ଔ⺖㗴ߩ࠻࡟࡯࠾ࡦࠣࠍⴕߞߡ
޿ࠆ‫ޕ‬
B-4 ࠨ࡞♽⛔ㅴൻߦ߅ߌࠆ਄⢇ߩ⸃೬ቇ⊛⊒㆐
⷏᧦ኼᄦ㧘TA Aversi-Ferreira㧘ၳᖝ㇢㧔ንጊᄢ࡮කቇ⮎ቇ⎇ⓥㇱ࡮ࠪࠬ࠹ࡓᖱേ⑼ቇ㧕 ᚲౝኻᔕ⠪㧦ਛ᧛స᮸
ࠝࡑࠠࠨ࡞ߪ‫ߣ࠻ࡅ߿࡯ࠫࡦࡄࡦ࠴ޔ‬ห᭽ߦ‫ޔ‬਄⢇ߦࠃࠅ㆏ౕࠍ↪޿ߡ㘩‛ࠍᓧࠆߎߣ߇⍮ࠄࠇߡ޿ࠆ‫ࠄࠇߎޕ‬
㆏ౕ૶↪ߦߪ‫ޔ‬਄⢇╭⡺ߩᓸ⚦ߥࠦࡦ࠻ࡠ࡯࡞߇ᔅⷐߢ޽ࠅ‫ޔ‬㆏ౕ૶↪ࠍⴕ߁ࠝࡑࠠࠨ࡞ߩᚻᜰߪ‫ࡄࡦ࠴߿࠻ࡅޔ‬
ࡦࠫ࡯ߣ⇣ߥࠆ╭ᡰ㈩ߦ߽㑐ࠊࠄߕ‫ߣ࡯ࠫࡦࡄࡦ࠴߿࠻ࡅޔ‬ห᭽ߥᲣᜰኻะᕈࠍ᦭ߒߡ޿ࠆߎߣ߇᣿ࠄ߆ߦߐࠇߡ
޿ࠆ‫৻ޕ‬ᣇ‫ⴕޔߪ࡞ࠩࡦࡎ࠾ޔ‬േቇ⊛⎇ⓥ߿␹⚻↢ℂቇ⊛⎇ⓥߦᄙߊ↪޿ࠄࠇߡ޿ࠆߦ߽㑐ࠊࠄߕ‫ߩߘޔ‬਄⢇ߩ⸃
೬ቇ⊛․ᓽߪᲧセ⊛ਇ᣿ߢ޽ࠆ‫⎇ᧄޕ‬ⓥߢߪ‫ࠆߌ߅ߦ࡞ࠩࡦࡎ࠾ޔ‬਄⢇ߩ⸃೬ቇ⊛․ᓽࠍ‫ޔ‬⡺⌒⸃೬ߦࠃࠅ⸃ᨆߒ
ߚ‫⚿ߩߘޕ‬ᨐ‫ߩ࡞ࠩࡦࡎ࠾ޔ‬਄⣨ੑ㗡╭‫ޔ‬਄⣨╭‫ޔ‬ὖญ⣨╭‫ޔ‬਄⣨ਃ㗡╭߅ࠃ߮ dorsoepitroclear muscle㧔ࠨ࡞㘃ߩ
ߺ‫ߢ࠻ࡅޔ‬ሽ࿷ߖߕ㧕ߦ߅ߌࠆ␹⚻ᡰ㈩߅ࠃ߮ⴊ▤ߩಽᏓߪ‫ߩઁޔ‬㔤㐳㘃ߣหߓߢ޽ߞߚ߇‫⿠ߩߘޔ‬ᆎߣ஗ᱛߩઃ
⌕ㇱ૏߇⇣ߥࠆߎߣ߇᣿ࠄ߆ߦߥߞߚ‫⃻ޕ‬࿷‫ޔ‬೨⣨╭ߩ⡺⌒⸃೬ࠍⴕߞߡ߅ࠅ‫੹ޔ‬ᓟో࠺࡯࠲ࠍ Comparative
Anatomical Index (IAC) ࠍ↪޿ߡᢙ୯ൻߔࠆߎߣߦࠃࠅ‫ࠍ࠻ࡅޔ‬฽߻ઁߩ㔤㐳㘃ߣᲧセߒ‫ޔ‬਄⢇ߩㅴൻㆊ⒟ࠍ᣿ࠄ
߆ߦߒߡ޿ߊ੍ቯߢ޽ࠆ‫ޕ‬
B-5 ࡑ࡯ࡕ࠮࠶࠻ߦ߅ߌࠆ㙃⢒୘૕ߩࠝࠠࠪ࠻ࠪࡦỚᐲ
㥲⮮ᘏሶ㧔᧲੩ᄢ࡮㒮࡮✚วᢥൻ㧕 ᚲౝኻᔕ⠪㧦ਛ᧛స᮸
ㄭᐕ‫␠ߣࡦࠪ࠻ࠪࠠࠝࠆ޽ߢߟ৻ߩ࠼࠴ࡊࡍ⚻␹ޔ‬ળᕈߦ㑐ߔࠆ⎇ⓥ߇ߍߞᱤ㘃ߢ⋓ࠎߦ߅ߎߥࠊࠇߡ޿ࠆ‫ࠝޕ‬
ࠠࠪ࠻ࠪࡦߪ‫␠ޔ‬ળᕈߩ╙৻ᱠߣ⠨߃ࠄࠇࠆ⹺⍮࡮ⴕേߦ㑐ࠊߞߡ޿ࠆߎߣ߇ࠊ߆ߞߡ߅ࠅ‫ࠍ࠻ࡅޔ‬ኻ⽎ߣߒߚ⎇
ⓥ߽⋓ࠎߣߥߞߡ޿ࠆ‫࠻ࡅߛ߹޿ޔߒ߆ߒޕ‬એᄖߩ㔤㐳㘃ߦ߅ߌࠆ␠ળⴕേߣࠝࠠࠪ࠻ࠪࡦߩ㑐ଥߦߟ޿ߡߩ⎇ⓥ
ߪᢙ߇ዋߥ޿‫⎇ᧄޔߢߎߘޕ‬ⓥߢߪ‫ޔ‬දห❥ᱺࠍ߅ߎߥ߁ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ࠍኻ⽎ߦ‫ޔ‬Უⷫߛߌߢߥߊῳⷫߩࠝ
ࠠࠪ࠻ࠪࡦỚᐲ߇‫ޔ‬ᅧᆼ࡮಴↥࡮㙃⢒ⴕേߦࠃࠅߤߩࠃ߁ߦᄌൻߔࠆ߆ࠍ⺞ߴࠆߎߣࠍ⋡⊛ߣߒߚ‫ޕ‬
੃ᐜఽ߇޿ࠆῳⷫ୘૕ߣන⁛㘺⢒ࠝࠬ୘૕߆ࠄណዩߒ‫ޔ‬Ꮢ⽼ߩࠝࠠࠪ࠻᷹ࠪቯ↪ EIA ࠠ࠶࠻㧔ࡅ࠻‫↪ࠬ࠙ࡑޔ‬㧕
ࠍ↪޿ߡ‫ޔ‬ዩਛߩࠝࠠࠪ࠻ࠪࡦ㊂ࠍᲧセߒߚ߇‫⟲ޔ‬㑆ߦᏅߪߺࠄࠇߥ߆ߞߚ‫ߚ߹ޕ‬಴↥೨ᓟߢࠝࠠࠪ࠻ࠪࡦ㊂ߦᄌ
ൻ߇ߺࠄࠇࠆ߆ࠍᬌ⸛ߔࠆߚ߼ߦ‫ޔ‬ᅧᆼਛ㨪಴↥ᓟߩ❥ᱺࡍࠕࠃࠅណዩࠍ߅ߎߥ޿‫ࡦࠪ࠻ࠪࠠࠝޔ‬㊂ߩ᷹ቯࠍ߅ߎ
ߥߞߚ‫⚿ߩߘޕ‬ᨐ‫ߪߢࠬࡔޔ‬಴↥ᒰᣣ㧔ⴊᶧ߽ዩߦᷙ౉ߒߡ޿ߚ㧕ߦࡇ࡯ࠢࠍ␜ߒ‫ޔ‬಴↥೨ࠃࠅ߽಴↥ᓟߢ୯߇㜞
޿௑ะ߇ߺࠄࠇߚ‫ߪߢࠬࠝޔߚ߹ޕ‬೨ᓟߢᄢ߈ߥᄌൻ߇ߺࠄࠇߥ߆ߞߚ‫ޕ‬
ߘߩᓟ‫࠼࠴ࡊࡍߩࡦࠪ࠻ࠪࠠࠝࠆߌ߅ߦ࠻࠶࠮ࡕ࡯ࡑޔ‬㈩೉߇ઁߩູ੃㘃ߣ⇣ߥࠆߎߣ߇⊒⴫ߐࠇߚߚ߼‫࡯ࡑޔ‬
ࡕ࠮࠶࠻ߩࠝࠠࠪ࠻ࠪࡦࠍวᚑߒ‫ޔ‬ౣᐲ᷹ቯ♽ߩᅷᒰᕈࠍᬌ⸛ߒߡ޿ࠆ‫ޕ‬
㧙㧙
B-6 㔤㐳㘃ߦ߅ߌࠆ␹⚻ᩕ㙃࿃ሶߩ♖␹ᯏ⢻⊒㆐ߦਈ߃ࠆᓇ㗀
㇊ᵄብਯ㧘ᄖጊ⧷๺㧘㔍ᵄኼ᣿㧔ᣂẟᄢ࡮⣖⎇࡮ಽሶ␹⚻↢‛㧕㧘᳓㊁⺈ ᚲౝኻᔕ⠪㧦ਛ᧛స᮸
⛔วᄬ⺞∝ߩ⊒∝ේ࿃ߩ߭ߣߟߣߒߡ‫ޔ‬⢝ఽ߿ᣂ↢ఽᦼߦ߅ߌࠆᧃ᪳ᕈࠨࠗ࠻ࠞࠗࡦߦࠃࠆ⣖⊒㆐㓚ኂ߇ᗐቯߐ
ࠇߡ޿ࠆ‫ޕ‬ᣂ↢੿ࡑ࠙ࠬߩ⊹ਅߦ␹⚻ᩕ㙃ᕈࠨࠗ࠻ࠞࠗࡦߢ޽ࠆ਄⊹ᚑ㐳࿃ሶ㧔EGF㧕߿࠾ࡘ࡯࡟ࠣ࡝ࡦ 1 ߥߤࠍ
ᛩਈߔࠆߎߣߢ‫⹺ޔ‬⍮ⴕേ⇣Ᏹ߇ᚑᾫᓟߦ⺃⊒ߐࠇࠆߎߣ߇⍮ࠄࠇߡ޿ࠆ‫ߩߎޔߒ߆ߒޕ‬઒⺑߇ࡅ࠻ࠍ฽߻㔤㐳㘃
ߦ߽ㆡ↪ߢ߈߁ࠆ߆‫⇼ޔ‬໧߽ᄙ޿‫ߢߎߘޕ‬ᚑ㐳ߩᣧ޿㔤㐳㘃ߢ޽ࠆࡑ࡯ࡕ࠮࠶࠻ࠍ↪޿‫ᧄޔ‬઒⺑ߩ㔤㐳㘃ߢߩᬌ⸽
ࠍ⹜ߺߚ‫␠ߪ࠻࠶࠮ࡕ࡯ࡑߦ․ޕ‬ળⴕേᕈߩ㜞޿㔤㐳㘃ߢ޽ࠅ‫␠ޔ‬ળⴕേ߇்ኂߐࠇࠆ⛔วᄬ⺞∝ࠍ⹏ଔߔࠆߦ
ߪ‫ޔ‬ℂᗐ⊛ߥታ㛎േ‛ߣ⠨߃ߚ‫ᧄޕ‬ᐕᐲߪ‫ޔ‬ᤓᐕᐲߦ⾆⎇ⓥᚲߦߡታᣉߒߚᣂ↢ఽࡑ࡯ࡕ࠮࠶࠻߳ߩ EGF ߩᛩਈ
േ‛ߩ⚻ㆊⷰኤࠍ⛮⛯ߔࠆߣߣ߽ߦ‫ޔ‬ᅧᆼࡑ࡯ࡕ࠮࠶࠻Უ૕߳ EGF ߩᛩਈ⹜㛎ࠍታᣉߒߚ‫ޕ‬
ᅧᆼਛߩࡑ࡯ࡕ࠮࠶࠻Უⷫߪ 2 ඘ࠍណ↪ߒߡᛩਈࠍⴕߞߚ‫↪ߩߎޕ‬㊂ߪ‫ޔ‬੐೨ߩᚑ㐳ࡑ࡯ࡕ࠮࠶࠻߳ߩ EGF ᛩ
ਈߦࠃࠆᓇ㗀⹏ଔ⹜㛎ߦࠃࠅ᳿ቯߐࠇߚࡊࡠ࠻ࠦ࡯࡞ߦᓥߞߚ‫৻ߣࠆࠃߦ࡞࡯ࠦ࠻ࡠࡊߩߎޕ‬ㆊᕈߩཌྷฯߪ⊒↢ߔ
ࠆ߇‫ޔ‬૕㊀Ⴧട߽޽ࠅ‫ⴕޔ‬േ߽ߔߋᱜᏱൻߔࠆ‫࠻࠶࠮ࡕ࡯ࡑޕ‬⢝ఽߦ߅޿ߡᲣሶ㑆ߩ Urero-Placentral Barrier (UPB)
߇ቢᚑߔࠆᅧᆼਛᦼ߹ߢࠍᮡ⊛ᤨᦼߣߒߚ‫৻ޕ‬඘ߩࡑ࡯ࡕ࠮࠶࠻Უ૕ߦߪ‫ޔ‬ᅧᆼ 9㨪11 ㅳߢ EGF ᛩਈ‫߁߽ޔ‬Უ૕
ߢߪᅧᆼ 12㨪14 ㅳࠍ⋡቟ߦ‫ޔ‬0.3mg/kg ૕㊀ߢ 1 ᣣ 1 ࿁‫ޔ‬㓒ᣣ⊹ਅᛩਈ㧔⸘ 5 ࿁ᛩਈ㧕ࠍታᣉߒߚ‫ޕ‬಴↥ᤨᦼ߽ᱜ
Ᏹߢ‫ޔ‬ᣂ↢ఽߩ૕㊀߽ᱜᏱߢ޽ߞߚ‫ߦ․ޕ‬ᣂ↢ఽߩⴕേ╬ߦ⇣Ᏹߪ⷗ࠄࠇߥ߆ߞߚ߇‫ޔ‬಴↢⋥ᓟߦ਄ਅ㗶ฦ‫੃ߦޘ‬
ಾᱤ߇ 4 ᧄ㧔╙ 1 ಾᱤ 2 ᧄ‫ ╙ޔ‬2 ಾᱤ 2 ᧄ㧕‫ ⸘ޔ‬8 ᧄ↢߃ߡ߅ࠅ‫ޔ‬EGF ᛩਈߩ෻ᔕߩ⽎ᓽߢ޽ࠆᱤ಴⧘ߩଦㅴ⃻⽎
߇ⷰኤߐࠇߚ‫ޔߪߣߎߩߎޕ‬Უ૕߆ࠄ UPB ࠍㅘㆊߒߡ EGF ߇⢝ఽߦ⒖ⴕߒߚߎߣࠍᗧ๧ߔࠆ‫੹ޕ‬ᓟ‫ޔ‬ᚑ㐳ࠍᓙߞ
ߡⴕേᜰᮡߩቯ㊂ൻࠍⴕ޿‫ޔ‬EGF ߣ⹺⍮ⴕേ⊒㆐ߩ㑐ㅪࠍᬌ⸽ߒߚ޿‫ޕ‬
B-7 ࡑ࡯ࡕ࠮࠶࠻⣖ߦ߅ߌࠆࠪ࠽ࡊࠬน႟ᕈ㑐ㅪⰮ⊕࠼࡟ࡉ࡝ࡦߩ఺∉⚵❱ൻቇ⊛⸃ᨆ
⊕የᥓ᣿㧘ఽፉિᒾ㧘᫃↰⵨⾆㧔⟲㚍ᄢቇ࡮㒮࡮ක㧕 ᚲౝኻᔕ⠪㧦ਛ᧛స᮸
࠼࡟ࡉ࡝ࡦߪ᮸⁁⓭⿠ࠬࡄࠗࡦߦዪ࿷ߔࠆࠕࠢ࠴ࡦ✢⛽⚿ว࠲ࡦࡄࠢ⾰ߢ޽ࠅ‫ߩࠬࡊ࠽ࠪޔ‬น႟ᕈߦߣߞߡ㊀ⷐ
ߥ௛߈ࠍߒߡ޿ࠆ‫⎇ᧄޕ‬ⓥߪ‫ߩࡦ࡝ࡉ࡟࠼ߡ޿↪ࠍ࠻࠶࠮ࡕ࡯ࡑޔ‬㜞ᰴ⣖ᯏ⢻ߦ߅ߌࠆ௛߈ߩ⸃᣿ࠍ⋡ᜰߒ‫࡯ࡑޔ‬
ࡕ࠮࠶࠻⣖ౝߩ࠼࡟ࡉ࡝ࡦߩಽᏓࠍ఺∉⚵❱ൻቇᨴ⦡ߢ⸃ᨆߒߚ‫ޕ‬㓽ߩࡑ࡯ࡕ࠮࠶࠻ 3 ඘ࠍᷓ㤗㈮ᓟ‫ࡦ࡝ࡑ࡞ࡎޔ‬
ṁᶧߢἠᵹ࿕ቯߒ⣖ࠍขࠅ಴ߒߚ‫ޕ‬ᶐẃ࿕ቯᓟ‫ࠬ࡯ࡠࠢࠬޔ‬ṁᶧ߳⟎឵ߒߚ‫ޕ‬ಓ⚿ᓟ‫ޔ‬౰⁁ᢿߢ 12m ߩಓ⚿ಾ ࠍ૞ᚑߒ‫ޔ‬᛫࠼࡟ࡉ࡝ࡦ᛫૕‫ޔ‬᛫ࠪ࠽ࡊ࠻ࡈࠖࠫࡦ᛫૕‫ޔ‬᛫࠳ࡉ࡞ࠦ࡞࠴ࡦ᛫૕ࠍ↪޿‫ޔ‬DAB ߦࠃࠅ఺∉⚵❱ൻ
ቇᨴ⦡ࠍⴕߞߚ‫ߩࡦ࡝ࡉ࡟࠼ߪߢࠬ࠙ࡑޕ‬ዪ࿷߇⹺߼ࠄࠇߥ޿ౝ஥ਛ㓒ߦ߅޿ߡ‫ߩࡦ࡝ࡉ࡟࠼ߪߢ࠻࠶࠮ࡕ࡯ࡑޔ‬
ᒝ޿ᨴ⦡߇⷗ࠄࠇߚ‫ޕ‬ㅢᏱߪ࠼࡟ࡉ࡝ࡦ߇Ớᨴߔࠆ㗔ၞߪࠪ࠽ࡊࠬࡑ࡯ࠞ࡯ߢ޽ࠆࠪ࠽ࡊ࠻ࡈࠖࠫࡦ޽ࠆ޿ߪᐜ⧯
␹⚻⚦⢩ߩࡑ࡯ࠞ࡯ߢ޽ࠆ࠳ࡉ࡞ࠦ࡞࠴ࡦ߇Ớᨴߔࠆ߇‫ߩߎޔ‬ౝ஥ਛᩭߦ߅޿ߡߪߎࠇࠄߩࡑ࡯ࠞ࡯ߪỚᨴߒߥ߆
ߞߚ‫৻ޕ‬ᣇ‫࡞ࡇࡠ࡯ࡘ࠾ߩઁޔ‬㗔ၞߢߪ࠼࡟ࡉ࡝ࡦߣࠪ࠽ࡊ࠻ࡈࠖࠫࡦߩỚᨴㇱ૏ߪ৻⥌ߒ‫ޔߚ߹ޔ‬⣖ቶਅᏪㇱߦ
߅ߌࠆ⒖േਛߩ␹⚻೨㚟⚦⢩ߪ࠼࡟ࡉ࡝ࡦߣ࠳ࡉ࡞ࠦ࡞࠴ࡦ߇Ớᨴߒߡ޿ߚ‫ޕ‬
B-8 ᛫߁ߟ⮎ߦࠃࠆࡑ࡯ࡕ࠮࠶࠻ᶏ㚍ᱤ⁁࿁㗰☸⚦⢩ߩ⣕ᚑᾫലᨐ
ᄢᐔ⠹ม㧔⮮↰଻ⴡᄢ࡮✚ක⎇࡮ࠪࠬ࠹ࡓක㧕㧘⪤ේ⧷㓶㧘᣽ሶᶈቁ㧘਄㊁෹↢㧔⮮↰଻ⴡᄢ࡮✚ක⎇㧕 ᚲౝኻ
ᔕ⠪㧦ਛ᧛స᮸
ᚒ‫⛔ޔߪޘ‬วᄬ⺞∝߿෺ᭂᕈ᳇ಽ㓚ኂߥߤ♖␹∔ᖚ᭽ߩⴕേ⇣Ᏹࠍ␜ߔᄙᢙ♽⛔ߩࡑ࠙ࠬߦ߅޿ߡᧂᚑᾫᶏ㚍ᱤ
⁁࿁㧔iDG㧕߇↢ߓߡ޿ࠆߎߣࠍ⷗಴ߒߡ޿ࠆ‫৻ޕ‬ᣇ‫ޔ‬㊁↢ဳࡑ࠙ࠬߦኻߒߡ‫ޔ‬᛫߁ߟ⮎ߩᘟᕈᛩਈ߿⣖㔚᠄࡚ࠪ
࠶ࠢࠍಣ⟎ߔࠆߣ‫ޔ‬iDG ߇↢ߓࠆߎߣ߇ಽ߆ߞߚ‫ޔߜࠊߥߔޕ‬iDG ߩ⺃ዉ߇߁ߟ∛ߦኻߔࠆᴦ≮ലᨐߢ޽ࠆߣ޿߁
ᣂߒ޿઒⺑߇␜ໂߐࠇࠆ‫ޕ‬iDG ߩੱᎿ⊛ߥᱜᏱൻߣ⺃ዉ߇ታ⃻ߢ߈ࠇ߫‫⛔ޔ‬วᄬ⺞∝‫ޔ‬෺ᭂᕈ᳇ಽ㓚ኂ‫ߥ∛ߟ߁ޔ‬
ߤߩ♖␹∔ᖚߩᴦ≮ᴺߦ⚿߮ߟߊߎߣ߇ᦼᓙߢ߈ࠆ‫⎇ᧄޕ‬ⓥߢߪ‫ޔ‬᛫߁ߟ⮎ߦࠃࠆ iDG ߩ⺃ዉߦߟ޿ߡ‫ࡕ࡯ࡑޔ‬
࠮࠶࠻ߦ᛫߁ߟ೷ߩ৻ߟߢ޽ࠆࡈ࡞ࠝࠠ࠮࠴ࡦߩ᡼಴ࡍ࡟࠶࠻㧔3 mg/kg/day㧘60 day-release㧕ࠍ⊹ਅߦၒ߼ㄟߺ‫ޔ‬
ߘߩ೨ᓟ 4 ㅳ㑆ߦ࿑ᒻᑯ೎⺖㗴‫ޔ‬ㅒォቇ⠌⺖㗴‫ޔ‬ᵴേ㊂᷹ቯࠍታᣉߒ‫ߩߘޔ‬ᓟ⣖ࠍ࿕ቯߒ⚵❱ቇ⊛⸃ᨆࠍⴕߞߚ‫ޕ‬
ߘߩ⚿ᨐ‫ޔ‬ቇ⠌⺖㗴߿ᵴേ㊂ߦᄢ߈ߥᄌൻߪⷰኤߐࠇߥ߆ߞߚ߇‫ޔ‬ᱤ⁁࿁ߦ߅޿ߡᧂᚑᾫ㗰☸⚦⢩ߩࡑ࡯ࠞ࡯ߢ޽
ࠆ Doublecortin ߇Ⴧടߒߡ޿ࠆߎߣ߇᣿ࠄ߆ߣߥߞߚ‫⚿ߩߎޕ‬ᨐߪ‫ޔ‬㔤㐳㘃ߢ߽ iDG ࠍ⺃ዉߢ߈ࠆߎߣࠍ␜ໂߒߡ
޿ࠆ‫੹ޕ‬ᓟ‫ޔ‬iDG ൻߦࠃࠆⴕേᄌൻߦߟ޿ߡ⸃ᨆࠍㅴ߼ߡ޿߈ߚ޿‫ޕ‬
B-9 ࠾ࡎࡦࠩ࡞ߩࠕࡔ࡯ࡃᗵᨴߦ㑐ߔࠆ∉ቇ⎇ⓥ
ᯌ⵨ม㧔᧲ᶏᄢ࡮ක㧕㧘ዊᨋᱜⷙ㧔ᘮᔕᄢ࡮ක㧕㧘ᩉື㓶㧔㐳ፒᄢ࡮ᾲ⎇㧕 ᚲౝኻᔕ⠪㧦ᐔ੗໪ਭ
ᦨㄭ‫ࡃ࡯ࡔࠕ∯⿒ޔ‬㧔Entamoeba histolytica㧕ߣߪ⇣ߥࠆ∛ේࠕࡔ࡯ࡃ E. nuttalli ߇‫࡞ࠨޔ‬㘃߆ࠄ⷗ߟ߆ߞߡ޿ࠆ‫ޕ‬
ᧄ⎇ⓥߩ⋡⊛ߪ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ⣺▤ነ↢ࠕࡔ࡯ࡃߩᗵᨴታᘒࠍ᣿ࠄ߆ߦߔࠆߎߣߢ޽ࠆ‫ޕ‬
ችፒ⋵ਠ㑆Ꮢߩᐘፉߦ߅޿ߡ‫ޔ‬㊁↢࠾ࡎࡦࠩ࡞ߩ♮ଢ 32 ᬌ૕ࠍណขߒߚ‫⋥ޕ‬ធ㏜ᬌߢߪ‫⯻✢♮ޔ‬ෆ‫ޔ‬㖊⯻ෆ‫ޔ‬
⢗⯻ෆ‫ޔ‬⣺⚿▵⯻ෆ‫ޔ‬R ဳ੿⯻߇ᬌ಴ߐࠇߚ߇‫ߪ࠻ࠬࠪࡃ࡯ࡔࠕޔ‬ᬌ಴ߐࠇߥ߆ߞߚ‫♮ޔߢߎߘޕ‬ଢ߆ࠄ DNA ࠍ
᛽಴ߒ‫ޔࡃ࡯ࡔࠕ∯⿒ޔ‬E. dispar㧘E. nuttalli㧘E. chattoni㧘ᄢ⣺ࠕࡔ࡯ࡃ㧔E. coli㧕ߦߟ޿ߡ PCR ᴺߦࠃࠆᬌ಴ࠍ⹜
ߺߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬E. chattoni ߇ 26 ᬌ૕㧔81㧑㧕ߦ߅޿ߡ㓁ᕈߣߥߞߚ‫ ߩઁߩߘޔߒ߆ߒޕ‬4 ⒳㘃ߩࠕࡔ࡯ࡃߪోߊ
ᬌ಴ߐࠇߥ߆ߞߚ‫♮ޔߚ߹ޕ‬ଢᬌ૕ߦߟ޿ߡ‫↰ޔ‬ㄝජ⪲ၭ࿾ߦࠃࠆࠕࡔ࡯ࡃߩಽ㔌ၭ㙃ࠍ⹜ߺߚ߇‫ߡߴߔޔ‬㒶ᕈߢ
޽ߞߚ‫ޕ‬
ߎࠇ߹ߢߩઁ࿾ၞߦ߅ߌࠆ⺞ᩏߢ߽ E. chattoni ᗵᨴߪ㜞₸ߦ⹺߼ࠄࠇߚ߇‫ઁޔ‬⒳ߩࠕࡔ࡯ࡃ߽หᤨߦᬌ಴ߐࠇࠆ
ߎߣ߇ᄙߊ‫ޔ‬࿾ၞߦࠃࠆᗵᨴࠕࡔ࡯ࡃ⒳ߩ㆑޿߇⏕⹺ߐࠇߚ‫ޔ߅ߥޕ‬ណଢߦ㑐ߒߡߪ‫ޔ‬ᐘፉⷰኤᚲߩ㋈᧛᳁ߣ౰࿾
᳁ߩදജࠍ޿ߚߛ޿ߚ‫ޕ‬
㧙㧙
B-10 ࠾ࡎࡦࠩ࡞࡮࠻࡜ࡦࠬࠨࠗ࡟࠴ࡦߩ↢ൻቇ⊛ᕈ⁁ߦ㑐ߔࠆ⎇ⓥ
ਛ᧛⚩৻ᦶ㧔ṑ⾐ක⑼ᄢ࡮േ‛↢๮⑼ቇ⎇ⓥ࠮ࡦ࠲࡯㧕 ᚲౝኻᔕ⠪㧦㋈ᧁ᮸ℂ
ⴊᷡ࠲ࡦࡄࠢߩ࠻࡜ࡦࠬࠨࠗ࡟࠴ࡦ㧔TTR㧕ߪ‫ోߩ࠻ࡅޔ‬りᕈ⠧㦂ᕈࠕࡒࡠࠗ࠼∝㧔SSA㧕ߩේ࿃ߢ޽ࠆ‫ޕ‬TTR
ߪ྾㊂૕ߢ቟ቯߒ‫ޔ‬න㊂૕ߦߥࠆߣ✢⛽ේᕈ߇Ⴧߒ‫ࠍ࠼ࠗࡠࡒࠕޔ‬ᒻᚑߒ߿ߔߊߥࠆ‫ߜࠊߥߔޕ‬න㊂૕ߩഀว߇Ⴧ
߃ࠆߎߣߢ SSA ⊒∝ߩน⢻ᕈ߇੍᷹ߢ߈ࠆ‫ޕ‬ട㦂ߪ TTR ߇ਇ቟ቯൻߔࠆ৻ߟߩⷐ࿃ߢ޽ࠅ‫ޔ‬એ೨ߩᧄ౒ห೑↪⎇
ⓥߦߡᔃ⤳߳ߩ TTR ᴉ⌕߇⏕⹺ߐࠇߚࠨ࡞⒳‫ ߽ߢ࡞ࠩࡦࡎ࠾ޔ‬TTR ߩ྾㊂૕߆ࠄන㊂૕߳ߩട㦂ᕈᄌൻ߇‫ࡒࠕޔ‬
ࡠࠗ࠼ᒻᚑߦ㑐ਈߒߡ޿ࠆ߆ߤ߁߆ᬌ⸛ߒߚ‫ޕ‬
ᐔᚑ 20‫ޔ‬21 ᐕᐲߩᧄ౒ห೑↪⎇ⓥߢ∛ℂ⚵❱ቇ⊛ᬌ⚝ࠍⴕߞߚ 33 ଀ߩ߁ߜ‫ޔ‬ᱷ૛ⴊᷡ߇଻▤ߐࠇߡ޿ߚ 11 ଀
㧔2 ᱦ߆ࠄ 29 ᱦ‫ޔ‬ᐔဋ 10.1 ᱦ㧕ࠍ↪޿ߡ‫ޔ‬TTR ߦኻߔࠆ࠙ࠚࠬ࠲ࡦࡉࡠ࠶࠻ߣ ELISA ࠍⴕߞߚ‫ޕ‬ㆊ෰ߩ౒ห೑↪
⎇ⓥߢᔃ⤳ߦ TTR ᴉ⌕ࠍ⹺߼ߚ୘૕㧔34 ᱦ㧕ߩⴊᷡߪᱷߞߡߥ߆ߞߚ‫ޕ‬
࠙ࠚࠬ࠲ࡦࡉࡠ࠶࠻ߢߪᦨ㜞㦂ߩ 29 ᱦߩേ‛ߢන㊂૕ߩࡃࡦ࠼߇ᒝ߆ߞߚ߇‫ߩઁߩߘޔ‬േ‛ߢߪᐕ㦂ߦ઻߁௑
ะߪ⷗ࠄࠇߥ߆ߞߚ‫ޕ‬ELISA ߢߪ 2ޯ11 ᱦߩ 8 ଀ߩᐔဋ߇ 36.8mg/dL ߛߞߚߩߦኻߒ‫ޔ‬13ޯ29 ᱦߩ 3 ଀ߩᐔဋ߇
27.1 mg/dL ߢ㜞㦂ߩേ‛ߩᣇ߇ⴊᷡỚᐲߪૐ߆ߞߚ‫ޕ‬
TTR ↥↢㊂ߪട㦂ߦ઻߁⢄⚦⢩ߩᯏ⢻ૐਅߥߤߢᷫዋߔࠆߎߣ߇ផ᷹ߐࠇߚ‫࠼ࠗࡠࡒࠕޕ‬ේᕈߩᒝ޿න㊂૕ߩ಴
⃻ߦߪ‫ޔ‬30 ᐕㄭ޿⛘ኻ⊛ߥᤨ㑆߇ᔅⷐߢ‫ޔ‬34 ᱦߩ㜞㦂࠾ࡎࡦࠩ࡞ߦߩߺ TTR ᴉ⌕߇⷗ࠄࠇߚ⚵❱ቇ⊛ᄌൻࠍᡰᜬ
ߔࠆ⚿ᨐߣߥߞߚ‫ޕ‬
B-11 ࠪࠞߩⓨ㑆ಽᏓߦ෸߷ߔࠨ࡞ߩᓇ㗀
឴ᆄ⋥᮸㧔ർᶏ㆏ᄢ࡮ർᣇ↢‛࿤ࡈࠖ࡯࡞࠼⑼ቇ࠮ࡦ࠲࡯࡮๺᱌ጊ⎇ⓥᨋ㧕 ᚲౝኻᔕ⠪㧦ඨ⼱๋㇢
ደਭፉోၞߩ⺞ᩏ߆ࠄ‫ޔ‬᥊ⷰࠬࠤ࡯࡞㧔ࡑࠢࡠࠬࠤ࡯࡞㧕ߦ߅ߌࠆࠪࠞኒᐲߦߪࠨ࡞ኒᐲ߇ᱜߩⷐ࿃ߣߒߡ௛ߊ
ߎߣ߇␜ߐࠇߡ޿ࠆ‫⎇ᧄߢߎߘޕ‬ⓥߢߪ↢ᕷ࿾ࠬࠤ࡯࡞㧔ࡒࠢࡠࠬࠤ࡯࡞㧕ߦ߅޿ߡ߽‫ߩࠞࠪޔ‬ⓨ㑆ಽᏓ߇ࠨ࡞ߩ
ಽᏓߦⷙቯߐࠇߡ޿ࠆ߆ࠍᬌ⸛ߒߚ‫ޕ‬
ደਭፉ⷏ㇱ࿾ၞߩඨጊ࿾඙߅ࠃ߮Ꮉේ࿾඙ߦߘࠇߙࠇ⚂ 5km ߩⷰኤ࡞࡯࠻ࠍ⸳ቯߒ‫ⷰޔ‬ኤ࡞࡯࠻ࠍᱠ߈ߥ߇ࠄ‫ޔ‬
ࠪࠞߣࠨ࡞ߩ⊒⷗ߦദ߼ߚ‫ޕ‬േ‛ࠍ⊒⷗ߒߚ႐วߦߪ‫ޔ‬േ‛⒳࡮ዻᕈ࡮૏⟎࡮ᤨ㑆ߥߤࠍ⸥㍳ߒߚ‫⺞ߩߎޕ‬ᩏߪฦ
࿾඙ߢ 3 ࿁ߕߟⴕߞߚ‫ޕ‬෼㓸ߒߚ࠺࡯࠲ߪ GIS ߢಽᨆߒߚ‫⺞ޕ‬ᩏ࡞࡯࠻߆ࠄ 20m એౝߦߟ޿ߡߪ߽ࠪࠞࠨ࡞߽⷗
⪭ߣߒ߇ߥ޿ߣ⠨߃ࠄࠇߚߩߢ‫▸ߩߘޔ‬࿐ࠍಽᨆኻ⽎ߣߒߚ‫⺞ޔߕ߹ޕ‬ᩏ࡞࡯࠻ਔ஥ߦ 20m ߩࡃ࠶ࡈࠔ࡯ࠍ૞ࠅ
㕙Ⓧࠍ᷹ߞߚ‫ߚߒ⷗⊒ࠍ࡞ࠨޔߚ߹ޕ‬૏⟎ߩ๟ㄝ 20m ߦ߽ࡃ࠶ࡈࠔ࡯ࠍ૞ᚑߒ‫ޔ‬㕙Ⓧࠍ᳞߼ߚ‫⺞ޔߡߒߘޕ‬ᩏ࡞
࡯࠻ࡃ࠶ࡈࠔ࡯㕙Ⓧߦኻߔࠆࠨ࡞ࡃ࠶ࡈࠔ࡯㕙ⓍߩᲧ₸ࠍ▚಴ߒߚ‫ߩߎޕ‬Ყ₸߇‫⺞ޔ‬ᩏ࡞࡯࠻ࡃ࠶ࡈࠔ࡯ౝߢ⊒⷗
ߒߚࠪࠞᢙߣࠨ࡞ࡃ࠶ࡈࠔ࡯ౝߢ⊒⷗ߒߚࠪࠞᢙߩᲧ₸ߣ᦭ᗧߦ⇣ߥࠆ߆ࠍᬌቯߒߚ‫ޕ‬
⺞ᩏ࿾඙ߏߣߦ 3 ࿁ಽߩ࠺࡯࠲ࠍࡊ࡯࡞ߒߡᬌቯࠍⴕߞߚᚲ‫ߩࠄߜߤޔ‬࿾඙ߢ߽᦭ᗧߦࠨ࡞ࡃ࠶ࡈࠔ࡯ౝߢࠪࠞ
߇ᄙߊ⊒⷗ߐࠇࠆߎߣ߇␜ߐࠇߚ‫ޔߒߛߚޕ‬1 ࿁ߏߣߦಽᨆߒߚ႐วߦߪ᦭ᗧᏅ߇ߢߥ޿႐ว߇޽ߞߚ‫ޕ‬᥊ⷰࠬࠤ
࡯࡞ߢ⷗ࠄࠇߚࠨ࡞߇ࠪࠞߩኒᐲߦਈ߃ࠆᓇ㗀ߪ‫↢ޔ‬ᕷ࿾ࠬࠤ࡯࡞ߦ߅ߌࠆࠨ࡞㧙ࠪࠞߩⓨ㑆㈩⟎ߦၮߠߊ߽ߩߢ
޽ࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬
B-12 ᦭ኂ㚟㒰୘૕ࠍ↪޿ߚ྾࿖ߩ㊁↢࠾ࡎࡦࠩ࡞୘૕⟲ߩ․ᓽಽᨆ
⼱࿾᫪⑲ੑ㧔྾࿖⥄ὼผ⑼ቇ⎇ⓥ࠮ࡦ࠲࡯㧕 ᚲౝኻᔕ⠪㧦ᷰㆻ㇌ᄦ
྾࿖ߢߪᄙߊߩ࿾ၞߢ࠾ࡎࡦࠩ࡞ߦࠃࠆㄘ૞‛ⵍኂ߇⊒↢ߒ‫߁઻ߦࠇߘޔ‬㚟㒰ᵴേ߇ⴕࠊࠇߡ޿ࠆ‫߇ߥߒ߆ߒޕ‬
ࠄ‫ޔ‬㚟㒰ߐࠇߚ୘૕߆ࠄߩᖱႎ෼㓸ߪ㚟㒰ᢙ‫ޔ‬ᚑ㐳Ბ㓏‫ޔ‬ᕈ೎⒟ᐲߦ㒢ࠄࠇ‫‛↢ޔ‬ቇ⊛ߥᖱႎߦ㑐ߒߡߪ߶ߣࠎߤ
⸥㍳ߐࠇߕߦၒ⸳ಣಽߐࠇߡ߈ߚ‫ޔߚ߹ޕ‬྾࿖↥࠾ࡎࡦࠩ࡞ߩᮡᧄᢙ߽㕖Ᏹߦዋߥߊ‫ޔ‬྾࿖↥࿾ၞ୘૕⟲ߩ⎇ⓥߪ
߶ߣࠎߤߥߐࠇߡ޿ߥ޿‫⎇ᧄޕ‬ⓥߪᐔᚑ 22 ᐕᐲࠃࠅߩ⛮⛯⺖㗴ߣߒߡ‫࡞ࠩࡦࡎ࠾ޔ‬྾࿖࿾ၞ୘૕⟲ߦߟ޿ߡ‫↢ޔ‬
‛ቇ⊛․ᓽߥࠄ߮ߦ᦭ኂ㚟㒰⁁ᴫࠍᛠីߔࠆߎߣࠍ⋡⊛ߦⴕߞߚ‫ޕ‬
ᐔᚑ 23 ᐕᐲߪ‫ޔ‬㜞⍮⋵ౝߦ⺞ᩏ࿾ၞࠍ 3 ࿾ၞ⸳ߌᖱႎࠍ෼㓸ߒߚ‫ޕ‬ኻ⽎࿾ၞߪ‫ޔ‬㚅⟤Ꮢ㧔⋵᧲ㇱ㧕‫ޔ‬ਛ࿯૒↸㧔ਛ
ᄩㇱ㧕߅ࠃ߮྾ਁචᏒ㧔⋵⷏ㇱ㧕ߢ޽ࠆ‫ޕ‬
ฦ⺞ᩏኻ⽎࿾ၞ߳ᐔᚑ 23 ᐕ 5 ᦬‫ޔ‬7 ᦬‫ޔ‬10 ᦬‫ޔ‬ᐔᚑ 24 ᐕ 1 ᦬߅ࠃ߮ 3 ᦬ߦ⿞߈‫᦭ޔ‬ኂ㚟㒰୘૕ࠍ 43 ୘૕ฃߌ
౉ࠇߚ‫ޕ‬
ฃߌ౉ࠇߚ୘૕ߦߟ޿ߡ‫ޔ‬㜞੗ᱜᚑᢎ᝼ߣදജߒߡ↢૕߅ࠃ߮㛽ᩰᮡᧄߩ⸘᷹ߣ㛽ᩰᮡᧄൻߒߚ⾗ᢱߩ଻▤ࠍ‫ޔ‬
੹੗໪㓶ಎᢎ᝼ߣදജߒߡಽሶ↢‛ቇ⊛ߥಽᨆࠍⴕߞߚ‫ޕ‬
ߘߩ⚿ᨐ‫ޔ‬྾࿖ߩ࠾ࡎࡦࠩ࡞ߪㆮવ⊛ߥᄌ⇣ᕈ߇ዋߥ޿ߥߤ‫․ޔ‬ᓽ߇ᓢ‫ߦޘ‬᣿ࠄ߆ߦߥߞߡ߈ߡ޿ࠆ‫ޕ‬
B-13 ࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ࠍ↪޿ߚ✛ౝ㓚ᕈ✂⤑࡮ⷞⷡਛᨔ㓚ኂ⊒∝ᯏᐨߩ⸃᣿
ේ⧷ᓆ㧘᎑Ỉ㓷శ㧘ਛ᧛ା੺㧔ጘ㒂⮎⑼ᄢ࡮⮎ല⸃ᨆቇ㧕 ᚲౝኻᔕ⠪㧦ਛ᧛స᮸
ᚒ߇࿖ߦ߅޿ߡ‫✛ޔ‬ౝ㓚ߪਛㅜᄬ᣿ේ࿃ߩ╙৻૏ࠍභ߼ࠆ∔ᖚߢ޽ࠆ‫✛ޔࠄ߇ߥߒ߆ߒޕ‬ౝ㓚ߩ⊒∝߅ࠃ߮✂⤑
㓚ኂㅴⴕߩᯏᐨߦߟ޿ߡߪ߶ߣࠎߤ⸃᣿ߐࠇߡ޿ߥ޿‫ޔߢߎߘޕ‬ᚒ‫✛ߪޘ‬ౝ㓚ᕈ✂⤑࡮ⷞⷡਛᨔ⊒∝㓚ኂߩᯏᐨ⸃
᣿ࠍ⋡⊛ߣߒߡ‫ߡ޿↪ࠍ࠻࠶࠮ࡕ࡯ࡑࡦࡕࠦޔ‬ᘟᕈ㜞⌒࿶✛ౝ㓚ࡕ࠺࡞ߩ૞⵾ࠍ⹜ߺߚ‫࡞࡯࠲ࡆ࡞ࡃ࠻ࡦࡍޕ‬㤗㈮
ਅߦࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ 4 㗡ߩᏀ⌒ߩ೨⌒ㇱ✢⛽ᩇᏪߦࠕ࡞ࠧࡦ࡟࡯ࠩ࡯ࠍᾖ኿ߒ‫⌒ޔ‬ᚱ᳓ߩឃ಴ࠍᛥ೙ߒߚ‫࡟ޕ‬
࡯ࠩ࡯ᾖ኿ߪ 2 ㅳ㑆㓒ߢ 2 ࿁ߦಽߌߡᾖ኿ߒߚ‫⌒ޕ‬࿶ߪࠤ࠲ࡒࡦ㧛ࡔ࠹࠼ࡒࠫࡦ૬↪㤗㈮ਅߢ⌒࿶⸘ࠍ↪޿ߡ᷹ቯ
ߒ‫⌒ޔ‬ᐩ౮⌀ߪᚻᜬߜᑼ⌒ᐩࠞࡔ࡜㧔Micron III㧕ࠍ↪޿ߡ᠟ᓇߒߚ‫࡯ࠩ࡯࡟ޕ‬ᾖ኿ 4 ㅳᓟࠃࠅᜬ⛯⊛ߥ⌒࿶਄᣹߇
ⷰኤߐࠇߚ‫⌒ߦࠄߐޕ‬ᐩᚲ⷗ࠃࠅ‫࡯ࠩ࡯࡟ޔ‬ᾖ኿ 9 ㅳᓟߦ߅޿ߡ㜞⌒࿶⌒ߩⷞ␹⚻੃㗡ㇱߩⴊ▤ߩ᣿ࠄ߆ߥ෼❗߅
㧙㧙
ࠃ߮ᵻ޿੃㗡㒱ಳߩ᜛ᄢ߇ⷰኤߐࠇߚ‫ߚ߹ޕ‬ណขߒߚⷞ␹⚻ߩ࠻࡞ࠗࠫࡦࡉ࡞࡯ᨴ⦡ߦࠃࠅ‫࡯ࠩ࡯࡟ޔ‬ᾖ኿⌒ߦ߅
ߌࠆⷞ␹⚻ゲ╷㓚ኂ߇ⷰኤߐࠇߚ‫ޕ‬
એ਄ߩ⚿ᨐ߆ࠄ‫ޔ‬਎⇇ߢೋ߼ߡࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻ߦ߅޿ߡᘟᕈ㜞⌒࿶ࡕ࠺࡞ࠍ૞⵾ߔࠆߎߣߦᚑഞߒߚ‫ޕ‬
B-14 DNA analysis of White Headed Langur and feeding plants.
Yin Lijie㧔ർ੩ᄢ࡮↢๮⑼ቇቇ㒮㧕㧘Qin Dagong㧘Pan Wenshi㧘Yao Jinxian㧘Xian Danlin ᚲౝኻᔕ⠪㧦੹੗໪㓶
From 2011 to 2012, we collected 22 feces samples on December 2010, January and March 2011, January 2012 from two
populations of white-headed langurs. And 42 plants species that langurs eating or not were collected in March 2011. We have
gotten about 15 plants species by DNA analyzing from two feces samples.
On March 2012, I have done DNA analysis of six feces samples of white-headed langurs under directing by Dr. Hiroo Imai
and his student Nami Suzuki in PRI. We extracted and cloned DNA in feces from six samples and selected three samples to do
DNA sequencing. From these three samples, we got several plant species that langurs ate. Based on the results, we discussed
next study plan in the future. It is important for the research that how to collect feces of langurs, sample size and sample
seasonality.
In addition, I did a presentation to introduce wildlife conservation in Peking University center for Nature and Society and
research of white-headed langurs when I visited PRI. Although my English is not so good, I hope it may expedite the
community and cooperation between PKU Center for Nature and society and Primates Research Institute㧘Kyoto University.
B-15 ࠕࠞࠥࠩ࡞ᣂ↢ఽߦ߅ߌࠆⷞⷡೝỗߦࠃࠆࠬ࠻࡟ࠬ✭๺ലᨐ
Ꮉ਄ᷡᢥ㧔⡛ᔃᅚሶᄢ࡮ᔃℂ㧕 ᚲౝኻᔕ⠪㧦෹᳗㓷Ꮖ
╩⠪ࠄߪ࠾ࡎࡦࠩ࡞ᣂ↢ఽ߇ណⴊࠍฃߌࠆ႐㕙ߦ‫ࠍ⥇࡯࠳ࡦࡌ࡜߿࠭ࠗࡁ࠻ࠗࡢࡎޔ‬๒␜ߔࠆߣࠬ࠻࡟ࠬ߇✭๺
ߐࠇࠆߎߣࠍ᣿ࠄ߆ߦߒߚ‫ޕ‬
㧔Kawakami, Tomonaga, Suzuki, Primates, 2002,43,73-85㧧Ꮉ਄࡮෹᳗࡮㋈ᧁ‫ੱޔ‬㑆ⅣႺቇ
⎇ⓥ 2009,7 ,89-93㧕ᧄ⎇ⓥߢߪ‫ߩߘޔ‬⍮⷗ࠍᐢߍࠆߚ߼ߦ‫ⷡⷞޔ‬ೝỗࠍ๒␜ߒߡߺࠆ‫ߩ࡞ࠩ࡮࠽࠻ࠝޔߕ߹ޕ‬㗻౮
⌀ࠍ૶߁ߎߣߦߒߚ‫ޕ‬
ᧄᐕᐲߪ࠾ࡎࡦࠩ࡞ߢߪߥߊ‫ ࠬࠝߩ࡞ࠩࠥࠞࠕޔ‬4 㗡ߩ࠺࡯࠲߇ᓧࠄࠇߚ‫ ╙ޕ‬1 ࿁⋡ߩታ㛎ᣣ߇ᐔဋ↢ᓟ 12 ᣣ‫ޔ‬
╙ 2 ࿁⋡ߪ↢ᓟ 19.5 ᣣߢ޽ߞߚ‫ⷡⷞޕ‬ೝỗࠍ๒␜ߒߚ᧦ઙߣ㗻౮⌀ࠍ࡜ࡦ࠳ࡓ࡮࠼࠶࠻ߦߒߚ⛔೙᧦ઙࠍᲧߴߚ‫ޕ‬
ⴕേ⹏ቯߩ⚿ᨐߢߪ‫ޔ‬㗻๒␜ലᨐߪߺࠄࠇߥ߆ߞߚ‫ޕ‬
B-16 㔤㐳㘃ߩฦ⚵❱ߦ߅ߌࠆ๧ⷡᖱႎવ㆐‛⾰ߩሽ࿷
ᮭ↰ᓀ㧘᧻᧛⑲৻㧔ጘ㒂ᄢ㧕 ᚲౝኻᔕ⠪㧦੹੗໪㓶
⃻࿷‫ߢࠬ࠙ࡑ߿࠻ࡅޔ‬๧ⷡฃኈ૕߇⥠ߛߌߢߥߊᶖൻ▤ߥߤߦ߅޿ߡ߽⊒⃻ߒߡ޿ࠆߎߣ߇ಽ߆ߞߡ߈ߡ޿ࠆ‫ޕ‬
ᧄ⎇ⓥߢߪࠦࡕࡦࡑ࡯ࡕ࠮࠶࠻߿ࠕࠞࠥࠩ࡞ߥߤࠍኻ⽎ߦ‫ࠇߘߣ⥠ޔ‬એᄖߩ⤳ེߦ߅޿ߡ‫ޔ‬๧ⷡᖱႎવ㆐‛⾰ߢ޽
ࠆ -gustducin ߣ TRPM5 ߩሽ࿷㊂ࠍ RT-qPCR ᴺߦࠃࠅ⛘ኻቯ㊂ߒߚ‫ࠆߥ⇣ޔߦࠄߐޕ‬⒳‫ࠆߥ⇣ޔ‬ᐕ㦂ߩ㔤㐳㘃ࠍ
Ყセߔࠆߎߣߦࠃࠅ‫ޔ‬๧ⷡᖱႎવ㆐‛⾰ߩ⊒⃻㊂ߩ⒳Ꮕ߿ᐕ㦂Ꮕࠍ⺞ᩏߒߚ‫ޕ‬
ߘߩ⚿ᨐ‫ޔ‬ቯ㊂ߒߚో㔤㐳㘃ߢߪ‫ߣེ⤳ߩઁޔ‬Ყߴߡᶖൻེ♽ߦ߅޿ߡ -gustducin ߣ TRPM5 ߇ᄙ㊂ߦሽ࿷ߔࠆ
ߎߣ߇ಽ߆ߞߚ‫⋤ޔߪߢ࠻࠶࠮ࡕ࡯ࡑࡦࡕࠦޕ‬⣺‫ޔ‬ᄢ⣺ߦ߅޿ߡ‫ߣ⥠ޔ‬ห㊂߽ߒߊߪߘࠇએ਄ߩ -gustducin ߇ሽ
࿷ߒߚ‫ޕ‬TRPM5 ߦ㑐ߒߡ߽‫ޔ‬ᐕ㦂Ꮕ߿୘૕Ꮕߪ޽ࠆ߽ߩߩ‫ޔ‬ዊ⣺‫⋤ޔ‬⣺‫ޔ‬ᄢ⣺ߦ߅޿ߡሽ࿷㊂߇ᄙ߆ߞߚ‫৻ޕ‬ᣇ‫ޔ‬
ࠕࠞࠥࠩ࡞ߥߤߢߪ‫⋤ޔ‬⣺ߦ߅޿ߡሽ࿷㊂߇ᄙ޿ߣ޿߁⚿ᨐߪᓧࠄࠇߥ߆ߞߚ‫⋤ޕ‬⣺ߦ߅ߌࠆ๧ⷡᖱႎવ㆐‛⾰ߩ
ሽ࿷㊂ߩᄙߐߪ‫․ߦ࠻࠶࠮ࡕ࡯ࡑࡦࡕࠦޔ‬ᓽ⊛ߥ⚿ᨐߢ޽ࠆน⢻ᕈ߇㜞޿‫ࠥࠞࠕߣ࠻࠶࠮ࡕ࡯ࡑࡦࡕࠦޔߒ߆ߒޕ‬
ࠩ࡞ߣߢߪ⺞ᩏߒߚᐕ㦂߇⇣ߥࠆߚ߼‫⚿ߩߎޔ‬ᨐߩᏅ߇ᐕ㦂Ꮕߦ⿠࿃ߔࠆน⢻ᕈ߽޽ࠆ‫੹ޕ‬ᓟ‫ߥ⚦⹦ࠅࠃޔ‬ಽᨆࠍ
߅ߎߥߞߡ⏕߆߼ߚ޿ߣ⠨߃ߡ޿ࠆ‫ޕ‬
B-17 㔤㐳㘃๧ⷡฃኈ૕ㆮવሶ⟲ߩ⊒⃻⸃ᨆ
⍹ਣ༑ᦶ㧘㒙ㇱ⟤᮸㧔᧲੩ᄢ࡮㒮࡮ㄘቇ↢๮⑼ቇ㧕 ᚲౝኻᔕ⠪㧦੹੗໪㓶
ࠕࠞࠥࠩ࡞ߩ๧ⷡฃኈ૕ߣਅᵹࠪࠣ࠽࡞વ㆐࿃ሶ⟲ߩ⨧⁁࡮᦭ㇳ੃㗡๧⭫ߦ߅ߌࠆ⊒⃻ࠍ in situ ࡂࠗࡉ࡝࠳ࠗ
࠯࡯࡚ࠪࡦᴺࠍ↪޿ߡ⸃ᨆߒߚ‫Ⱟޔߚ߹ޕ‬శੑ㊀ in situ ࡂࠗࡉ࡝࠳ࠗ࠯࡯࡚ࠪࡦᴺࠍ↪޿ߡ‫ޔ‬ฦㆮવሶࠍ⊒⃻ߔࠆ
⚦⢩ߩ⋧㑐㑐ଥࠍ⺞ߴߚ‫ޕ‬
TAS1R‫ޔ‬TAS2R‫ޔ‬PKD1L3 ߩฦ๧ⷡฃኈ૕ㆮવሶߪ‫ޔ‬๧⭫ਛߩߘࠇߙࠇ⇣ߥࠆ๧⚦⢩ߢឃઁ⊛ߦ⊒⃻ߒߡ߅ࠅ‫ޔ‬ၮ
ᧄ๧ߏߣߦฃኈߔࠆ๧⚦⢩߇ಽ߆ࠇߡ޿ࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬ᰴߦ‫ޔ‬ฦㆮવሶߦߟ޿ߡ⹦⚦ߦ⸃ᨆߒߚߣߎࠈ‫ޔ‬
TAS1R ࡈࠔࡒ࡝࡯ߦ㑐ߒߡߪ‫੃ߩࠇߕ޿ޔ‬㗡ߢ߽ TAS1R1 ߣ TAS1R2 ߇੕޿ߦ೎‫ߩޘ‬๧⚦⢩ߦ⊒⃻ߒߡ޿ߚ‫ޕ‬PKD1L3
ߪ‫᦭ޔ‬ㇳ੃㗡ߛߌߢߥߊ⨧⁁੃㗡ߢ߽⊒⃻ߒߡ޿ߚ‫ޕ‬G ࠲ࡦࡄࠢ⾰ߦ㑐ߒߡߪ‫ޔ‬GNAT3 ߇⨧⁁੃㗡ߣ᦭ㇳ੃㗡ߩ
ਔᣇߢ⊒⃻ߔࠆߩߦኻߒߡ‫ޔ‬GNA14 ߪ᦭ㇳ੃㗡ߛߌߢ⊒⃻ߔࠆߎߣߪࡑ࠙ࠬߣห᭽ߢ޽ߞߚ‫᦭ޔߒ߆ߒޕ‬ㇳ੃㗡
ߦ߅޿ߡ GNA14 ࠍ⊒⃻ߔࠆ⚦⢩ߪ‫ߩߘޔ‬ഀว߇ࡑ࠙ࠬߣᲧセߒߡ㗼⪺ߦዊߐߊ‫ޔ‬TAS1R2 ⊒⃻⚦⢩ߣឃઁ⊛ߢ޽ࠆ
ὐߪ⇣ߥߞߡ޿ߚ‫ޕ‬એ਄ߩታ㛎⚿ᨐ߆ࠄ‫ޔ‬หߓູ੃㘃ߦዻߔࠆ㔤㐳㘃ߣߍߞᱤ㘃ߢߪ‫ޔ‬๧ⷡ㑐ㅪㆮવሶߩ⊒⃻᭽ᑼ
ߦ㑐ߒߡ‫౒ޔ‬ㅢὐߣ⋧㆑ὐ߇޽ࠆߎߣ߇᣿ࠄ߆ߣߥߞߚ‫ޕ‬
B-18 ੩ㇺ⋆࿾ർ✼ߦ↢ᕷߔࠆ࠾ࡎࡦࠩ࡞ߩ଻ో↢ᘒቇ⊛⎇ⓥ
⷏㇍㗼㆐㧔੩ㇺᐭ㠽₞໧㗴⎇ⓥળ㧕 ᚲౝኻᔕ⠪㧦ᷰㆻ㇌ᄦ
⥄ቛߣߘߩㄭㄝ㧔ⴕ᡽⊛ߦߪ੩ㇺᏒᏀ੩඙㕒Ꮢ‫ޔ‬㕷㚍‫ޔ‬ጤୖ‫߮ࠃ߅ޔ‬ർ඙਄⾐⨃ߥߤ㧕ߦߪ࠾ࡎࡦࠩ࡞ߩ⟲ࠇ߇
㧙㧙
಴ᴚߒ‫₎“ޔ‬ኂ”߇ࠃߊ↢ߓߡ޿ࠆ‫₎ߪ⑳ޕ‬ኂߩታᘒࠍ⍮ࠅ‫ߩߘޔ‬シᷫࠍ࿑ࠆߚ߼‫ޔ‬2004 ᐕߦߎߩ⟲ࠇߩ⺞ᩏࠍᆎ߼
ߚ‫ޕ‬2008 ᐕએᓟߪᲤᄛ‫ߩߘޔ‬ᣣߩ⟲ࠇߩ⒖േߒߚ࡞࡯࠻ߣᴱߞߚ႐ᚲ‫߮ࠃ߅ޔ‬㧔น⢻ߥࠄ߫㧕⠉ᣣߩ੍ᗐㅴ〝ࠍ
ࡉࡠࠣߢ⊒ାߔࠆߎߣࠍᆎ߼ߚ‫ ߪߦࠣࡠࡉޕ‬2012 ᐕ 3 ᦬ 31 ᣣ⃻࿷ߢ⚂ 37000 ੱ૛߇ࠕࠢ࠮ࠬߒߡ޿ࠆ‫ࡠࡉߩߎޕ‬
ࠣߦࠃߞߡ 2 ߟߩჿ‫ޔ‬
‫₎ޟ‬ኂኻ╷ࠍ┙ߡ߿ߔߊߥߞߚ‫ޔߣޠ‬
‫↢ߩ࡞ࠨޟ‬ᘒߦ⥝๧߇ࠊ޿ߡ߈ߚ‫ࠍޠ‬⡞ߊࠃ߁ߦߥߞߚ‫ޕ‬
⟲ࠇߩࠨࠗ࠭ߪ 2005 ᐕ㧦⚂ 50 㗡‫ޔ‬2011 ᐕ㧦⚂ 60 㗡ߢ‫ ߩߎޔ‬7 ᐕ㑆ߩᄌൻߪዋߥ޿‫ޕ‬หᦼ㑆‫ ߩ࡯ࡃࡦࡔࠇ⟲ޔ‬52
㨪59㧑㧦ࠕࠞࡦࡏ࠙ߣࠦ࠼ࡕ‫ޔ‬30㨪38㧑㧦↢ᱺน⢻㧔5 ᱦએ਄㧕ࡔࠬߢ‫ޔ‬᭴ᚑߦ㑐ߒߡ߽ᄌൻߪዋߥ޿‫⟲ޔߛߚޕ‬
ࠇߩਛߦ޿ࠆᚑࠝࠬ߇ᦨೋ 2 㗡ߛߌߢ޽ߞߚ߇‫ޔ‬2008 ᐕએ㒠ߪ 6-7 㗡ߦჇ߃ߚ‫ޕ‬
ߎߩ⟲ࠇߩሽ࿷߇ೋ߼ߡ⏕⹺ߐࠇߚߩߪ 1970 ᐕઍ೨ඨߢ޽ࠅ‫ޔ‬ᒰᤨߩⴕേၞߪ㕷㚍‫ޔ⦁⾆ޕ‬ੑࡁἑ‫ޔ‬㔕ࡩ⇌ߢ
޽ߞߚ‫⺞߇⑳ޕ‬ᩏࠍᆎ߼ߚߎࠈߪ⾆⦁‫ޔ‬㔕ࡩ⇌ߦⴕߊߎߣߪߥߊ‫ߢ߁߸ߞ޿ߩߘޔ‬Ꮢේ‫ޔ‬ጤୖ‫ޔ‬਄⾐⨃߇ⴕേၞߦ
฽߹ࠇ‫ߪ࠭ࠗࠨߩߘޔ‬ᩰᲑߦᄢ߈ߊߥߞߡ޿ߚ‫ߪࠇ⟲ޔߜࠊߥߔޕ‬⠹૞࿾ߩዋߥ޿ਛጊ㑆࿾ၞࠃࠅ߽↰࿦ㇱߩᄙ޿
ᐔ㊁ㇱࠍࠃߊേߊࠃ߁ߦߥߞߡ޿ߚ‫ߩߎޕ‬௑ะߪߎߩ 7 ᐕ㑆ߢ߽ࠃࠅ㗼⪺ߦߥࠅ‫߫߃଀ޔ‬㕷㚍ࠍ⸰ࠇࠆ㗫ᐲߪᐕ‫ޘ‬
ૐਅߒ‫ޔ‬ᤓᐕߪߘߎߢᴱߞߚߎߣߪ৻࿁߽ߥ߆ߞߚ‫ޕ‬
B-19 ࠾ࡎࡦࠩ࡞㊁↢⟲ߩᐕ㦂᭴ᚑߣᚑ㐳࡮ട㦂ࡄ࠲࡯ࡦ
ጯᧄ⌀ᑿ㧔ࢃ㊁↢േ‛଻⼔▤ℂ੐ോᚲ㑐⷏ಽቶ㧕 ᚲౝኻᔕ⠪㧦ự↰ⓛ
⟲ࠇߩඨᢙ߇᝝₪ߐࠇߚ↲⾐ A ⟲㧔ṑ⾐⋵㧕ߩో㛽ᩰᮡᧄ 124 ୘૕ߦߟ޿ߡ‫ޔ‬ᐕ㦂ᩏቯࠍታᣉߔࠆߣߣ߽ߦ‫ޔ‬
ᚑ㐳࡮ട㦂ߦ઻ߞߡᄌൻߔࠆߣផ᷹ߐࠇࠆㇱ૏ߩ⸘᷹ࠍⴕߞߚ‫ޕ‬
ᐕ㦂᭴ᚑ㧔ḩᐕ㦂㧕ߪ 0 ᚽ㧦23 㗡‫ޔ‬1 ᚽ㧦27 㗡‫ޔ‬2 ᚽ㧦20 㗡‫ޔ‬3 ᚽ㧦14 㗡‫ޔ‬4 ᚽ㧦5 㗡‫ޔ‬5 ᚽ㧦11 㗡‫ޔ‬6 ᚽ㧦2
㗡‫ޔ‬7 ᚽ㧦1 㗡‫ޔ‬8 ᚽ㧦6 㗡‫ޔ‬9 ᚽ㧦4 㗡‫ޔ‬11 ᚽ㧦2 㗡‫ޔ‬12 ᚽ㧦3 㗡‫ޔ‬13 ᚽ㧦2 㗡‫ޔ‬14 ᚽ‫ޔ‬16 ᚽ‫ޔ‬17 ᚽ‫ޔ‬25 ᚽ߇ฦ 1
㗡ߢ޽ߞߚ‫ޕ‬4 ᚽ߅ࠃ߮ 5 ᚽߣߒߚ୘૕ߩᱤߩ⪚಴੤឵ߣ࠮ࡔࡦ࠻⾰ᐕベߘࠇߙࠇ߆ࠄផቯߐࠇࠆᐕ㦂ߦߪ‫ޔ‬ዋߥ
ߊߣ߽ 1 ᚽߩ߫ࠄߟ߈߇޽ߞߚ‫ޕ‬
ᚑ㐳ߦߣ߽ߥ߁ᄌൻߣߒߡ‫ޔ‬྾⢇㐳㛽㐳㧔ᐳ㜞ߦኻߔࠆ␜ᢙ㧕ߪ 7 ᚽ߹ߚߪ 8 ᚽߦ⥋ࠆ߹ߢჇടߒߟߠߌߚ‫ޕ‬㗻
㐳㧔㗡⬄㐳ߦኻߔࠆ␜ᢙ㧕ߪᚑ㐳ߔࠆߦߒߚ߇޿Ⴧടߒ‫ޔ‬ᚑ₞ߦ㆐ߒߚᓟ߽ട㦂ߣߣ߽ߦჇടߒߟߠߌߚ‫ޕ‬
㜞㦂ߣ૏⟎ߠߌࠄࠇࠆ 25 ᚽ㧔ࡔࠬ㧕ߣ‫ޔ‬7 ᚽ߆ࠄ 17 ᚽߩࡔࠬ 15 ୘૕ಽߩฦ⣶ᬁߩ⣻஥㗡የ㐳㧔⢛஥㗡የ㐳ߦ
ኻߔࠆ␜ᢙ㧕ࠍᲧセߒߚߣߎࠈ‫ޔ‬ඨᢙߢዊߐߊ‫ޔ‬ᱷࠅߩඨᢙߢߪᄢ߈߆ߞߚ‫ޕ‬
B-20 ਅർඨፉ⣁㊁ᴛߦ߅ߌࠆ㊁↢࠾ࡎࡦࠩ࡞ߩ୘૕⟲േᘒߣ଻ోߩߚ߼ߩ⻉໧㗴
᧻ጟผᦶ㧘ਛጊ⵨ℂ㧔ਅർඨፉߩࠨ࡞⺞ᩏળ㧕 ᚲౝኻᔕ㓶⠪㧦ᷰㆻ㇌ᄦ
ਅർඨፉ⣁㊁ᴛ A-87 ⟲㧔ጊߩ⟲ࠇ㧕ߣ A2-85 ⟲㧔᳃ኅ๟ㄝߩ⟲ࠇ㧕ߩ୘૕ᢙߪଐὼჇട௑ะߦ޽ࠅ‫ޔ‬಴↥₸ߪ
ߘࠇߙࠇ 63㧑㧔ᤓᐕ 30%㧕ߣ 90㧑㧔ห 50㧑㧕ߢ޽ߞߚ‫ޕ‬
A2-85 ⟲ߩ 0 ᱦߩᱫ੢₸߇ᤓᐕᐲ 50㧑ߣ଀ᐕߦᲧߴ㜞ߊ‫࡞ࠨޔ‬ㅊ޿›ߩዉ౉ߦࠃࠆ⟲ࠇߩᡬੂේ࿃ߣ⠨߃ࠄࠇߚ
߇‫੹ޔ‬ᐕᐲߩᱫ੢₸ߪ 0㧑ߢ޽ߞߚ‫࡞ࠨޕ‬ㅊ޿›ߩዉ౉ߦࠃࠅㄘ૞‛߳ߩଐሽߪᷫዋߒߚ߇‫ࠆࠃߦࠇߘޔ‬ᩕ㙃⁁ᘒ
ߩᖡൻ߇‫ޔ‬಴↢₸߿ᱫ੢₸ߦᓇ㗀෸߷ߒߡߪ޿ߥ޿ߣ⠨߃ࠄࠇࠆ‫ޕ‬
୘૕ᢙߩჇടߦ઻޿ㆆേၞ߽ଐὼ‫᧲ޔ‬ᣇะߦ᜛ᄢ௑ะߦ޽ࠅ‫ޔ‬೑↪㗫ᐲߩ㜞޿࿾ၞ߽᧲߳ࠪࡈ࠻ߒߡ޿ࠆ‫ޕ‬
ᤓᐕᐲߩ౻ߪ‫ޔ‬27 ᐕ߱ࠅߣ߽޿ࠊࠇࠆ㒠㔐㊂ߩᄙߐߢ‫ޔ‬2012 ᐕ 4 ᦬⃻࿷‫ޔ‬ᄢ᏷ߦᬀ‛ߩ⧘็߈߇ㆃࠇߡ޿ࠆ߇‫ޔ‬
ᄢ㊂ᱫߪ⿠߈ߡ߅ࠄߕ‫ߩߎޔ‬࿾ၞߩ࠾ࡎࡦࠩ࡞୘૕⟲ߩჇട௑ะߦᓇ㗀ߪߥߐߘ߁ߢ޽ࠆ‫ޕ‬
B-21 ࠾ࡎࡦࠩ࡞㑐▵ฃേᛶ᛫․ᕈߩ⸘᷹
⩆ේ⋥㆏㧔ᘮᙥ⟵Ⴖᄢ࡮ℂᎿ࡮ᯏ᪾Ꮏ㧕 ᚲౝኻᔕ⠪㧦ᐔፒ㍈⍫
࠾ࡎࡦࠩ࡞ߩੑ⿷ᱠⴕߪ‫ߣࠇߘߩ࠻ࡅޔ‬Ყセߒߡਅ⢇߇ో૕⊛ߦᦛ߇ߞߚ⁁ᘒߢⴕࠊࠇࠆ‫╭ޔߪࠇߎޕ‬㛽ᩰ♽ߩ
ᒻᘒ࡮᭴ㅧ⊛೙⚂ߦࠃࠅⷙቯߐࠇࠆਅ⢇㧔ᓟ⢇㧕㑐▵ߩนേ․ᕈ߇‫ޔ‬ਔ⠪ߢ⇣ߥߞߡ޿ࠆߎߣߦ⿠࿃ߔࠆߣ⠨߃ࠄ
ࠇࠆ߇‫ߤࠎߣ߶ߪࡓ࠭࠾ࠞࡔߥ⚦⹦ߩߘޔ‬᣿ࠄ߆ߦߥߞߡ޿ߥ޿‫ࠍߣߎߩߎޕ‬᣿ࠄ߆ߣߔࠆߚ߼ߦߪ‫࡞ࠩࡦࡎ࠾ޔ‬
ᣂ㞲ዱ૕ࠍ↪޿ߡਅ⢇╭㛽ᩰ♽ߩฃേᛶ᛫․ᕈࠍ⸘᷹ߒ‫ޔ‬ੑ⿷ᱠⴕߩㅴൻࠍ⠨ኤߔࠆ਄ߢ㊀ⷐߥਅ⢇㑐▵ߩนേ೙
⚂ߣ╭㛽ᩰᒻᘒ࡮᭴ㅧߩ㑐ଥࠍ᣿ࠄ߆ߦߔࠆߎߣ߇㊀ⷐߢ޽ࠆ‫ޕ‬
ᧄᐕߪߘߩ╙৻Ბ㓏ߣߒߡ‫ޔ‬ਅ⢇╭㛽ᩰ♽ߩ㑐▵ฃേᛶ᛫․ᕈߩ⸘᷹ᚻᴺߩౕ૕⊛ᬌ⸛ࠍⴕߞߚ‫ޕ‬ᬌ⸛ߦߪ‫࠾ޔ‬
ࡎࡦࠩ࡞ 1 ୘૕㧔ࠝࠬ࡮7.3kg㧕ߩᣂ㞲ዱ૕ࠍ↪޿ߚ‫ౕޕ‬૕⊛ߦߪᧄᮡᧄࠍ╙৻⣶ᬁߢ೨ᓟߦಽ㔌ߒ‫ޔ‬ቢోߥ࠾ࡎ
ࡦࠩ࡞ਅ⢇╭㛽ᩰ♽ߩ㑐▵ࠍฃേ⊛ߦേ߆ߔߎߣߢ⸘᷹ᣇᴺࠍᬌ⸛ߒߚ‫ޕ‬ᬌ⸛ߩ⚿ᨐ‫ޔ‬㑐▵ߩㄭ૏ㇱࠍⓨ㑆ߦ࿕ቯ
ߒ‫ޔ‬㆙૏ㇱߦขࠅઃߌߚࡅࡕࠍ࠺ࠫ࠲࡞ᚻ߫߆ࠅࠍ੺ߒߡᒁߞᒛࠆߎߣߢ‫ޔ‬㑐▵ⷺᐲߩᄌൻߦߣ߽ߥ߁㑐▵ฃേᛶ
᛫․ᕈߩᄌൻࠍቯ㊂ൻߢ߈ࠆߣ⠨߃ࠄࠇߚ‫੹ޕ‬ᓟᧄᚻᴺߦၮߠ޿ߡ⸘᷹ࠍㅴ߼ࠆ੍ቯߢ޽ࠆ‫ޕ‬
B-22 㔤㐳㘃ߦ߅ߌࠆᤨⓨ㑆⊛ߥኻ⽎㑆㑐ଥߩℂ⸃ߦ㑐ߔࠆ⎇ⓥ
᧛੗ජኼሶ㧔₹Ꮉᄢ࡮⣖⑼ቇ⎇ⓥᚲ㧕 ᚲౝኻᔕ⠪㧦෹᳗㓷Ꮖ
ᄙߊߩኻ⽎߇ሽ࿷ߔࠆᄖ⇇ߢㆡᔕ⊛ߦ߰ࠆ߹߁ߚ߼ߦߪ‫ޔ‬ኻ⽎ࠍߘࠇߙࠇಣℂߔࠆߩߢߪߥߊ‫ߩߘޔ‬㑆ߦ૗ࠄ߆
ߩ㑐ଥᕈࠍ⷗ߟߌ‫ޔ‬㑐ㅪઃߌࠍߒߡᛒ߁ߩ߇ല₸⊛ߢ޽ࠆ‫ޔ߫߃଀ߪࠇߘޕ‬ኻ⽎㑆ߩ⷗ߚ⋡߿ᯏ⢻ߩ㘃ૃᕈ‫ޔ‬ⓨ㑆
⊛ߥㄭធߥߤߢ߽น⢻ߢ޽ࠆ‫♽ᤨޔߚ߹ޕ‬೉਄ߩ㑐ଥᕈߢ߽ኻ⽎ࠍ㑐ㅪઃߌࠆߎߣ߇ߢ߈ࠆ‫ޕ‬A ߣ޿߁ኻ⽎߇಴⃻
ߒߚᓟߦ‫ޔ‬B ߣ޿߁ኻ⽎߇಴⃻ߔࠆߣ‫ ߪߜߚ⑳ޔ‬A ߣ B ߩ㑆ߦ㑐ଥࠍ⷗಴ߒ‫ޔ‬੕޿ࠍ㑐ㅪઃߌࠆ‫ߡߒߘޕ‬㕙⊕޿
ߎߣߦ‫ޟޔ‬AB‫৻ߩޠ‬ᣇะߛߌࠍ⚻㛎ߒߚߦ߽㑐ࠊࠄߕ‫ޟޔ‬B ߩᓟߦߪ A ߇಴⃻ߔࠆ㧔BA㧕
‫߁޿ߣޠ‬ኻ⒓⊛ߥ㑐
ଥࠍᦼᓙߔࠆ‫⺰ߪࠇߎޕ‬ℂ⊛ߦᔅߕߒ߽ᱜߒ޿⸶ߢߪߥ޿߇‫ੱޔ‬㑆߇߽ߟᒝ޿ࡃࠗࠕࠬߩ߭ߣߟߢ‫ߩߜߚ⑳ޔ‬ᕁ⠨
㧙㧙
ࠍᣇะߠߌࠆ߽ߩߢ޽ࠆ‫ߥ߁ࠃߩߎޕ‬௑ะߪࡅ࠻એᄖߩ㔤㐳㘃㧔࠴ࡦࡄࡦࠫ࡯߇㧕ߢ߽⷗ࠄࠇࠆߩߛࠈ߁߆‫߹ߟޕ‬
ࠅ‫ޟ߽࡯ࠫࡦࡄࡦ࠴ޔ‬AB‫ࠅ➅ࠍޠ‬㄰ߒ⚻㛎ߒߚᓟߦ‫ޔ‬
‫ޟ‬BA’‫ޟߊߥߪߢޠ‬BA‫ᦼࠍޠ‬ᓙߔࠆߩߛࠈ߁߆‫ᧄޕ‬ᐕ
ᐲߩ⎇ⓥߢߪ‫ߩߎޔ‬ᬌ⸛ߩ╙৻ᱠߣߒߡ‫ޟ߇࡯ࠫࡦࡄࡦ࠴ߕ߹ޔ‬AB‫߁޿ߣޠ‬ᕭᗧ⊛ߥ㑐ଥࠍ⸠✵ߢߪߥߊ⢻േ
‫ޟ‬AB‫ࠅ➅ߦߢߔ߁޿ߣޠ‬㄰ߒ⚻㛎ߒߚ‫ᦼޔ‬ᓙㅢࠅߩ
⊛ߦቇ⠌ߔࠆ߆ߤ߁߆ࠍ⺞ߴߚ‫⺖ޕ‬㗴ߪᵈⷞ⺖㗴ࠍ↪޿ߡ‫ޔ‬
㑐ଥࠍឭ␜ߒߚᤨߣ‫ޟޔ‬AB’‫⚻߁޿ߣޠ‬㛎ߒߡ޿ߥ޿‫ᦼޔ‬ᓙߦ㆑෻ߒߚ㑐ଥࠍឭ␜ߒߚᤨߣߢᵈⷞᤨ㑆߇㆑߁߆
ߤ߁߆ࠍ⺞ߴߚ‫ޟ߇࡯ࠫࡦࡄࡦ࠴ߒ߽ޕ‬A ߩ޽ߣߪ B ߇಴ࠆ‫ߦࠇߘޔ߫ࠄߥࠆ޿ߡߒ⼂⹺ࠍߣߎ߁޿ߣޠ‬෻ߒߚ㑐
ଥࠍࠃࠅ㐳ߊ⷗ࠆߣ੍ᗐߐࠇࠆ‫ޕ‬ታ㛎ߩ⚿ᨐߪߎࠇࠍᡰᜬߒ‫ ࠄ߆ߕߺޔ߽ߡߊߥߒࠍ✵⸠ߪ࡯ࠫࡦࡄࡦ࠴ޔ‬A ߣ B
ߩኻ⽎㑆ࠍ㑐ㅪઃߌߒ‫ޟޔ‬A ߩᓟߪ B‫߁޿ߣޠ‬ᕭᗧ⊛ߥ࡞࡯࡞ࠍᦼᓙߒߡ޿ࠆߎߣ߇ࠊ߆ߞߚ‫⚿ߩߎޕ‬ᨐࠍ〯߹߃
੹ᓟߪ‫ޔ‬
‫ޟ‬AB‫⚻ߩޠ‬㛎ߛߌߢ‫ޟޔ‬BA‫߁޿ߣޠ‬ኻ⒓⊛ߥ㑐ଥࠍ࠴ࡦࡄࡦࠫ࡯߇᭴▽ߔࠆ߆ߤ߁߆ࠍ⺞ߴߡ޿ߊ‫ޕ‬
㧨ቇળ⊒⴫㧪
1) Murai C, Tomonaga M. Monkeys understand other's attentional state by reading gaze. The 15th annual meeting of the
ASSC.
2) ᧛੗ජኼሶ࡮෹᳗㓷Ꮖ. ߎߞߜࠍ⷗ߡࠆ㧫࠾ࡎࡦࠩ࡞ߦࠃࠆઁ⠪ߩᵈᗧ⁁ᘒߩ⹺⼂. Animal 2011 ᣣᧄേ‛ᔃ
ℂቇળ㧔╙ 71 ࿁㧕࡮ᣣᧄേ‛ⴕേቇળ㧔╙ 30 ࿁㧕࡮ᔕ↪േ‛ⴕേቇળ㧛ᣣᧄኅ⇓▤ℂቇળ㧔2011 ᐕᐲ㧕ว
หᄢળ.
B-23 㔤㐳㘃ߦ߅ߌࠆࠛࡇࠥࡁࡓㅴൻߩ⸃᣿
৻ᩉஜม㧘૒‫⵨ᧁޘ‬ਯ㧘⑔↰ᷧ㧔਻Ꮊᄢ࡮↢ක⎇㧕 ᚲౝኻᔕ⠪㧦ㇹᐽᐢ
ᚒ‫ߪޘ‬㔤㐳㘃ߦ߅ߌࠆࠥࡁࡓㅴൻߣࠛࡇࠥࡁࡓㅴൻߩ㑐ଥࠍ⸃᣿ߔࠆߚ߼‫࡯ࠫࡦࡄࡦ࠴ߣ࠻ࡅޔ‬㧔㔤㐳㘃⎇ⓥᚲ
ߩ㘺⢒୘૕㧕ߩᧃ᪳⊕ⴊ⃿ߩDNAࡔ࠴࡞ൻᲧセ⎇ⓥࠍⴕߞߡ޿ࠆ‫ޔߦߢ߹ࠇߎޕ‬21‫ޔ‬22⇟ᨴ⦡૕ߩࠥࡁࡓ࠲ࠗ࡝ࡦ
ࠣࠕ࡟ࠗࠍ↪޿ߡ16ࠞᚲߩࡔ࠴࡞ൻᏅ⇣㗔ၞ ࠍหቯߒ‫߇ࠄࠇߘޔ‬ㆮવሶ⊒⃻ߣᒝ޿⋧㑐ࠍ␜ߔߎߣࠍ᣿ࠄ߆ߦߒ
ߡ߈ߚ‫ߩࠄࠇߎޔߦࠄߐޕ‬㗔ၞߩࡔ࠴࡞ൻ⁁ᘒࠍࠧ࡝࡜߿ࠝ࡜ࡦ࠙࡯࠲ࡦߩDNAߢ߽⺞ߴࠆߎߣߢ‫ޔ‬CTCF࠲ࡦࡄ
ࠢ⾰ߩ⚿ว㈩೉ߩ಴⃻࡮ᶖᄬ߿ࡑࠗࠢࡠࠨ࠹࡜ࠗ࠻࡝ࡇ࡯࠻ౝߩCpGኒᐲߩᄌൻߦࠃߞߡ‫ޔ‬DNAࡔ࠴࡞ൻ⁁ᘒ߇ᄌ
ൻߒ‫ޔ‬ォ౮⁁ᘒߦᓇ㗀ࠍਈ߃ߡ޿ࠆߎߣࠍ਎⇇ߢೋ߼ߡ␜ߒߚ‫ޕ‬㧔⑔↰ࠄ‫⺰ޔ‬ᢥᛩⓂਛ㧕
ᧄᐕᐲߪߎߩ⎇ⓥࠍߐࠄߦᐢߍࠆߚ߼‫࡞࠴ࡔޔ‬ൻDNA⚿ว࠲ࡦࡄࠢ⾰㧔MBD1㧕ߣᄢⷙᮨࠪ࡯ࠤࡦࠨ࡯ࠍ↪޿ߡ‫ޔ‬
ࡅ࠻ߣ࠴ࡦࡄࡦࠫ࡯ߩࡔ࠴࡞ൻᏅࠍࠥࡁࡓࡢࠗ࠼ߦ⸃ᨆߒߚ㧔MBD-seq㧕
‫ᧃޔߚ߹ޕ‬᪳ⴊ⊕ⴊ⃿߆ࠄRNAࠍ⺞⵾ߒ
ߡ㧔࠴ࡦࡄࡦࠫ࡯ߪ㔤㐳⎇㘺⢒୘૕㧕‫ޔ‬ォ౮㐿ᆎὐߩ႐ᚲߣ૶↪㗫ᐲࠍ✂⟜⊛ߦ⸃ᨆߔࠆߚ߼ߩ࡜ࠗࡉ࡜࡝࡯ࠍ૞
ᚑߒߚ㧔⃻࿷‫ޔ‬ᄢⷙᮨࠪ࡯ࠤࡦࠨ࡯ߦߡ⸃ᨆਛ㧕‫ޕ‬MBD-seqߩ⸃ᨆ߆ࠄ‫ޔ‬CTCFߩ⚿วㇱ૏ߩᄌൻߦࠃࠆࡔ࠴࡞ൻᄌ
ൻ߇᥉ㆉ⊛ߥ߽ߩߢ޽ࠆߎߣࠍ⏕⹺ߒߡ޿ࠆ‫⚿ߩࠄࠇߎޕ‬ᨐߪ‫ޔ‬ㅴൻߩㆊ⒟ߢࠫࠚࡀ࠹ࠖ࠶ࠢߥᄌൻ߆ࠄ⴫⃻ဳߩ
ᄌൻ࡮ᄙ᭽ᕈ߇↢ߓࠆ㓙ߦࠛࡇࠫࠚࡀ࠹ࠖ࠶ࠢߥࡔࠞ࠾࠭ࡓ߇㊀ⷐߥᓎഀࠍṶߓߡ޿ࠆߎߣࠍ␜ߒߡ޿ࠆ‫ޕ‬
㧨ቇળ⊒⴫㧪
⑔↰ᷧ㧘߶߆ ‫ޟ‬㔤㐳㘃㑆ߩDNAࡔ࠴࡞ൻᄙဳߩ⸃ᨆ‫╙ ޠ‬83࿁ᣣᧄㆮવቇળ㧔2011/09, ੩ㇺ㧕
B-24 㘃ੱ₎ߩ㗡⬄ᐩࠍ⽾ㅢߔࠆ␹⚻ⴊ▤ሹߦ㑐ߔࠆ⥃ᐥၮ␆කቇ⊛࡮Ყセ⸃೬ቇ⊛⎇ⓥ
Ỉ㊁໪৻㧔␹ᄹᎹᱤ⑼ᄢ࡮ੱ૕᭴ㅧቇ㧕 ᚲౝኻᔕ⠪㧦ự↰ⓛ
੹࿁ߪ Canalis caroticus ߣ‫⽾ࠍߎߘޔ‬ㅢߔࠆ᭴ㅧ‛‫ޔ‬෸߮ ICA ߣ Foramen lacerum ߣߩ㑐ଥߦߟ޿ߡ⎇ⓥࠍⴕߞߚ‫ޕ‬
Canalis caroticus ࠍ⽾ㅢߔࠆᦨᄢߩ᭴ㅧ‛ߪ൩⺰ Arteria carotis interna (ICA) ߢ޽ࠆ߇‫ࠍࠇߘޔ‬ขࠅᏎߊ Sinus
cavernosus ߽㊀ⷐߢ޽ࠆ‫࠻ࡅޕ‬෸߮㘃ੱ₎ߢߪ Canalis caroticus ߪᒝߊዮᦛߔࠆ‫ⴊߪࠇߎޕ‬ᵹߩᷫ࿶ߣ‫ޔ‬A.ophthalmica,
A.c.anterior, A.c.media ╬߳ߩⴊᵹ㈩ಽߦᷓߊ᜔ࠊߞߡ޿ࠆߣ⠨߃ࠄࠇࠆ‫ ߛߚޕ‬Artiodactyla ╬ߦ⷗ࠄࠇࠆ Rete mirabile
ߣᲧセᬌ⸛ߔࠆߣ‫ޔ‬ICA ౝߩⴊᵹ߇ Hagen- Poiseuille’s equation ࠍᦨఝవߣߒߚ⸳⸘ߢ޽ࠆߣߪᔅߕߒ߽⸒߃ߥ޿‫ޕ‬
Perissodactyla ߦ߅ߌࠆ Canalis caroticus ߣઁߩ Foramina ߣߩว૕ߪ‫ޔ‬୘‫ ߩޘ‬Foramina ߿ Canales ࠍ⚦ಽൻߔࠆ௑ะ
ߩᒝ޿ Hominoidea ߣኻぞ⊛ߢ޽ࠅ‫ߦᦝޔ‬Ყセᬌ⸛ߩᔅⷐᕈ߇᦭ࠆ‫ޕ‬ᓥ᧪‫ޔ‬Canalis caroticus ߣ Foramen lacerum ߣߩ
㑐ㅪᕈߦ㑐ߔࠆ⎇ⓥߪ߶ߣࠎߤሽ࿷ߒߥ߆ߞߚ‫࠻ࡅޔߒ߆ߒޕ‬෸߮ Pongo ߦ߅ߌࠆ Foramen lacerum ߩ⊒㆐ߪ‫ޔ‬ICA
߇ Canalis caroticus ࠍ⽾ㅢߒߚᓟߦ‫ޔ‬਄ᣇߦ┙ߜ਄߇ࠆㇱಽߢߩᐥ㕙ࠍ‫⎬ޔ‬㛽ߢߪήߊ‫ޔ‬エ㛽ߦߔࠆߣ޿߁ὐߢ‫ޔ‬
↹ᦼ⊛ߥ᭴ㅧᄌൻߢ޽ࠆߎߣࠍᒝ⺞ߒߚ޿‫ޕ‬Canalis caroticus ౝ‫ޔ‬෸߮ߘࠇߦᒁ߈⛯ߊ⎬⤑ਅߦ߅޿ߡ‫ޔ‬ICA ࠍขࠅ
Ꮞߊ Sinus cavernosus ߦᣈߌࠆㅒⴕᕈߩ㕒⣂ⴊᵹߩሽ࿷߇‫ޔ࠻ࡅޔ‬ᰴ޿ߢ㘃ੱ₎ߦ߅޿ߡ㗼⪺ߦᄢ߈޿ߎߣࠍᜰ៰
ߒߚ޿‫ޕ‬
㧨⊒⴫᭎ⷐ㧪
Sawano K, Neurovascular tunnels in the midcranial base and their relative structure in the evolution of the cerebral blood
supply. (1): Kanagawa Dental College, (2): Edogawa Hospital, (3): Dept. Radiology, Yokohama City Univ. (Yokohama), (4):
Dept. Neurosurgery, Yokohama City Univ. Sch. Med.Anthropological Science, Volume 119, Number 3, p. 314, 2011.
B-25 ູ੃㘃ߦ߅ߌࠆ⣖ᒻᘒߩᐞ૗ቇ⊛ಽᨆ
ᴡㇱჽ৻㇢㧔᧲੩ᄢ࡮ℂ㧕 ᚲౝኻᔕ⠪㧦⷏᧛೰
㔤㐳㘃ߦ߅޿ߡ‫ޔ‬㗡⬄ၮᐩⷺߣ⣖ൻᜰᢙߣߩ㑆ߦ⋧㑐߇޽ࠆߎߣ߇⍮ࠄࠇߡ޿ࠆ߇‫੃ູߩઁޔ‬㘃ߢߩ⎇ⓥߪᄙߊ
ߥ޿‫ߩߘޕ‬ὑ‫ߩߎޔ‬㑐ଥߪູ੃㘃ో⥸⊛ߦ⷗ࠄࠇࠆ߽ߩߢ޽ࠆߩ߆᣿⏕ߦߪࠊ߆ߞߡ޿ߥ޿‫ޕ‬㔤㐳㘃એᄖߩູ੃㘃
ߦ߅ߌࠆ㗡⬄ၮᐩⷺߣ⣖ൻߣߩ㑐ଥࠍ᣿ࠄ߆ߦߔࠆߎߣߪ‫ޔ‬⣖ࠍ⊒㆐ߐߖࠆ߁߃ߢߩ㗡⬄ᒻᚑߩࡔࠞ࠾࠭ࡓߩℂ⸃
ߦ㕖Ᏹߦ᦭⋉ߢ޽ࠅ‫ޔ‬㔤㐳㘃ߢ⷗ࠄࠇࠆ⣖ൻߩᦝߥࠆℂ⸃ߦ❬߇ࠆ‫⺧↳ޕ‬⠪ߪ․ߦ㔤㐳㘃ߢ⷗ࠄࠇࠆ㗡⬄ၮᐩߣ⣖
㧙㧙
ൻᜰᢙߩ㑐ଥᕈ߇‫ޔ‬㔤㐳㘃એᄖߩູ੃㘃ߢ߽ห᭽ߦ⷗ࠄࠇࠆ߆ᬌ⸽ࠍⴕߞߚ‫ޕ‬
㔤㐳㘃ࠍ฽߻ູ੃㘃㗡㛽ߩ CT ࠬࠠࡖࡦࠍⴕ޿‫ޔ‬⣖ࠛࡦ࠼ࠠࡖࠬ࠻ࠍ૞⵾ߒߘߩ૕Ⓧࠍ⸘᷹ߒ⣖ൻᜰᢙࠍ▚಴ߒ
ߚ‫ ߚ߹ޕ‬CT ࠺࡯࠲߆ࠄ㗡⬄ၮᐩⷺߩ⸘᷹߽ⴕ޿‫ޔ‬⣖ൻᜰᢙߣߩ⋧㑐ࠍ⺞ߴߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬㔤㐳㘃એᄖߩູ੃㘃ߦ
߅޿ߡ‫ޔ‬ᒙ޿ߥ߇ࠄ߽㗡⬄ၮᐩⷺߣ⣖ൻᜰᢙߦ⋧㑐߇⷗ࠄࠇߚ‫ޕ‬
ߎߩߎߣ߆ࠄ⣖ൻߦࠃࠆ㗡⬄ၮᐩⷺߩᄌൻߪ‫ޔ‬㔤㐳㘃ߛߌߢ⷗ࠄࠇࠆ߽ߩߢߪߥߊ‫੃ູߩઁߩߘޔ‬㘃ߦ߅޿ߡ߽
ห᭽ߢ޽ࠆߎߣ߇ࠊ߆ߞߚ‫ޕ‬
B-26 ࠾ࡎࡦࠩ࡞ߩࠕࠞࡦࡏ࠙ߣߘߩᲣⷫ㑆ߢ↢ߓࠆ㘩ᕈߩ㆑޿ߦ㑐ߔࠆ‛ℂ⊛ⷐ࿃ߩᬌ⸛
⼱ญ᥍㚅㧔੩ㇺᄢ࡮㒮࡮ℂ㧕 ᚲౝኻᔕ⠪㧦ඨ⼱๋㇢
ᧄ⎇ⓥߢߪ࠾ࡎࡦࠩ࡞ߩ㘩‛ຠ⋡ߩ߆ߚߐߩ⸘᷹ࠍⴕ޿‫߇࠙ࡏࡦࠞࠕޔ‬ᲣⷫߣᲧセߒߤߩࠃ߁ߥ߆ߚߐࠍㆬᅢߔ
ࠆߩ߆ࠍ⺞ᩏߒߚ‫ޕ‬౻ቄߦ‫ޔ‬㣮ఽፉ⋵ደਭፉߣ㕍᫪⋵ਅർඨፉߦ߅޿ߡ‫ߩߘߪߚ߹࠙ࡏࡦࠞࠕޔ‬Უⷫࠍㅊ〔ߒ‫ޔ‬ណ
㘩ᤨߦ㘩‛ຠ⋡ߩណ㓸ࠍⴕ޿‫࡯࠲࡯ࡔࠝ࡟ޔ‬㧔ᩣᑼળ␠ࠨࡦ⑼ቇ‫ޔ‬MODEL:COMPAC-100Τ㧕ߦߡ 6 ᤨ㑆એౝߦ߆
ߚߐ⸘᷹ࠍⴕߞߚ‫੍ޕ‬஻⊛ߥ⸃ᨆߢߪ޽ࠆ߇‫ޔ‬Უሶߩ㘩‛ຠ⋡ߩ೑↪ߩ㆑޿ߣᦨᄢ⎕უ⩄㊀୯㧔ⵙᢿᤨߦ↢ߓߚ⩄
㊀ߩᦨᄢ୯ࠍᢿ㕙Ⓧߢഀߞߚ୯㧕ߣߩ㑐ଥߦߟ޿ߡᬌ⸛ߒߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬ਔ࿾ၞ౒ߦᲣⷫߣᲧセߒߡࠕࠞࡦࡏ࠙ߪ
ᦨᄢ⎕უ⩄㊀୯߇ૐ޿ຠ⋡ࠍㆬᅢߔࠆ௑ะߦ޽ߞߚ‫ޔ߫߃଀ޕ‬ਅർඨፉߢߪ‫߇࡞ࠨޔߪ࠙ࡏࡦࠞࠕޔ‬೑↪ߔࠆ᮸⊹
ߩਛߢ߽ᦨᄢ⎕უ⩄㊀୯߇ૐ޿࠷࡝ࡃ࠽߿ࠢ࠭ߩ᮸⊹ࠍ೑↪ߔࠆ௑ะߦ޽ߞߚ‫ޕ‬ደਭፉߢߪ‫ޔ‬Უⷫ߇࿾㕙ߢࡕࠢ࠲
࠴ࡃ࠽ߩ⒳ሶࠍណ㘩ߒߡ޿ࠆ㓙ߦ‫ߪ࠙ࡏࡦࠞࠕޔ‬᮸਄ߢ‫ޔ‬⒳ሶࠃࠅᦨᄢ⎕უ⩄㊀୯߇ૐ޿ห⒳ߩᧂᾫ⒳ሶࠍណ㘩ߔ
ࠆ႐㕙߇ߚ߮ߚ߮ⷰኤߐࠇߚ‫⚿ߩߎޕ‬ᨐ߆ࠄ‫ޔ‬ᲣⷫߣᲧセߒߡࠕࠞࡦࡏ࠙ߪ๰ྨ⢻ജ߇ૐ޿ߚ߼‫޿߆ࠄࠊ߿ޔ‬ຠ⋡
ࠍㆬᅢߒߚน⢻ᕈ߇␜ໂߐࠇߚ‫੹ޕ‬ᓟ‫ߩߐߚ߆ޔ‬ಽᨆࠍㅴ߼ࠆߣ౒ߦ‫ޔ‬㘩‛ຠ⋡ߩᄢ߈ߐ߿ߘߩᓧࠄࠇࠆ㜞ߐ‫ޔ‬ᠲ
૞ᢙߥߤઁߩ‛ℂ⊛ᕈ⾰߽⠨ᘦߦ޿ࠇ‫ޔ‬Უሶߩ㘩‛೑↪ߩ㆑޿ࠍᬌ⸛ߒߡ޿߈ߚ޿‫ޕ‬
B-27 ㊁↢࠾ࡎࡦࠩ࡞ߩࡢࠞࡕࡁࠝࠬߩ಴⥄⟲㔌⣕೨ᓟߩ↢ᵴผߦ㑐ߔࠆ㐳ᦼㅊ〔⺞ᩏ
ፉ↰዁༑㧔Ꮲ੩⑼ቇᄢ࡮ࠕ࠾ࡑ࡞ࠨࠗࠛࡦࠬ㧕 ᚲౝኻᔕ⠪㧦ඨ⼱๋㇢
ችၔ⋵㊄⪇ጊߩ࠾ࡎࡦࠩ࡞ߩࠝࠬߦߪ‫ࠇ⟲ޔ‬ౝࠝࠬ‫ࠆߥ⇣߁޿ߣࠬࠝ࡝࠻ࡅޔࡊ࡯࡞ࠣࠬࠝޔ‬ሽ࿷᭽ᑼ߇޽ࠆ‫ޕ‬
ࠝࠬߪᚑ㐳ߦ઻޿಴⥄⟲ࠍ㔌⣕ߒ‫ߡ߇߿ޔ‬೎ߩ⟲ࠇߦ⒖☋ߔࠆ߇‫ߦࠇ⟲ޔ‬ㅊ㓐ߔࠆࠝࠬߣ⟲ࠇࠝࠬߣߩ㑆ߦߪẜ࿷
⊛ߥࠦࡦࡈ࡝ࠢ࠻߇ሽ࿷ߒ‫ޔ‬ㅊ㓐ࠝࠬ߇ᱫߦ⥋ࠆߎߣ߽᦭ࠅᓧࠆߚ߼‫ޔ‬㔌⣕࡮⒖☋ߪ‫ޔ‬ᒰ⹥୘૕ߦߣߞߡߪ࡝ࠬࠢ
ߩ㜞޿ⴕേߩߪߕߛ‫ߩࠬࠝߩߎޕ‬ሽ࿷᭽ᑼߩᄌൻߩ⥋ㄭⷐ࿃߆ࠄ᣿ࠄ߆ߦߔࠆߚ߼ߦ‫ޔ‬㊄⪇ጊߩ࠾ࡎࡦࠩ࡞ A ⟲
಴りߩ 2011 ᐕᤨὐߢ 7-8 ᱦߩࡢࠞࡕࡁࠝࠬ 3 㗡㧔ࠕࠪࡕ࡮ࠗࠞࡠࠬ࡮ࡈࡒࡗ㧕ࠍ⺞ᩏኻ⽎ߣߒ‫ⴕޔ‬േ࡮␠ળ㑐ଥ
ߩ࠺࡯࠲ࠍ 2009 ᐕ߆ࠄ 2011 ᐕ㧔ਃᐕ㑆㧕ߦࠊߚࠅᢿ⛯⊛ߦ෼㓸ߒߚ‫ޕ‬
ࠗࠞࡠࠬ㧔8 ᱦ㧕ߪ 09 ᐕᄐએ೨ߦߪ‫ޔ‬ፉߩർ⷏ㇱߩ A ⟲ౝߣߘߩ๟ㄝߩࠝࠬࠣ࡞࡯ࡊߢ⏕⹺ߐࠇߡ޿ߚ߇‫ޔ‬09
ᐕ⑺એ㒠 A ⟲ߩ᧲ߩ C2 ⟲ߩ๟ㄝߩࠝࠬࠣ࡞࡯ࡊߢ⏕⹺ߐࠇߡ޿ࠆ‫ࡕࠪࠕޕ‬㧔8 ᱦ㧕ߪ 09 ᐕએ㒠‫ޔ‬A ⟲ߩධߩ B1
⟲ߩ๟ㄝߩࠝࠬࠣ࡞࡯ࡊߢ⏕⹺ߐࠇ⛯ߌߡ޿ࠆ‫ࡗࡒࡈޕ‬㧔7 ᱦ㧕ߪ 09 ᐕᄐએ೨ߦߪ A ⟲ౝߢ⏕⹺ߐࠇߡ޿ߚ߇‫ޔ‬
09 ᐕ⑺એ㒠৻ᤨ⊛ߦ B1 ⟲ߩ๟ㄝߩࠝࠬࠣ࡞࡯ࡊߦ಴⃻ߔࠆࠃ߁ߦߥࠅ‫ޔ‬10 ᐕᄐએ㒠ቯ⌕ߒߡ޿ࠆߣផቯߐࠇߚ‫ޕ‬
ࠕࠪࡕߪ‫ޔ‬B1 ㅊ㓐ࠝࠬࠣ࡞࡯ࡊౝߢ‫ࠆ޽ߩ✼ⴊޔ‬ᐕ㐳୘૕ߣ␠ળ㑐ଥ߇ᒝ޿‫ޕ‬
ࠦ࠼ࡕᦼ߆ࠄ⟲ࠇࠍ಴ߡࠁߊ㗫ᐲࠍ਄ߍߡࠁ߈‫ߦࡊ࡯࡞ࠣࠬࠝߡ߇߿ޔ‬ቯ⌕ߔࠆߎߣߢ‫ޔ‬಴⥄⟲߆ࠄߩ㔌⣕ࠍቢ
ᚑߔࠆ‫߁޿ߣޔ‬࿑ᑼߪ㊄⪇ጊߦ߅޿ߡ߽ⷰኤߐࠇߚ‫৻ޕ‬ᣇߤߎߦቯ⌕ߔࠆ߆ߪࠝࠬߚߜߣߩⴊ✼‫ޔ‬ㆊ෰ߩ㑐ଥ‫ࠗޔ‬
ࡦ࠲࡜࡚ࠢࠪࡦߩⓍߺ㊀ߨ‫ߡߞࠃߦߤߥޔ‬ᓇ㗀ࠍฃߌࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬
B-28 දജ⺖㗴ߦ߅ߌࠆ⥄Ꮖ⹺⍮ߩታ㛎⊛ಽᨆ
⨲ጊᄥ৻㧔Ꮲ੩ᄢ࡮ᢥ࡮ᔃℂ㧕 ᚲౝኻᔕ⠪㧦⣁↰⌀ᷡ
േ‛ߦ㏜ࠍឭ␜ߒ‫⥄ߩߘޔ‬Ꮖߩ෻኿௝ࠍ⥄Ꮖߣ⹺⍮ߔࠆ߆ߤ߁߆ࠍ⺞ߴࠆ⎇ⓥߪ⥄Ꮖ㏜௝⹺⍮ߣ๭߫ࠇ‫⃻ޔ‬࿷߹
ߢߦᄙߊߩേ‛⒳ࠍኻ⽎ߦᬌ⸛ߐࠇߡ޿ࠆ‫ޕ‬ᄙߊߩవⴕ⎇ⓥߢߪ‫ޔ‬⒳ࠍ㒢ቯߒߥ޿ߢⷞⷡ⊛ߦઁ୘૕ࠍ⼂೎ߢ߈ࠆ
ߎߣ߇ಽ߆ߞߡ޿ࠆ৻ᣇ‫⥄ޔ‬Ꮖ㏜௝ࠍ⥄Ꮖߩ෻኿‛ߣ⹺⼂ߔࠆߎߣߪ㔍ߒ޿ߣႎ๔ߒߡ޿ࠆ‫౒ޔߢߎߘޕ‬ห૞ᬺࠍ
ㅢߓߚઁ⠪ߩⴕേߩࡕ࠾࠲࡯߇‫⥄⊛ⷡⷞޔ‬Ꮖ⹺⍮ߩᚑ┙ࠍଦㅴߔࠆߣ઒ቯߒ‫⥄ޔ‬Ꮖ⹺⍮ߩᚑ┙ⷐ࿃ߦߟ޿ߡ‫ࡕࠦޔ‬
ࡦࡑ࡯ࡕ࠮࠶࠻ 4 ୘૕ࠍኻ⽎ߦታ㛎ࠍ߅ߎߥߞߚ‫ޕ‬
౒ห૞ᬺߦߟ޿ߡ‫ޔ‬ೋ߼ߦ୘૕ߏߣߦ‫ߚ޿ߟߩࡕࡅޔ‬บゞࠍᚻ೨ߦᒁ߈ነߖࠆߣ‫ޔ‬บゞߦਸ਼ߞߚࠛࠨࠍᚻߦ౉ࠇ
ࠆ⸠✵ࠍ߅ߎߥߞߚ‫ߩߡోޕ‬୘૕ߪ‫ޔ‬10 ⹜ⴕ⒟ᐲߢߔߋߦࡅࡕࠍߚߋࠅነߖߡ‫ࠍࠨࠛޔ‬ขࠆߎߣ߇ߢ߈ߚ‫ߎߘޕ‬
ߢ‫࠻ࠗࡔࠫ࡯ࠤޔ‬ห჻ࠍࡍࠕߣߒߡ‫ޔ‬2 ୘૕߇หᤨߦࡅࡕࠍᒁ߆ߥ޿ߣบゞࠍᒁ߈ነߖࠆߎߣ߇ߢ߈ߥ޿ࠃ߁ߥද
ജ⺖㗴ߩ⸠✵ࠍታᣉߒߚ‫ޕ‬1 ୘૕ߛߌߩ⸠✵ߢߪߔߋߦࡅࡕࠍᒁߊߎߣ߇ߢ߈ߚ߇‫ޔ‬2 ୘૕߇หߓ࠲ࠗࡒࡦࠣߢᒁ
ߊߎߣߪ㔍ߒߊ‫ޔ‬ታ㛎ⵝ⟎ߩᡷༀ߇ᔅⷐߣߥߞߚ‫ޕ‬
߹ߚ‫߇࠻࠶࠮ࡕ࡯ࡑޔ‬㏜௝ߦኻߒߡ‫ߥ߁ࠃߩߤޔ‬෻ᔕࠍ␜ߔ߆‫ⴕޔ‬േⷰኤࠍⴕߞߚߣߎࠈ‫ޔ‬ታ㓙ߩࠤ࡯ࠫࡔࠗ࠻‫ޔ‬
㕖ࠤ࡯ࠫࡔࠗ࠻ߣߪ⇣ߥࠆ෻ᔕ߇⹺߼ࠄࠇߚ‫ޕ‬
B-29 ㆮવሶ⸃ᨆߦࠃࠆਃ㊀⋵ౝߩ࠾ࡎࡦࠩ࡞ߩ୘૕⟲⺞ᩏ
౐ᵄ⟜⡡㧘㋈ᧁ⟵ਭ㧔NPO ᴺੱࠨ࡞ߤߎࡀ࠶࠻㧕 ᚲౝኻᔕ㓶⠪㧦Ꮉᧄ⧐
ਃ㊀⋵ౝߩ࠾ࡎࡦࠩ࡞ߦߟ޿ߡ‫▤⼔଻ޔ‬ℂࠍᬌ⸛ߔࠆߚ߼‫⃻ޔ‬ሽߔࠆ⟲ࠇߩㆮવ⊛᭴ㅧࠍᛠីߔࠆߎߣ‫ޔ‬๺᱌ጊ
⋵߆ࠄߩ࠲ࠗࡢࡦࠩ࡞ㆮવሶߩ᜛ᢔ⁁ᴫߩࡕ࠾࠲࡝ࡦࠣࠍ⎇ⓥ⋡⊛ߣߒ‫ᧄޔ‬ᐕᐲߪࠝࠬ 75 ୘૕ߦߟ޿ߡ Y-STR ᬌ
ᩏ‫ ࠬࡔޔ‬8 ୘૕ߦߟ޿ߡ D-loop ╙㧝นᄌၞ‫ ╙ޔ‬2 นᄌၞߩႮၮ㈩೉ߩಽᨆࠍⴕߞߚ‫ޕ‬
ࠝࠬߩ Y ᨴ⦡૕ߪ 13 ࠲ࠗࡊߦಽ㘃ߐࠇߚ‫ޕ‬ㆊ෰ߦᬌᩏߐࠇߚฦ⋵ߩ⚿ᨐߣᲧセߒߡᄙ᭽ߥ࠲ࠗࡊ߇⏕⹺ߐࠇߚ‫ޕ‬
ฦ࠲ࠗࡊߣ߽࿾ၞዪ࿷ᕈߪ⹺߼ࠄࠇߕ‫ޔ‬ᄙ᭽ߥ Y ㆮવሶ߇ᐢ▸࿐ߦಽᏓߒߡ޿ࠆߣផኤߐࠇࠆ‫↱࡞ࠩࡦࡢࠗ࠲ޕ‬
᧪ߣߺࠄࠇࠆ࠲ࠗࡊߪ⏕⹺ߐࠇߥ߆ߞߚ‫ޕ‬
㧙㧙
ࡔࠬߦߟ޿ߡߪ‫ޔ‬D-loop ╙ 1 นᄌၞߦߟ޿ߡ 7 ࠲ࠗࡊ߇⏕⹺ߐࠇ‫ޔ‬㋈㣮Ꮢߩ⟲ࠇߩ࠲ࠗࡊߪㆊ෰ߩ⎇ⓥ⚿ᨐߩṑ
⾐⋵ౝߩ⟲ࠇߣหߓ࠲ࠗࡊߢ޽ߞߚ‫ ߩઁޕ‬6 ࠲ࠗࡊߪ‫ઁޔ‬࿾ၞߢ⏕⹺ߐࠇߡ޿ߥ޿࠲ࠗࡊߢ޽ߞߚ‫ޕ‬
੹ᓟࠨࡦࡊ࡞ᢙࠍჇ߿ߒߡ޿ߊߎߣߢ‫ޔ‬ਃ㊀⋵ౝߩ࠾ࡎࡦࠩ࡞ߩ⟲ࠇߩᚑ┙߿᜛ᢔ⁁ᴫࠍ⚦߆ߊ᣿ࠄ߆ߦߒߡ޿
ߊ੍ቯߢ޽ࠆ‫ޕ‬
B-30 ⊹⤏ߩㅴൻ↢ℂቇ
㘖↰⪲ሶ㧘Ꮉ᎑ᓀᄐ㧘ੇߎࠁࠆ㧘ᴡ㊁⟤ᕺሶ㧘᩵᦭ടሶ㧔✚⎇ᄢ࡮వዉ⎇㧕 ᚲౝኻᔕ⠪㧦ㇹᐽᐢ
ࡅ࠻ߣ㔤㐳㘃ߩᒻᘒ⊛ߥ㆑޿ߩ৻ߟߣߒߡ‫ߩ⤏⊹ޔ‬᭴ㅧ߇޽ࠆ‫ޕ‬૕Ძߩ᦭ήࠍ฽߼ߡ‫ޔ‬᳨⣼߿‫⊹ޔ‬ਅ⢽⢌ߩ㊂‫ޔ‬
᷷ᐲᗵⷡฃኈ૕‫ߥ߹ߑ߹ߐޔߤߥ♽∉఺ޔ‬ᒻ⾰ߦ㑐ࠊࠆಽሶߩಽᏓ߇ࡅ࠻ߣઁߩ㔤㐳㘃ߩ㑆ߢߪ⇣ߥࠆߎߣ߇੍ᗐ
ߐࠇࠆ‫ޔߢߎߘޕ‬Ԙ⊹⤏ߢߩߎࠇࠄߩᒻ⾰ߦ㑐ࠊࠆㆮવሶߩ⊒⃻㊂ࠍࡅ࠻ߣ㔤㐳㘃ߢᲧセߔࠆ‫ޕ‬ԙ⊒⃻ߩ㆑޿߇ࠊ
߆ߞߡ޿ࠆㆮવሶߦߟ޿ߡߪ‫ߩߘޔ‬๟ㄝߩࠥࡁࡓߩႮၮ㈩೉߽᳿ቯߒ‫ߩߣ࠻ࡅޔ‬Ყセࠍⴕ޿‫⃻⊒ޔ‬Ꮕߦነਈߔࠆㆮ
વ⊛ⷐ࿃ߩ․ቯࠍ⹜ߺࠆ‫ޕ‬Ԛࠥࡁࡓߩࡔ࠴࡞ൻߥߤߩࠛࡇࠫࠚࡀ࠹ࠖࠢࠬߩലᨐࠍ᣿ࠄ߆ߦߔࠆߚ߼ߩ⸃ᨆ߽ⴕ߁‫ޕ‬
એ਄ߩ 3 ߟߩ⋡⊛ࠍᜬߞߡ⎇ⓥࠍㆀⴕߒߚ‫ޕ‬
ᧄ⎇ⓥߢߪ‫᧚ޔ‬ᢱߣߒߡ㔤㐳㘃ߩ⊹⤏⚵❱߆ࠄ mRNA ߅ࠃ߮ DNA ࠍන㔌ߔࠆ‫ ߩߎޕ‬mRNA ߣࡅ࠻ߩ⊹⤏↱᧪
ߩ mRNA ߩ⊒⃻㊂ࠍᲧセߔࠆ‫ޕ‬⒳ౝߩ୘૕㑆ߩᄌ⇣ߣߩ㆑޿ࠍ᣿⏕ߦߔࠆߚ߼ߦ‫ޔ‬㔤㐳㘃ߢߪ৻ߟߩ⒳߆ࠄⶄᢙ
୘૕ߩࠨࡦࡊ࡞ߦߟ޿ߡ⺞ߴࠆߎߣࠍ⋡ᮡߣߔࠆ‫ޔ࡜࡝ࠧޔ࡯ࠫࡦࡄࡦ࠴ޕ‬ᣥ਎⇇ࠩ࡞‫ޔ‬ᣂ਎⇇ࠩ࡞↱᧪ߩ⊹⤏ࠨ
ࡦࡊ࡞㧔ᢙᐔᣇ࠮ࡦ࠴ࡔ࡯࠻࡞྾ᣇ⒟ᐲ㧕ࠍ೑↪ߔࠆ੍ቯߢ޽ߞߚ‫ޕ‬
ߎࠇ߹ߢߦ‫౒ޔ‬ห⎇ⓥߢឭଏ޿ߚߛ޿ߚ‫ⶄ࡞ࠩࠥࠞࠕޔ‬ᢙ୘૕ߩ⊹⤏ߦട߃ߡ‫ᧄޔ‬ᐕᐲߪ‫ࡄࡦ࠴ޔ࡜࡝ࠧࠪ࠾ޔ‬
ࡦࠫ࡯‫ࠍ࡞ࡊࡦࠨ⤏⊹ߩ࡞ࠩࠟ࠽࠹ࡠࠢࡈޔ‬ଏਈ޿ߚߛ޿ߚ‫ ߡ޿ߟߦ࡞ࡊࡦࠨߩࠄࠇߎޕ‬RNA ࠍ᛽಴ߒߚ‫⃻ޕ‬࿷
ߪߎࠇࠄߩ RNA ࠍ↪޿ߚ‫ޔ‬NGS (next generation sequencing) ߦࠃࠆ⊒⃻⸃ᨆࠍ⸘↹ߒߡ߅ࠅ‫ࡄࡦ࠴ޔߩߢ߹ࠇߎޔ‬
ࡦࠫ࡯෸߮ࠕࠞࠥࠩ࡞ߩࡑࠗࠢࡠࠕ࡟ࠗߢߩ⚿ᨐߣߩᲧセࠍⴕ޿ߚ޿‫ޕ‬
B-31 ₎ኂ⟲ߦ߅ߌࠆ࿯࿾೑↪․ᕈߣㄘ࿾೑↪ߦᓇ㗀ࠍਈ߃ࠆⷐ࿃ߩᬌ⸛
⼱ᄢテ㧔ጊญᄢ࡮ㄘ㧕 ᚲౝኻᔕ⠪㧦ඨ⼱๋㇢
ᧄ⎇ⓥߪ‫ޔ‬ㄘ૞‛ⵍኂࠍᒁ߈⿠ߎߒߡ޿ࠆ࠾ࡎࡦࠩ࡞ 2 ⟲ࠍኻ⽎ߦ‫ޔ‬ណ㘩႐ᚲߩㆬᛯߦ㑐ࠊࠆⶄᢙߩ㗄⋡ࠍ⸳ቯ
ߒߡߎࠇࠍቯ㊂⊛ߦ⹏ଔߒ‫ߩ࡞ࠨޔ‬ㄘ࿾߳ߩଐሽᐲߣߩ㑐ㅪߦߟ޿ߡ᣿ࠄ߆ߦߔࠆߎߣࠍ⋡⊛ߣߒߚ‫ޕ‬ጊญ⋵ߦ↢
ᕷߔࠆ A ⟲㧔ጊญᏒ A ࿾඙㧕ߣ B ⟲㧔ቝㇱᏒ B ࿾඙㧕ߦߟ޿ߡ‫ ࠇߙࠇߘޔ‬2 㗡ߩ୘૕ߦ⊒ାᯏࠍⵝ⌕ߒߡ‫ࠫ࡜ޔ‬
ࠝ࠹࡟ࡔ࠻࡝࡯ᴺߦࠃࠅⴕേၞߣ࿯࿾೑↪㗫ᐲࠍ⺞ߴߚ‫ⴕߩࠇ⟲ߩࠇߙࠇߘޔߚ߹ޕ‬േၞߦ฽߹ࠇࠆ㓸⪭ߦ߅޿ߡ‫ޔ‬
ㄘ࿾ߩ⒳㘃ߣ㕙Ⓧ‫ޔ‬ㄘ࿾ߣᨋ✼ߣߩ〒㔌‫ޔ‬ᩑ߅ࠃ߮ᩙߩᨐ᮸ᧁߩᧄᢙ‫ⵍޔ‬ኂኻ╷ߩ⒳㘃ߣߘߩ᦭ήߦߟ޿ߡ⺞ߴߚ‫ޕ‬
ᦨᄖㇳᴺ㧔100%㧕ߦࠃࠆⴕേၞ㕙Ⓧߪ A ߅ࠃ߮ B ⟲ߦ߅޿ߡߘࠇߙࠇ 27.3km2 ߣ 9.5km2 ߢ޽ߞߚ‫ߪ࠭ࠗࠨࠇ⟲ޕ‬
ᱜ⏕ߥ㗡ᢙߪਇ᣿ߢ޽ࠆ߇‫ ߽ࠇߕ޿ޔ‬70ޯ100 㗡ߢ޽ߞߚ‫ޕ‬࿯࿾೑↪㗫ᐲߦߟ޿ߡ‫ޔ‬A ⟲ߢߪቄ▵ߦࠃࠆ೑↪႐ᚲ
߅ࠃ߮㗫ᐲߪᄌൻߒߚ߇‫ޔ‬B ⟲ߢߪ㧝ᐕࠍㅢߓߡหߓ႐ᚲࠍ➅ࠅ㄰ߒ೑↪ߔࠆ௑ะ߇ߺߣ߼ࠄࠇߚ‫ޕ‬ㄘ࿾ߩ࠲ࠗࡊ
೎ഀวߣ‫ޔ‬ㄘ࿾ߣᨋ✼ߣߩ〒㔌ߢߪ‫ޔ‬A ࿾඙ߣ B ࿾඙ߢᄢ߈ߥ㆑޿ߪߥ߆ߞߚ߇‫ⵍޔ‬ኂኻ╷ߦߟ޿ߡߪ A ࿾඙ߩ
ᣇ߇ᣇᴺߩᢙ߿ⴕ߁㗫ᐲ߇ᄙ߆ߞߚ‫ޕ‬B ࿾඙ߢߪ․ቯߩㄘ࿾ߩ㓸ਛ⊛ߥⵍኂ߇ᄙߊ‫ ߁߸ߞ޿ޔ‬A ࿾඙ߢߪ‫ⵍޔ‬ኂ߇
ಽᢔߒߡ߅ࠅ‫ߩࠢ࠶࠼࡯ࠠࡦࡕߦߊߣޔ‬ᵴേ႐ᚲߦ߅޿ߡߪⵍኂ߇ዋߥ߆ߞߚ‫ޔߚ߹ޕ‬ਔ࿾඙ߦ߅޿ߡ‫ޔ‬9 ᦬߆ࠄ
11 ᦬ߦߪᨐ᮸ᧁߩᧄᢙ߇ᄙ޿႐ᚲ߶ߤ೑↪㗫ᐲ߇㜞߆ߞߚ‫ޕ‬એ਄ࠃࠅ‫ޔ‬᫪ᨋⅣႺ߇߶߷หߓߢ޽ࠆㄭ㓞ߩ↢ᕷ࿾ࠍ
߽ߟ 2 ⟲ߦ߅޿ߡ‫ޔ‬࿯࿾೑↪ߩ㘃ૃᕈߣ⋧㆑↢߇᣿ࠄ߆ߣߥࠅ‫ޔ‬ㄘ࿾߳ߩଐሽߩᒝߐ߇⇣ߥࠆߎߣ߇ಽ߆ߞߚ‫ޕ‬
B-32 㔤㐳㘃ߩ⤖ౝᏱ࿷⚦⩶ฌߦ߅ߌࠆ੃㉄⩶ߩᨐߚߔᓎഀ
㊁ญ๺ᶈ㧔ᾢᧄᄢ࡮㒮࡮↢๮⑼ቇ㧕㧘ᶆ㊁ᔀ㧔ᾢᧄᄢ࡮↢๮⾗Ḯ㧕 ᚲౝኻᔕ⠪㧦ᐔ੗໪ਭ
ࡅ࠻ߩ⤖ౝߢߪ‫ᦨࠍ⩶㉄੃ޔ‬ఝ൓ߣߔࠆᏱ࿷⚦⩶ฌ߇ᒻᚑߐࠇ‫ޔ‬ᄖㇱ߆ࠄߩ∛ේ⩶ߩଚ౉࡮Ⴧᱺࠍ㒐㒰ߒߡ޿ࠆ
ߣ⠨߃ࠄࠇߡ޿ࠆ‫⤖࠻ࡅޔߢߎߘޕ‬ౝߢߩ੃㉄⩶ߩᓎഀࠍ⎇ⓥߔࠆߚ߼ߩࡕ࠺࡞േ‛ߣߒߡߩ㔤㐳㘃ߩ᦭↪ᕈࠍ᣿
ࠄ߆ߦߔࠆߚ߼‫࡞ࠩࡦࡎ࠾ޔ‬㧔㓽 15 㗡㧧3㨪21 ᱦ㧕ߩ⤖ౝ⚦⩶ฌߩᬌ⚝ࠍⴕߞߚ‫ⷙޕ‬ೣ⊛ߥ᦬⚻๟ᦼ߇ⷰኤߐࠇ
ߚ 11 㗡ߩ⤖ౝ⚦⩶ฌߩᚑ❣ࠍ⸃ᨆߒߚߣߎࠈ‫⤖ߩ࡞ࠩࡦࡎ࠾ޔ‬ౝ߆ࠄߪ 5 ⒳㘃ߩㅢᕈህ᳇ᕈ⩶㧔Enterobacteriaceae,
Strept ococci, Staphylococci, Corynebacteria, Lactobacilli 㧕 ෸ ߮ 4 ⒳ 㘃 ߩ ህ ᳇ ᕈ ⩶ 㧔 Bacteroidaceae, Veillonellae,
Gram-positive anaerobic cocci (GPAC), Gram-positive anaerobic rods 㧕 ߇ ಽ 㔌 ߐ ࠇ ߚ ‫ ߜ ߁ ߩ ߘ ޕ‬Streptococci,
Corynebacteria, Bacteroidaceae ෸߮ GPAC ߪ 80%એ਄ߩ୘૕߆ࠄಽ㔌ߐࠇ‫ߟ߆ޔ‬ಽ㔌⩶ᢙ߽㜞߆ߞߚߎߣ߆ࠄ‫ࠇߎޔ‬
ࠄߩ⩶⒳ߪ࠾ࡎࡦࠩ࡞ߩ⤖ౝߦ߅ߌࠆਥⷐߥ᭴ᚑ⩶⒳ߢ޽ࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬Lactobacilli ߩᬌ಴₸ߪ 56%ߣਛ⒟
ᐲߢ޽ߞߚ߇‫ޔ‬ಽ㔌⩶ᢙߪ 105.4 (CFU/vagina) ߣ࠴ࡦࡄࡦࠫ࡯ߩ႐วߣห᭽ߦᲧセ⊛㜞޿୯ࠍ␜ߒߡ޿ߚ‫ޔߚ߹ޕ‬
ಽ㔌ߐࠇߚ⤖ౝ✚⚦⩶ᢙࠍ᦬⚻๟ᦼ೎㧔ෆ⢩ᦼ‫ޔ‬㤛૕ᦼ෸߮᦬⚻ᦼ㧕ߦಽߌߡᲧセߒߚߣߎࠈ‫ࡦࠥࡠ࠻ࠬࠛޔ‬Ớᐲ
߇㜞ߊߥࠆෆ⢩ᦼߢᦨ߽㜞ߊߥࠆ௑ะࠍ␜ߒߚ‫ޕ‬ᓟᬌᩏ଀ᢙࠍჇ߿ߔߎߣߦࠃࠅ‫⚻᦬ޔ‬๟ᦼ߿ട㦂╬ߩⷐ࿃߇࠾ࡎ
ࡦࠩ࡞⤖ౝ⚦⩶ฌߦߤߩࠃ߁ߥᓇ㗀ࠍਈ߃ߡ޿ࠆ߆ࠍ᣿⏕ߦߒߚ޿‫ޕ‬
B-33 ࠾ࡎࡦࠩ࡞੃ఽߦ߅ߌࠆㆇേ್ᢿ ̆⛘ኻ್ᢿ߆⋧ኻ್ᢿ߆̆
ᷰㄝഃᄥ㧘⮮↰๺↢㧔੩ㇺᄢ࡮㒮࡮ᢥ㧕 ᚲౝኻᔕ⠪㧦෹᳗㓷Ꮖ
‛૕ߩㆇേࠍⷞⷡ⊛ߦ⹺⼂ߔࠆ㓙‫ޔ‬๟࿐ೝỗߩㆇേߣ㑐ㅪઃߌߡ್ᢿߔࠆ⋧ኻ್ᢿߣ‫ߩߘޔ‬ೝỗߘߩ߽ߩߩ‛ℂ
⊛ߥ૏⟎ߩᄌൻࠍ್ᢿߔࠆ⛘ኻ್ᢿ߇ሽ࿷ߔࠆ‫⎇ᧄޕ‬ⓥߪ‫ޔ‬න⚐࿑ᒻࠍ↪޿ߡ‫⋡߇ఽ੃࡞ࠩࡦࡎ࠾ޔ‬ᮡೝỗߩേ߈
ࠍ್ᢿߔࠆ㓙‫ޔ‬ᨒೝỗߩᓇ㗀ࠍฃߌࠆ㧔⋧ኻ್ᢿ㧕߆ฃߌߥ޿㧔⛘ኻ್ᢿ㧕߆ࠍᲧセ⹺⍮⊒㆐ߩⷰὐ߆ࠄ‫ޔ‬ᘠൻᴺ
ࠍ↪޿ߡಽᨆߒߚ‫⋡ޕ‬ᮡೝỗߢ޽ࠆචሼ࿑ᒻߣߘߩ๟࿐ߦ޽ࠆᱜᣇᒻᨒ߆ࠄᚑࠆ࠮࠶࠻ࠍೝỗߣߒߡ૶↪ߒ‫ޔ‬2 ࠮
㧙㧙
࠶࠻ࠍᏀฝኻ๒␜ߒߚ‫ޕ‬᡼㘺႐୘૕ߩ৻ᢧᬌᩏߩ㓙ߦ‫ޔ‬0 ᱦఽߩ࠾ࡎࡦࠩ࡞੃ఽࠍ⍴ᤨ㑆Უࠩ࡞߆ࠄಽ㔌ߒ࠲ࠝ࡞
ߢ଻ቯߒߚ߁߃ߢ‫ޔ‬೨㕙ߦ⸳⟎ߐࠇߚࡕ࠾࠲࡯ߦᤋߒ಴ߐࠇࠆೝỗࠍ๒␜ߒߚ‫ޕ‬ᨒೝỗߪ‫ޔ‬6࡮7 ࿁⋡ߩ๒␜ᤨߦ
ㆇേߒߚ‫ ߅ߥޕ‬6࡮7 ࿁⋡ߩߺ‫ޔ‬Ꮐฝߢೝỗߩㆇേᣇะߪ⇣ߥߞߚ㧔⋧ኻਇ৻⥌ VS ⛘ኻਇ৻⥌㧕
‫ߩࠄࠇߎޕ‬ೝỗ๒
␜ᤨߦ޿ߕࠇߩೝỗ࠮࠶࠻ࠍࠃࠅᵈⷞߔࠆ߆ࠍⷰኤߒߚ‫ⴕߩఽ੃ޕ‬േࠍࡆ࠺ࠝࠞࡔ࡜ࠍ↪޿ߡ⸥㍳ߒ‫ޔ‬ᵈⷞᤨ㑆ࠍ
᷹ቯߒᏀฝߢᲧセߒߚ‫⚿ޕ‬ᨐ‫⛘ޔ‬ኻਇ৻⥌ೝỗߦኻߔࠆᵈⷞᤨ㑆ߪ⋧ኻਇ৻⥌ೝỗߦኻߔࠆᵈⷞᤨ㑆ࠃࠅ߽᦭ᗧߦ
㐳ߊ㧔Z = 2.451‫ ޔ‬p = .014㧕‫ޔ‬0 ᱦఽߩ࠾ࡎࡦࠩ࡞ߪ⋡ᮡೝỗߩേ߈ࠍ๟࿐ೝỗߣ㑐ㅪઃߌߡ⹺⍮ߒߥ޿㧔⛘ኻ⊛⹺
⍮௑ะ㧕ߎߣ߇␜ໂߐࠇߚ‫ޕ‬
B-34 ࠹࠽ࠟࠩ࡞ᄢ⥓ᱤ 3 ᰴరᒻ⁁ߩಽᨆ
ᴡ㊁␞ሶ㧔⑼ඳ࡮ੱ㘃㧕 ᚲౝኻᔕ⠪㧦㜞੗ᱜᚑ
⃻↢ߩࡅ࠻ߣᄢဳ㘃ੱ₎ߦߟ޿ߡ‫ޔ‬ᄢ⥓ᱤਃᰴరᒻ⁁ࠍ⹦⚦ߦಽᨆߒߚ⚿ᨐ‫ߩ⾰࡞ࡔ࠽ࠛޔ‬ෘߐߣಽᏓߩ․ᓽ߇‫ޔ‬
ฦ⒳ߩ㘩ᕈߦᔕߓߚㆡᔕ⊛ߥ߽ߩߢ޽ࠆߎߣ߇‫ߦߢ߹ࠇߎޔ‬᣿ࠄ߆ߦߥߞߡ޿ࠆ‫ߚߒ߁ߎߚ߹ޕ‬ᚻᴺࠍᔕ↪ߒߡ‫ޔ‬
ൻ⍹ੱ㘃߿ൻ⍹㘃ੱ₎ߦߟ޿ߡ߽ห᭽ߩᬌ⸛ࠍߒߚߣߎࠈ‫ߩࠬࠢ࠹ࡇࠖ࠺࡞ࠕޔ‬ᄢ⥓ᱤᒻ⁁߇⃻↢࠴ࡦࡄࡦࠫ࡯ߣ
ߪᯏ⢻⊛ߦ⇣ߥࠆ߽ߩߢ޽ࠆߎߣ߿‫ޔ‬ਛᣂ਎ᄢဳ㘃ੱ₎ߩ࠴࡚ࡠ࡯࡜ࡇ࠹ࠢࠬߩᄢ⥓ᱤ߇⃻↢ࠧ࡝࡜ߣ㘃ૃߒߚᯏ
⢻ㆡᔕࠍ⷗ߖࠆߎߣߥߤ߇᣿ࠄ߆ߣߥߞߚ‫⎇ᧄޕ‬ⓥߢߪ࠹࠽ࠟࠩ࡞ᄢ⥓ᱤᒻ⁁ࠍ CT ࠬࠠࡖ࠽ߢ᠟௝ߒߡౣ᭴▽ߒ‫ޔ‬
⃻↢ᄢဳ㘃ੱ₎╬ߣᲧセߒߚ‫↢⃻ޕ‬㘃ੱ₎ߩߥ߆ߢߪ໑৻‫ޔ‬૕ࠨࠗ࠭߇ዊဳߢ޽ࠆ࠹࠽ࠟࠩ࡞ߪ‫ޔ‬ᄢ⥓ᱤߦߟ޿ߡ
߽‫ޔ‬ᄢဳߩ㘃ੱ₎ߣၮᧄᒻᘒࠍ౒᦭ߒߡߪ޿ࠆ߽ߩߩ‫ޔ‬ᄢ߈ߐߩ㆑޿ߪᱧὼߣߒߡ޿ࠆ‫ޔ‬ಽᨆߩ⚿ᨐ‫࡞ࠩࠟ࠽࠹ޔ‬
ᄢ⥓ᱤߪࠛ࠽ࡔ࡞⾰߇⛘ኻ⊛ߦ⭯ߊ‫ߟ߆ޔ‬Ყセ⊛ဋ৻ߦಽᏓߒߡ޿ࠆߎߣ߇ࠊ߆ߞߚ‫ߚ߹ޕ‬ຏว㕙┃ߩࠛ࠽ࡔ࡞⾰
߇஥ოߩࠛ࠽ࡔ࡞⾰ߦኻߒߡ⭯޿․ᓽ߇⷗ࠄࠇ‫ߩߎޔ‬ὐߦ߅޿ߡᨐታ㘩ߩ࠴ࡦࡄࡦࠫ࡯ߣ㘃ૃߔࠆߎߣ߽᣿ࠄ߆ߣ
ߥߞߚ‫ޕ‬
B-35 ⦡⋤ࠩ࡞ߩ⦡ⷡ․ᕈߩⴕേ⊛⎇ⓥ
ዊ᧻⧷ᒾ㧔↢ℂ⎇࡮✚⎇ᄢ㧕㧘ㇹ↰⋥৻㧘ᮮ੗ഞ㧘㜞ᧁᱜᶈ㧔↢ℂ⎇㧕㧘ጟỈ೰⿠㧘ᵄ㑆ᥓⴕ㧔✚⎇ᄢ࡮↢๮⑼ቇ㧕㧘
㟃↰ቁ๺㧔⼾ᯅᛛ⑼ᄢ㧕 ᚲౝኻᔕ⠪㧦ች࿾㊀ᒄ
ࠗࡦ࠼ࡀࠪࠕ↱᧪ߩ L ㍙૕ᰳ៊ߦࠃࠆ 2 ⦡ဳ⦡⋤ࠩ࡞ߩ⦡ⷡ․ᕈࠍ᣿ࠄ߆ߦߔࠆߚ߼ߦ‫ޔ‬ㆮવ⊛ߦหቯߐࠇߡ޿
ࠆ 2 ⦡ဳ⦡⋤ࠩ࡞ߣ 3 ⦡ဳᱜᏱࠩ࡞ࠍ↪޿ߡⴕേታ㛎ࠍⴕߞߚ‫ޕ‬2 ⒳㘃ߩᵄ㐳㧔592nm ߣ 660nm㧕ߩ LED ࠍ▫ᒻ
ߩⷞⷡೝỗ๒␜ⵝ⟎ߦ౉ࠇ‫ޔ‬೨㕙ߦ⸳ߌߚⓣ㧔⋥ᓘ 8mm㧕߆ࠄ࠺ࠖࡈࡘ࡯ࠩ࡯ࠍ੺ߒߡᾖ኿ߒߚ‫ޕ‬᭽‫ߥޘ‬ノᐲߩ
ೝỗࠍ↪޿ߡᬌ಴㑣୯ࠍ᷹ቯߒߚ‫ޕ‬3 ⦡ဳߣᲧセߔࠆߣ 2 ⦡ဳߢߪ 660nm ߩߣ߈ߦᬌ಴㑣୯ߩ਄᣹߇⷗ࠄࠇߚ‫ߐޕ‬
ࠄߦ⍹ේᑼᬌᩏ⴫ࠍᮨߒߚⷞⷡೝỗࠍ↪޿ߡ⦡ᑯ೎⺖㗴ࠍⴕߞߚ‫ⷡⷞޕ‬ೝỗߪⶄᢙߩ࠼࠶࠻ߦࠃߞߡ᭴ᚑߐࠇᱜᣇ
ᒻߩᄖᒻࠍᜬߟ‫ⷡⷞߩߎޕ‬ೝỗࠍ᳓ᐔߦ 3 ߟਗߴߡᶧ᥏࠺ࠖࠬࡊ࡟ࠗ਄ߦ๒␜ߒ‫ ߩߜ߁ߩߘޔ‬1 ߟߦߟ޿ߡⅣ⁁ߩ
ㇱಽߦ฽߹ࠇࠆ࠼࠶࠻ߩ⦡ࠍᄌൻߐߖ࠲࡯ࠥ࠶࠻ೝỗߣߒߚ‫࠻࠶ࠥ࡯࠲ߩ⋧⦡ߥ߹ߑ߹ߐޕ‬ೝỗࠍ↪޿ߡታ㛎ࠍⴕ
ߞߚ‫ޕ‬2 ⦡ဳߢߪᷙห⦡✢਄ߩ⦡⋧ߢᬌ಴₸ߩૐਅ⷗ࠄࠇ‫ޔ‬3 ⦡ဳߣߪ⇣ߥࠆ௑ะࠍ␜ߒߚ‫⚿ߩࠄࠇߎޕ‬ᨐߪ 2 ⦡
ဳ⦡⋤ࠩ࡞ߩ⦡ⷡ․ᕈࠍ෻ᤋߒߡ޿ࠆߣ⠨߃ࠄࠇࠆ‫ޕ‬
㧨ቇળ⊒⴫㧪
1) Koida K, et al. (2011) Color discrimination performance of genetically identified dichromatic macaques.The 21st
symposium of the International colour vision society, ICVS, (2011/07, Kongsberg Norway)
2) Koida K, et al. (2011) ੑ⦡ᕈࡑࠞࠢࠩ࡞ߩⴕേታ㛎ߦࠃࠆ⦡ⷡ࠹ࠬ࠻, Behavioral test of dichromatism in
genetically identified dichromatic macaques.╙ 34 ࿁ᣣᧄ␹⚻⑼ቇᄢળ,㧔2011/09, ᮮᵿ㧕
3) Koida K, et al. (2011) Color discrimination performance of dichromatic macaque monkey. The Asia-Pacific
Interdisciplinary Research Conference 2011, AP-IRC, (2011/11, Toyohashi, Japan)
B-36 ࡑࠞࠢߩᕈ⊹⣲⣘ߦ㑐ߔࠆಽሶၮ⋚⎇ⓥ
ዊ㊁⧷ℂ㧘⍹↰⾆ᢥ㧔᧲ᄢ࡮㒮࡮↢‛⑼ቇ㧕 ᚲౝኻᔕ⠪㧦㋈ᧁ᮸ℂ
㔤㐳㘃ߦߪߘߩ⊒ᖱᦼߦ᣿⏕ߥᕈ⊛ࠪࠣ࠽࡞ࠍ⊒ߔࠆ⒳߇޽ࠆ‫ࠢࠞࡑ߫߃଀ޕ‬ዻߩ޿ߊߟ߆ߩ⒳ߢߪࡔࠬߩᕈ⊹
⣲⣘㧔ߎߎߢߪ૕ⓍჇടߣ⚃ầࠍ฽߻㧕߇⿠ߎࠆߎߣ߇⍮ࠄࠇߡ޿ࠆ‫ޕ‬ᚒ‫ߩߎߪޘ‬ᕈ⊹⣲⣘ߦ⌕⋡ߒ‫ޔ‬ਥߦ૕ⓍჇ
ടߣ⚃ầ߇⋡┙ߟࠕࠞࠥࠩ࡞ߣ‫ߩߘޔ‬ㄭ✼⒳ߢ޽ࠆ࠾ࡎࡦࠩ࡞ࠍኻ⽎ߣߒߡ‫ޔ‬ᕈ⊹⦡‫ޔ❱⚵ޔ‬ㆮવሶߩᄌൻࠍㅊߞ
ߡ޿ࠆ‫ޕ‬ਔ⒳ߪᕈ⊹ߩ૕Ⓧᄌേߦ߅޿ߡᏅ߇⷗ࠄࠇ‫ߩ࡞ࠩࡦࡎ࠾ޔ‬ኙ಄ㆡᔕ߽⠨߃ࠄࠇࠆ‫ᧄޕ‬ᐕᐲߪ‫ޔ‬ᤓᐕᐲ⏕┙
ߒߚታ㛎♽ߦၮߠ޿ߡฦ⒳ታ㛎ࠍⴕߞߚ‫ޔߡߒߣߟߣ߭ߩߘޕ‬HE ᨴ⦡⚵❱ࠍ↪޿ߡⴊ▤ߩᢙࠍ⸃ᨆߒߚߣߎࠈ‫ޔ‬
ࠕࠞࠥࠩ࡞ߩᕈ⊹⚃ầߣⴊ▤ᢙߦᱜߩ⋧㑐߇⷗ࠄࠇߚ‫ޕ‬㧔⋧㑐ଥᢙ r=0.85㧕ߒ߆ߒ࠾ࡎࡦࠩ࡞ߢߪ⋧㑐߇⷗ࠄࠇߥ
߆ߞߚߎߣ߆ࠄ‫ޔ‬⒳ߦࠃߞߡ⚃ầ߇⇣ߥࠆࡊࡠ࠮ࠬߢ⃻ࠇߡ޿ࠆน⢻ᕈ߇޽ࠆ‫ޕ‬ᕈ⊹ߪౝಽᴲ♽ߦࠃࠅ⺞▵ߐࠇߡ
޿ࠆ߇‫ࠅࠃߦࡦࠥࡠ࠻ࠬࠛ߫߃଀ޔ‬ሶችౝ⤑ߩⴊ▤ᣂ↢߇⿠ߎࠆߥߤ‫ޔ‬ౝಽᴲ♽ಽሶၮ⋚ߦ㑐ߒߡࡅ࠻ߣߩᯏ⢻㘃
ૃᕈߪ⥝๧ᷓ޿‫੹ޕ‬ᓟߪౝಽᴲ♽ฃኈ૕ߩㆮવሶ⊒⃻ࠍ⺞ߴ‫ࠍ࡞ࠗࠔࡈࡠࡊߥ⚦⹦ࠅࠃޔ‬ᓧࠆ‫ޕ‬
㧨⊒⴫᭎ⷐ㧪
ಽశ᷹⦡⸘ࠍ↪޿ߚࡑࠞࠢᕈ⊹⦡ᄌൻߩ CIELAB ⦡ⓨ㑆ߦ߅ߌࠆ⴫␜㧔╙ 27 ࿁㔤㐳㘃ቇળ㧔›ጊ㧕㧕
ዊ㊁⧷ℂ㧔᧲ᄢ࡮㒮࡮↢‛⑼ቇ㧕‫ޔ‬⍹↰⾆ᢥ㧔หᏀ㧕‫ޔ‬㋈ᧁ᮸ℂ㧔㔤㐳⎇㧕
ࡑࠞࠢߢߪ⊒ᖱᦼߦࡔࠬߩᕈ⊹ᄌൻ㧔⣲⣘߆ߟ/߹ߚߪ⚃ầ㧕߇⿠ߎࠆ⒳߇޽ࠆ‫ߩߎޕ‬ᄌൻߪౝಽᴲ♽ߦࠃߞߡ
⺞▵ߐࠇ‫ߩࠬࡔޔ‬ᕈ⊹⦡߇ࠝࠬߩ❥ᱺⴕേߦᓇ㗀ߔࠆߣߩႎ๔߇޽ࠆ߇‫ߩࠬࡔޔ‬ᅧᕈߣߩ㑐ㅪߪᧂ⸃᣿ߥㇱಽ߇ᄙ
㧙㧙
޿‫ޕ‬వⴕ⎇ⓥߢߪᕈ⊹⦡᷹ቯߦࠞ࡜࡯࠴ࡖ࡯࠻ࠍ↪޿ࠆߎߣ߇ᄙߊ‫ߩߎޔ‬ᣇᴺߢߪⅣႺశߩᓇ㗀‫⹏ޔ‬ଔᤨߩⷰኤ⠪
ߩਥⷰࠍឃ㒰ߢ߈ߥ޿‫ߚߒ߁ߎޕ‬ẜ࿷⊛ᓇ㗀ࠍ㒰ᄖߔࠆߚ߼ߦ‫⎇ᧄޔ‬ⓥߢߪಽశ᷹⦡⸘㧔MINOLTA CG-411C㧕ࠍ
↪޿‫ޔ‬CIELAB ⦡ⓨ㑆ߢ⹏ଔߒߚ‫ޕ‬CIELAB ⦡ⓨ㑆ߪ 3 ゲ㧔⿒✛‫ޔ‬㕍㤛‫ޔ‬᣿ᥧ㧕ߢ᭴ᚑߐࠇࠆ‫࡞ࠩࠥࠞࠕޕ‬㧔Macaca
mulatta㧕9 㗡ߣ࠾ࡎࡦࠩ࡞㧔Macaca fuscata㧕5 㗡ࠍᬌ⚝ኻ⽎ߣߒ‫ޔ‬㕖⊒ᖱᦼ㧔7 ᦬㧕ߣ⊒ᖱᦼ㧔10 ᦬㧕ߦ‫ޔ‬ห৻୘
૕ߩᕈ⊹⦡ࠍ᷹ቯߒߚ‫⦡ࠆߥ⇣ߢ࡞ࠩࡦࡎ࠾ߣ࡞ࠩࠥࠞࠕޕ‬ᄌൻ߇⷗ࠄࠇ‫ޔ‬CIELAB ߩ‫ޟ‬㕍㤛‫ޠ‬ゲߦ߅ߌࠆჇട
߇‫᦭ߢߺߩ࡞ࠩࠥࠞࠕޔ‬ᗧߢ޽ߞߚ㧔t ᬌቯ‫᦭ޔ‬ᗧ᳓Ḱ 5%㧕‫ⴊߩ࡞ࠩࠥࠞࠕߪࠇߎޕ‬ᵹᄌേ߇࠾ࡎࡦࠩ࡞ࠃࠅ߽ᄢ
߈޿ߎߣࠍ␜ໂߔࠆ‫ⴊޕ‬ᵹߩჇᄢߪ⚃ầߩߺߥࠄߕ⣲⣘ߦߟߥ߇ࠆߚ߼‫ⴊߩ࡞ࠩࡦࡎ࠾ޔ‬ᵹ߇ᄌേߒߥ޿ߎߣߪ‫ޔ‬
ኙ಄ㆡᔕߣߩ㑐ㅪ߆ࠄ⥝๧ᷓ޿‫ޕ‬ಽశ᷹⦡⸘ࠍ↪޿‫ޔ‬ᕈ⊹⦡ᄌൻࠍ CIELAB ⦡ⓨ㑆ߦ߅޿ߡቴⷰ⊛ߦ⴫␜ߢ߈‫ޔ‬ㄭ
✼ߥࡑࠞࠢ⒳㑆ߢ⚃ầߩ㆑޿ࠍᬌ಴ߢ߈ߚ‫ޕ‬
B-37 ࡑࠞࠢዻ㔤㐳㘃ߦ߅ߌࠆᗵᨴ∝ᛶ᛫ᕈߩᄙဳߣࠥࡁࡓㅴൻ
቟ᵄ㆏㇢㧔㐳ፒᄢ࡮ᾲᏪකቇ⎇࡮⥃ᐥᗵᨴ∝ቇ㧕 ᚲౝኻᔕ⠪㧦ᐔ੗໪ਭ
ࡑࠞࠢዻ㔤㐳㘃ߪ⒳ಽൻߩㆊ⒟ߢ᫮ᕷⅣႺߩᓇ㗀ਅߦߘࠇߙࠇߩࠥࡁࡓࠍㅴൻߐߖ⛯ߌߡ޿ࠆߣ⠨߃ࠄࠇࠆ‫ޕ‬ᗵ
ᨴ࿃ሶߩ࿾ℂ⊛ಽᏓߪࠥࡁࡓᄙဳߩ࿾ℂ⊛⋧㆑↢ᚑߩේേജߣ⠨߃ࠄࠇ‫⸃ߩߘޔ‬᣿ߪ↢‛⒳߇޿߆ߦലᨐ⊛ߦᗵᨴ
࿃ሶߦኻಣߒߡ޿ࠆ߆ߩℂ⸃ࠍᷓ߼ࠆߎߣߦߟߥ߇ࠆ‫⎇ᧄޕ‬ⓥߢߪ Toll ᭽ฃኈ૕ TLR2‫ޔ‬TLR4‫ޔ‬TLR9 ߩᄌ⇣߿ᄙ
ဳ߇ࡅ࠻߿ࡑ࠙ࠬߢߪ∛ේᓸ↢‛↱᧪ߩ‛⾰ߩ⹺⼂ࠍᄌൻߐߖࠆߎߣ߆ࠄ‫ࠢࠞࡑޔ‬ዻ㔤㐳㘃ߦߟ޿ߡߘࠇࠄߩႮၮ
㈩೉ࠍ⸃ᨆߒ‫ޔ‬⒳ౝ߅ࠃ߮⒳㑆ߢߩ㕖ห⟵⟎឵ࠍ⹏ଔߒߚ‫ ߜ߁ߩߘޕ‬TLR2 ߦ㑐ߒߡ࠾ࡎࡦࠩ࡞ߢߪࠦ࡯࠼㗔ၞߩ
ో⥸ߦਗ਼ߞߡ㕖ห⟵⟎឵ߪ㗫ᐲ߇ૐ޿௑ะߦ޽ࠆߩߦኻߒߡࠕࠞࠥࠩ࡞ߢߪ‫⚦ߩ⊕Ⱞ⤑ޔ‬⢩ᄖㇱಽߦ⋧ᒰߔࠆ㗔ၞ
ߩ৻ㇱߦዪᚲ⊛ߦ㕖ห⟵⟎឵ߩ㓸Ⓧߔࠆㇱಽ߇⹺߼ࠄࠇ‫ޔ‬ᄙ᭽ᕈ₪ᓧㅴൻߩነਈ߇ផቯߐࠇߚ‫ࠞࠕߣ࡞ࠩࡦࡎ࠾ޕ‬
ࠥࠩ࡞ߩ㑆ߢ 326 ⇟⋡ߩࠕࡒࡁ㉄߇ߘࠇߙࠇ࠴ࡠࠪࡦ‫ߦࡦࠡ࡜ࡄࠬࠕޔ‬࿕ቯߒߡ߅ࠅ‫ߩߎޔ‬ㇱ૏ߪࡅ࠻ߩಽሶ᭴ㅧ
⸃ᨆ߆ࠄ࡝ࠟࡦ࠼⚿วߦ㑐ਈߔࠆߣߐࠇߡ޿ࠆߎߣ߆ࠄߎߩᄌൻ߇ࠕࠞࠥࠩ࡞ߢߩᄙ᭽ᕈߩⓍᭂ⊛ߥ⫾Ⓧࠍ߽ߚ
ࠄߒߡ޿ࠆߣផ᷹ߒ‫ޔ‬ಽሶࡕ࠺࡝ࡦࠣߦࠃࠆಽᨆࠍⴕߥߞߚ‫ޕ‬㧔1㧕߹ߚ‫ࠕ࡝࡜ࡑ࡞ࠨޔ‬ේ⯻ Plasmodium coatneyi
ߩታ㛎ᗵᨴߦ߅޿ߡ㜞ᗵฃᕈߢ޽ࠆ࠾ࡎࡦࠩ࡞ߣᛶ᛫ᕈߢ޽ࠆࠞ࠾ࠢࠗࠩ࡞ߩ⒳㑆ߦ߅޿ߡࡑ࡜࡝ࠕ⦡⚛ࠍ⹺⼂
ߔࠆ TLR9 ߩㆮવሶߦⶄᢙߩ㕖ห⟵⟎឵ࠍ⹺߼ߚ‫߇ࠄࠇߎޕ‬୘૕࡟ࡌ࡞ߢߩᗵฃᕈߩ⋧㆑ࠍ⺑᣿ߔࠆ߆ࠍᬌ⸛ߒߡ
޿ࠆ‫ާޕ‬ᢥ₂ި1㧚Takaki A, et al㧚Immunogenetics 64:15- 29 (2012)㧚
B-38 ࠾ࡎࡦࠩ࡞㓽ߩᩕ㙃⁁ᘒߣ㙄₪ᓧ㊂ߩ㗅૏ᩰᏅߦ㑐ߔࠆ㜞ፒጊ⟲ߣᐘፉ⟲ߩᲧセ
ᩙ↰ඳਯ㧔ᄢಽᏒᢎ⢒ᆔຬળ࡮ᢥൻ⽷⺖㧕 ᚲౝኻᔕ⠪㧦ự↰ⓛ
ᐘፉߢߪ 8 ᦬ߦ‫ޔ‬㜞ፒጊߢߪ 9 ᦬ߦ‫ޔ‬౮⌀⸘᷹ᴺߦࠃࠆᚑᾫ㓽ߩ૕㐳⸘᷹ࠍⴕߞߚ㧔ᐘፉ⟲㧦11 㗡㧧㜞ፒጊ⟲㧦
10 㗡㧕
‫ޕ‬㜞ፒጊ⟲ߩ૕㐳⸘᷹ߪ⚂ 10 ᐕಽߩ࠺࡯࠲߇޽ࠅ‫ޔ‬ᚑᾫᓟߪട㦂ߦ઻߁⍴❗߇⹺߼ࠄࠇߥ޿௑ะ߇ࠊ߆ࠅ
ߟߟ޽ࠆ߇‫ޔ‬ᐘፉ⟲ߩ૕㐳⸘᷹ߪ 2008 ᐕᐲ߆ࠄߩ㐿ᆎߢ޽ࠅ‫⺞ߛ߹ޔ‬ᩏᐕᢙ߇ዋߥߊ‫ޔ‬ᐕ㦂ᄌൻߩ௑ะࠍ᣿ࠄ߆
ߦߔࠆߦߪ⥋ߞߡ޿ߥ޿‫ޕ‬
㜞ፒጊ㓽ߩ૕㊀ߪ‫⥄ޔ‬りߦࠃࠆ࠺࡯࠲ߩ⫾Ⓧࠍ߶߷ 10 ᐕ㑆ⴕߞߡ߈ߡ߅ࠅ‫ޔ‬㜞㗅૏㓽ߪૐ㗅૏㓽ࠃࠅ߽㊀ߊ‫ޔ‬
⢒ߡ਄ߍࠆሶߩ૕㊀߽㊀޿ߎߣ߇ࠊ߆ߞߡ޿ࠆ‫৻ޕ‬ᣇ‫ޔ‬ᐘፉ㓽ߢߪ‫ޔ‬૕㐳⸘᷹ኻ⽎୘૕ߦ㒢ߞߡ‫ޔ‬૕㊀෸߮❥ᱺᚑ
❣ࠍ੩ㇺᄢቇ㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯ࠃࠅ࠺࡯࠲ࠍ୫↪ߒ‫ޔ‬ಽᨆࠍⴕߞߡ޿ࠆ߇‫ޔ‬2011 ᐕᐲߦ᣿ࠄ߆ߦߥߞߚߎߣߪ‫ޔ‬
ᰴߩߣ߅ࠅߢ޽ࠆ‫ޕ‬
Ԙ ᐘፉᚑᾫ㓽ߩ૕㊀ߪ⪺ߒ޿ᐕᄌേࠍ␜ߔ߇‫ߪࠇߎޔ‬୘૕㑆ߢ᣿⍎ߦห⺞ߒߡ߅ࠅ‫⥄ޔ‬ὼ㘩‛ߩ⼾ರ߇ᓇ㗀
ߒߡ޿ࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬
ԙ ᐘፉ㓽ߩ૕㊀ߪ㜞ፒጊ㓽ߩߘࠇࠃࠅ߽᦭ᗧߦシ߆ߞߚ‫ޕ‬
Ԛ ߶ߣࠎߤߔߴߡߩᐕ㦂ߦ߅޿ߡ‫ޔ‬㜞ፒጊ㓽ࠃࠅ߽ᐘፉ㓽ߩᣇ߇಴↥₸߇ૐ߆ߞߚ‫ޕ‬
߹ߚ‫ޔ‬㙄₪ᓧ㊂⺞ᩏߢߪ‫ޔ‬㜞ፒጊ㓽ߢߪ㜞㗅૏୘૕ߪૐ㗅૏୘૕ߩ⚂ 2.2 ୚ߩࠞࡠ࡝࡯ࠍ㙄㧔ࠦࡓࠡߣࠨ࠷ࡑࠗ
ࡕ㧕߆ࠄᓧߡ޿ࠆߎߣ߇ᣢߦࠊ߆ߞߡ޿ࠆ‫ޕ‬2011 ᐕᐲߪᐘፉ⟲ߦ߅޿ߡ㙄₪ᓧ㊂ߩ⺞ᩏࠍ㐿ᆎߒߚ߇‫ޔ‬บ㘑ធㄭߥ
ߤߦࠃࠅ 2 ᣣ㑆ߒ߆⺞ᩏ߇ߢ߈ߥ߆ߞߚ‫ޕ‬㜞㗅૏㓽ߣૐ㗅૏㓽ฦ 1 㗡ߕߟߩ㙄㧔ࠦࡓࠡ㧕₪ᓧ㊂⺞ᩏ⚿ᨐߦߔ߉ߥ
޿߇‫ޔ‬ᐘፉߢ߽‫ޔ‬㜞㗅૏㓽ߩᣇ߇ᄙߊߩ㙄ࠍ₪ᓧߒߡ޿ߚ‫ޕ‬
੹ᓟ‫ޔ‬ᐘፉ⟲ߣ㜞ፒጊ⟲ߩ㑆ߢߩ㙄₪ᓧ㊂࡮૕ࠨࠗ࠭࡮❥ᱺᚑ❣ߦߟ޿ߡߩ⺞ᩏࠍ⛮⛯ߒ‫ߩࠇߙࠇߘޔ‬ታᘒࠍࠃ
ࠅ⹦⚦ߦ⸃᣿ߒߡࠁ߈ߚ޿‫ޕ‬
㧨ቇળ⊒⴫㧪
1) ᩙ↰ඳਯ㧘߶߆ ࠾ࡎࡦࠩ࡞㓽ߩ૕ࠨࠗ࠭ߣ಴↥ᚑ❣ߩᐕ㦂ᄌൻ㧙㜞ፒጊ୘૕⟲ߣᐘፉ୘૕⟲ߩᲧセ㧙㧚
ᣣᧄ㔤㐳㘃ቇળ╙ 27 ࿁ᄢળ 2011 ᐕ 7 ᦬
2) ᩙ↰ඳਯ㧘߶߆ ࠾ࡎࡦࠩ࡞㓽ߦ߅ߌࠆ૕㊀࡮૕㐳࡮಴↥₸ߩᐕ㦂ᄌൻ᭽ᑼߩ୘૕⟲㑆Ყセ㧙㜞ፒጊߣᐘ
ፉ㧙㧚ᣣᧄູ੃㘃ቇળ 2011 ᐕᐲᄢળ 2011 ᐕ 9 ᦬
㧨⺰ᢥ⊒⴫㧪
Kurita H, et al. (2012) A photogrammetric method to evaluate nutritional status without capture in habituated
free-ranging Japanese macaques (Macaca fuscata): a pilot study Primates㧘53(1):7-11.
B-39 ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ࠻࡜࠶ࠢࡏ࡯࡞ᑼജ⸅ⷡ࠺ࠖࠬࡊ࡟ࠗࠍ↪޿ߚᲧセ⹺⍮⎇ⓥ
㈬੗ၮⴕ㧔ฬฎደᎿᬺᄢ࡮㒮࡮ᯏ⢻Ꮏቇ㧕
㧘↰ਛ↱ᶈ㧘૒㊁᣿ੱ㧔ฬฎደᎿᬺᄢ࡮ᯏ⢻Ꮏቇ㧕㧘⮮ᧄ⧷㓶㧔ฬฎደᎿ
ᬺᄢ࡮ᖱႎᎿቇ㧕 ᚲౝኻᔕ⠪㧦෹᳗㓷Ꮖ
㧙㧙
ᧄ⎇ⓥߪ‫ޔ‬ᣢߦ࠴ࡦࡄࡦࠫ࡯ߢ૶↪ታ❣ߩ޽ࠆ࠻࡜࠶ࠢࡏ࡯࡞ࠍ߽ߣߦ‫ޔ‬ജ⸅ⷡߩឭ␜߇น⢻ߥⵝ⟎ࠍ㐿⊒ߒ‫ޔ‬
ߎࠇࠍ↪޿ߡ࠴ࡦࡄࡦࠫ࡯ߦࠃࠆ⹺⍮ታ㛎ࠍⴕ߁ߎߣࠍ⋡⊛ߣߒߚ‫ߦ․ޕ‬ജ⸅ⷡߩᑯ೎߿ද⺞૞ᬺߦߟ޿ߡᲧセ⹺
⍮⑼ቇߩⷰὐ߆ࠄ⠨ኤߔࠆ‫੹ޕ‬ᐕᐲߪജ⸅ⷡߩᑯ೎ታ㛎ߩㆀⴕߩߚ߼ߩ⸠✵ࠍ㐿ᆎߒߚ‫࡯ࡏࠢ࠶࡜࠻ޔߦ߼ߓߪޕ‬
࡞ࠍㅢߓߡᠲ૞ߔࠆ↹㕙਄ߩࠞ࡯࠰࡞ࠍ‫ޔ‬㕒ᱛߔࠆ࠲࡯ࠥ࠶࠻ߦวࠊߖࠆ࠲ࠬࠢࠍⴕߞߚ‫࡞࡯ࡏࠢ࠶࡜࠻ޔ߅ߥޕ‬
ߩᠲ૞ߦ߅޿ߡߪ‫ޔ‬ജⷡࡈࠖ࡯࠼ࡃ࠶ࠢ߇ឭ␜ߐࠇߡ޿ࠆ‫⃻ޕ‬࿷ߪ‫ޔ‬ᰴߩࠬ࠹࠶ࡊߣߒߡ‫ࠍ࡞࠰࡯ࠞޔ‬േߊ࠲࡯ࠥ
࠶࠻ߦวࠊߖ‫ޔ‬ㅊᓥߔࠆታ㛎ࠍⴕߞߡ޿ࠆ‫ ߦߢ߹ࠇߎޔ߅ߥޕ‬6 ฬߩ࠴ࡦࡄࡦࠫ࡯߇‫ߩߎޔ‬ജⷡࡈࠖ࡯࠼ࡃ࠶ࠢઃ
߈࠻࡜࠶ࠢࡏ࡯࡞ࠍ↪޿ߡࠞ࡯࠰࡞ࠍ࠲࡯ࠥ࠶࠻ߦวࠊߖࠆߎߣ߇ߢ߈ߚ‫ޔߪߦ⊛⚳ᦨޕ‬ᑯ೎ታ㛎ࠍ᜛ᒛߒ‫ޔ‬ജ⸅
ⷡࠍ฽߻૞ᬺߩㆀⴕ߿ੑ୘૕ߩ࠴ࡦࡄࡦࠫ࡯ߩද⺞૞ᬺߦߟ޿ߡߩታ㛎ࠍⴕ߁੍ቯߢ޽ࠆ‫ޕ‬
B-40 ࡑ࠳ࠟࠬࠞ࡞↥⒘ዋේ₎㘃ߩㆮવ⊛▤ℂᴺߩ⏕┙
ቬㄭഞ㧔(⽷)ㅴൻ↢‛ቇ⎇ⓥᚲ㧕 ᚲౝኻᔕ⠪㧦↰ਛᵗਯ
ࡑ࠳ࠟࠬࠞ࡞↥ේ₎ߩߥ߆ߢ߽‫⛘ߦ․ޔ‬Ṍ߇ෂᗋߐࠇߡ޿ࠆ Eulemur macaco ߣ Varecia rubra ࠍኻ⽎ߣߒߡㆮવ
ሶࡑ࡯ࠞ࡯ࠍ૶ߞߚⴊ⛔▤ℂᴺࠍ⏕┙ߔࠆߚ߼‫ޔ‬࿖ౝߢ㘺⢒ߐࠇߡ޿ࠆ୘૕⟲㧔Varecia rubra 14 ୘૕‫ޔ‬Varecia v.
variegata 5 ୘૕‫ޔ‬Varecia spp 3 ୘૕‫ޔ‬Eulemur macaco 50 ୘૕㧕߆ࠄญౝ⚦⢩ࠍណขߒ‫ޔ‬DNA ࠨࡦࡊ࡞ࠍ⺞ᢛߒߚ‫ޕ‬
ߎࠇ߹ߢߩ౒ห೑↪⎇ⓥߢ⏕┙ߒߚ microsatellite ㆮવሶᐳ૏ߦߟ޿ߡ Multiplex ᴺߢಽᨆࠍⴕߞߚ‫ޔߚ߹ޕ‬ᣂߒߊ
ㆮવ⊛▤ℂߦ૶߃ࠆㆮવሶᐳࠍ᳞߼ߡ‫ޔ‬Varecia rubra ߦ߅޿ߡ 51HDZ247‫ޔ‬598‫ޔ‬646‫ޔ‬833‫ޔ‬985 ㆮવሶᐳࠍჇ᏷ߔ
ࠆࡊ࡜ࠗࡑ࡯ࠍ⹜ߺߚߣߎࠈ‫ޔ‬833 ߩჇ᏷߇ਇ቟ቯߢ޽ߞߚ߇‫ޔ‬247‫ޔ‬598‫ޔ‬646‫ޔ‬998 ߩ 4 ㆮવሶᐳ૏ߦᄙဳ߇ߺࠄ
ࠇ‫᦭ޔ‬ലߢ࿷ࠆ੐߇್᣿ߒߚ‫ޕ‬Varecia ߢߪ‫ޔ‬Eulemur fulvus ↪ߦ㐿⊒ߐࠇߚ Efr09 ߣ Eulemur mongoza ↪ߦ㐿⊒ߐࠇ
ߚ Em9 ߩ 2 ߟߩࡊ࡜ࠗࡑ࡯߇᦭ലߢ޽ࠆ੐߇್᣿ߒߚ‫ޕ‬ಽᨆߩ⚿ᨐ‫ޔ‬Varecia rubra ߩ㘺⢒୘૕⟲ߦ‫ޔ‬ῳⷫ߇㑆㆑ߞ
ߡ⊓㍳ߐࠇߡ޿ࠆ୘૕߇᣿ࠄ߆ߦߥࠅ‫ߩߎޔ‬᭽ߥ⚿ᨐ߆ࠄ‫⊓⛔ⴊޔ‬㍳ߦㆮવሶࡑ࡯ࠞ࡯ࠍ૶߁▤ℂᴺߩᔅⷐᕈࠍ∩
ᗵߒߚ‫ޕ‬
B-41 㔤㐳㘃ߩశᗵⷡࠪࠬ࠹ࡓߦ㑐ࠊࠆ࠲ࡦࡄࠢ⾰ߩ⸃ᨆ
ዊፉᄢテ㧘᫪ථ㧘㠽ዬ㓷᮸㧔᧲੩ᄢ࡮㒮ℂ࡮↢‛ൻቇ㧕 ᚲౝኻᔕ⠪㧦੹੗໪㓶
⣄ᬁേ‛ߦ߅޿ߡ‫ߡૃߪߣ⾰‛ⷞޔ‬㕖ߥࠆశฃኈⰮ⊕⾰㧔㕖ⷞⷡဳࠝࡊࠪࡦ㧕߇ᢙᄙߊหቯߐࠇߡ޿ࠆ‫ߪ౒⑳ޕ‬
ᦨㄭ‫ޔ‬㕖ⷞⷡဳࠝࡊࠪࡦߩ৻ߟ OPN5 ߇ࡑ࠙ࠬߩ✂⤑㜞ᰴ࠾ࡘ࡯ࡠࡦ߿✂⤑ᄖ⚵❱㧔⣖߿ᄖ⡊㧕ߦ⊒⃻ߔࠆߎߣ‫ޔ‬
ߐࠄߦࡑ࠙ࠬ߿ࡅ࠻ߩ OPN5 ߇ UV ᗵฃᕈߩశฃኈⰮ⊕⾰ߢ޽ࠆߎߣࠍ⷗಴ߒߚ‫[ޕ‬Kojima et al. (2011) PLoS ONE, 6,
e26388] ߎߩߎߣ߆ࠄ‫ޔ‬ᓥ᧪ UV ᗵⷡ߇ߥ޿ߣߐࠇߡ޿ߚ㔤㐳㘃ߦ߽‫ޔ‬UV ᗵฃᕈߩశࠪࠣ࠽࡞⚻〝߇ሽ࿷ߔࠆߎ
ߣ߇␜ໂߐࠇߚ‫⎇ᧄߢߎߘޕ‬ⓥߢߪ‫ޔ‬OPN5 ࠍ੺ߒߚశฃኈ߇㔤㐳㘃ߦ߅޿ߡߤߩࠃ߁ߥ↢ℂ⊛ᓎഀࠍᜂ߁ߩ߆ࠍ
ផቯߔࠆߚ߼‫ޔ‬㔤㐳㘃ߦ߅ߌࠆ OPN5 ߩ⊒⃻ㇱ૏ߩหቯࠍ⹜ߺߡ޿ࠆ‫ޔߦߢ߹ࠇߎޕ‬᡼ⴊ߽ߒߊߪἠᵹ࿕ቯߒߚࠨ
࡞୘૕ߩ⚵❱㧔⌒⃿࡮ᄖ⡊ߥߤ㧕ࠃࠅ࿕ቯᮡᧄࠍ૞⵾ߒߚ‫ࠅࠃ⃿⌒ޔߜ߁ߩߎޕ‬૞⵾ߒߚ⚵❱ಾ ߦኻߒߡࡑ࠙ࠬ
OPN5 ᛫૕ࠍ෻ᔕߐߖߚߣߎࠈ‫৻ޔ‬ㇱߩ⚦⢩ߦ㓁ᕈࠪࠣ࠽࡞߇ᬌ಴ߐࠇߚ߇‫ߩ⃿⌒ࠬ࠙ࡑޔ‬႐วߣߪ⇣ߥࠆ⚵❱ౝ
ዪ࿷ࠍ␜ߒߚ‫৻ޕ‬ᣇߎߩࡑ࠙ࠬ OPN5 ᛫૕ߪ‫࠻ࡅޔ‬⢦↱᧪ߩၭ㙃⚦⢩ (HEK293) ߦౝ࿷ߔࠆ㕖 OPN5 ࠲ࡦࡄࠢ⾰
ߦኻߒߡ߽‫ޔ‬ᒝ޿੤Ꮕ෻ᔕࠍ␜ߔߎߣ߇ࠊ߆ߞߚ‫ߩߎޕ‬᛫૕ࠍ↪޿ߚ OPN5 ⊒⃻⚦⢩ߩหቯߪ㔤㐳㘃⹜ᢱߦ߅޿ߡ
ߪ࿎㔍ߢ޽ࠆߣ⠨߃ࠄࠇࠆߚ߼‫ޔ‬ᣂߚߥ OPN5 ᛫૕ߩ૞⵾ߣ‫ޔ‬mRNA ࡟ࡌ࡞ߢߩ⊒⃻⸃ᨆࠍᬌ⸛ߒߡ޿ࠆ‫ޕ‬
B-42 ᣣᧄߢ㊁↢ൻߒߚ࠲ࠗࡢࡦࠩ࡞ߣบḧ࿷᧪⒳ߩᲧセ⎇ⓥ
⯃⑲ᘢ㧔บḧ࿡┙ዳ᧲⑼ᛛᄢቑ࡮㊁↢േ‛଻⢒⎇ⓥᚲ㧕૒ષ⌀⟤㧔㊁↢േ‛଻⼔▤ℂ੐ോᚲ㧕 ᚲౝኻᔕ⠪㧦Ꮉᧄ
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บḧ߆ࠄャ಴ߐࠇ‫ޔ‬ᣣᧄߢ㊁↢ൻߒߚ࠲ࠗࡢࡦࠩ࡞㧔Macaca cyclopis㧕ߪ‫ޔ‬㕍᫪⋵㊁ㄝ࿾ߣ๺᱌ጊ⋵ᄢᳰߢ࿷᧪
ߩ࠾ࡎࡦࠩ࡞ߣ੤㔀ߒߚߎߣ߇⏕⹺ߐࠇߡ޿ࠆ‫᧲ޔߚ߹ޕ‬੩ㇺદ⼺ᄢፉߢߪ࠾ࡎࡦࠩ࡞ߩ޿ߥ޿ⅣႺߢ㊁↢ൻߒߚ
࠲ࠗࡢࡦࠩ࡞߇ోፉߦಽᏓߔࠆߎߣ߇ႎ๔ߐࠇߡ޿ࠆ‫ޕ‬ᣣᧄߢ㊁↢ൻߒߚ࠲ࠗࡢࡦࠩ࡞ߣบḧ࿷᧪ߩ࠲ࠗࡢࡦࠩ࡞
ߩ↢‛ቇ⊛․ᓽࠍᲧߴࠆߎߣࠍ⋡⊛ߦ‫੹ޔ‬ᐕᐲߩ⎇ⓥߢߪ‫ߕ߹ޔ‬಴⥄ߩ໧㗴ࠍ⺞ߴࠆߚ߼‫ ࠕ࡝࠼ࡦࠦ࠻ࡒޔ‬DNA
㧔mtDNA㧕ߩ㈩೉ࠍ⸃⺒ߒ‫ޔ‬ㆮવ⊛․ᓽࠍ⸃ᨆߒߚ‫ޔߚ߹ޕ‬દ⼺ᄢፉࠍ⸰ࠇ‫ޔ‬ⅣႺ߿ࠨ࡞ߩಽᏓ⁁ᴫࠍⷰኤߒ‫ޔ‬
บḧߣߩⅣႺߩ⋧㆑ࠍᬌ⸛ߔࠆၮ␆⾗ᢱࠍᓧߚ‫ޕ‬
บḧߩ 4 ࿾ၞ㧔ർㇱ‫ޔ‬ਛㇱ‫ޔ‬ධ⷏ㇱ‫ޔ‬ධ᧲ㇱ㧕ߢᓧߚ♮⹜ᢱ߆ࠄ DNA ࠍ᛽಴ߒ‫ޔ‬mtDNA ߩ㕖ࠦ࡯࠼㗔ၞߦ޽ࠆ
╙ 2 นᄌၞߩㇱಽ㈩೉ࠍ⸃⺒ߒߚ‫ޕ‬㕍᫪‫ޔ‬๺᱌ጊ‫ޔ‬દ⼺ᄢፉߩ࠲ࠗࡢࡦࠩ࡞ߩ㈩೉ߣᲧセߒߚߣߎࠈ‫ޔ‬mtDNA ߆
ࠄ್ᢿߒߡ‫ޔ‬ᣣᧄߢ㊁↢ൻߒߚࠨ࡞ߪߘࠇߙࠇ಴⥄߇⇣ߥࠅ‫ޔ‬บḧᧄፉߩਛㇱၞ߇๺᱌ጊߦ‫ޔ‬ධㇱၞ㧔ධ᧲ㇱ‫ޔ‬ධ
⷏ㇱߩ඙೎ߪ᣿⍎ߦߪߟ߆ߥ޿㧕߇ਅർߣદ⼺ᄢፉߦ㑐ଥߔࠆ‫␜ߩߣޔ‬ໂࠍᓧߚ‫ޕ‬
ኻᔕ⠪߇㐿⊒ߒߚ♮ DNA ಽᨆᣇᴺߪᣢሽߩ߽ߩߣ⇣ߥࠆ‫ޕ‬㊁ᄖߢណขߔࠆ♮⹜ᢱߩㆮવሶಽᨆߦലᨐ⊛ߥᣇᴺ
߇⠌ᓧߢ߈ߚߩߢ‫੹ޔ‬ᓟߐࠄߦ೎ߩㆮવᮡ⼂ߦߟ޿ߡ߽บḧߢ㊁↢ߩࠨ࡞ߚߜࠍ⺞ᩏߔࠆ㆏߇㐿ߌߚ‫ߩߎޕ‬ᣇᴺࠍ
บḧߢߩ⺞ᩏߦ੹ᓟᔕ↪ߔࠆ⎇ⓥ⸘↹ࠍኻᔕ⠪ߣᬌ⸛ߒ‫ޔ‬ᣣᧄߦᜬߜㄟ߹ࠇߚ࠲ࠗࡢࡦࠩ࡞ߣㆮવ⊛߅ࠃ߮↢ᘒ⊛
․ᓽࠍᲧセߔࠆ⸘↹ߦߟ޿ߡ߽⼏⺰ߒߚ‫ޕ‬
B-43 ࠨ࡞ߩ൬޿ߦኻߔࠆవᄤ⊛ߥᕟᔺ෻ᔕߩ⸃ᨆ
ዊᣧᎹ઎ሶ㧘ዊᣧᎹ㜞㧘દᣧဈᥓሶ㧘ㄞశ⟵㧔ᄢ㒋ࡃࠗࠝ࡮␹⚻ᯏ⢻ቇ㧕 ᚲౝኻᔕ⠪㧦ਛ᧛స᮸
⑳ߚߜߪࡑ࠙ࠬߦవᄤ⊛ߥᕟᔺ෻ᔕࠍ⺃⊒ߔࠆ༦ⷡ␹⚻࿁〝ߩᯏ⢻ߦ⌕⋡ߒ‫ޔ‬ᣢ⍮ߩ൬޿ಽሶࠃࠅᒝജߦవᄤ⊛
ߥᕟᔺ෻ᔕࠍ⺃⊒ߔࠆੱᎿ‛↱᧪ߩ൬޿ಽሶࠍ⊒⷗ߒߚ‫ߩߎޕ‬൬޿ಽሶࠍ↪޿ߡవᄤ⊛ߥᕟᔺߣᓟᄤ⊛ߥᕟᔺߣߢ
ߪ⇣ߥࠆ↢ℂᔕ╵ࠍ઻߁ߎߣࠍೋ߼ߡ⊒⷗ߒߚ‫ߦࠬ࠙ࡑޔߪߢ↹⸘ᧄޕ‬ኻߒߡవᄤ⊛ߥᕟᔺ෻ᔕࠍ⺃⊒ߔࠆ൬޿ಽ
㧙㧙
ሶ߿ᕟᔺࠍ✭๺ߔࠆ൬޿ಽሶߩലᨐ߇ࠨ࡞ߦኻߒߡ߽଻ሽߐࠇߡ޿ࠆน⢻ᕈࠍᬌ⸽ߔࠆߎߣࠍ⋡⊛ߣߒߚ‫ᧄޕ‬ᐕᐲ
ߪ൬޿ಽሶߩൻቇวᚑߣࡑ࠙ࠬࠍ↪޿ߚࠬࠢ࡝࡯࠾ࡦࠣࠍㅴ߼‫ޔ‬వᄤ⊛ߥᕟᔺᖱേࠍ⺃⊒ߔࠆ൬޿ಽሶߩ࡟ࡄ࡯࠻
࡝࡯ࠍჇടߐߖߚ‫ޕ‬൬޿ಽሶߩലᨐߪേ‛⒳ߦࠃߞߡ⇣ߥࠆߩߢ‫ߦ࡞ࠨޔ‬ኻߒߡᦨㆡߥ൬޿ಽሶࠍ⊒⷗ߢ߈ࠆน⢻
ᕈ߇㜞ߊߥߞߚ‫ޔߚ߹ޕ‬൬޿ಽሶߦࠃࠆవᄤ⊛ߥᕟᔺࠍ✭๺ߔࠆ૞↪ߩ޽ࠆ൬޿ಽሶࠍ⊒⷗ߒߚ‫ߩࠄࠇߎޕ‬൬޿ಽ
ሶߩࠨ࡞ߦኻߔࠆലᨐࠍᬌ⸽ߔࠆ‫ޕ‬వᄤ⊛ߣᓟᄤ⊛ߥᕟᔺߦ઻߁↢ℂᜰᮡߩᄌൻߩ⸃ᨆࠍㅴ߼ߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬వᄤ
⊛ߥᕟᔺߢߩߺ૕⴫㕙᷷ᐲߣ૕ᷓㇱ᷷ᐲߩหᤨૐਅ߿ᔃᜉᢙߩᕆᷫ߇⺃⊒ߐࠇࠆߎߣࠍ⸃᣿ߒߚ‫ޔߚ߹ޕ‬వᄤ⊛ߥ
ᕟᔺߢߪ⣖ᵄߥߤߩ↢ℂᜰᮡߦ߽ᄌൻ߇⷗ࠄࠇࠆߎߣࠍ⸃᣿ߒߚ‫↢ߩࠄࠇߎޕ‬ℂᜰᮡߩਛߢࠨ࡞ߦ߅޿ߡ⸘᷹ߒ߿
ߔ޿ᜰᮡࠍㆬᛯߒߡ⸘᷹ߔࠆ‫ޕ‬
B-44 ࡀࡄ࡯࡞ࡅࡑ࡜ࡗߦ߅ߌࠆ㔤㐳㘃ߩ࿾ၞಽൻߦ㑐ߔࠆ⎇ⓥ
ࡓࠤࠪࡘ K ࠴ࡖ࡝࠮㧔࠻࡝ࡉࡃࡦᄢ࡮േ‛ቇ⑼㧕 ᚲౝኻᔕ⠪㧦Ꮉᧄ⧐
ࡀࡄ࡯࡞ߦߪૐ࿾߆ࠄጊጪ࿾Ꮺ߹ߢߩߐ߹ߑ߹ߥⅣႺߦࠕ࠶ࠨࡓࡕࡦࠠ࡯㧔Macaca assamensis㧕‫࡞ࠩࠥࠞࠕޔ‬㧔M.
mulatta㧕‫࡞࡯ࠣࡦ࡜ࠗ࡟ࠣޔ‬㧔Semnopithecus entellus㧦೎ฬࡂ࠿ࡑࡦ࡜ࡦࠣ࡯࡞㧕߇ᐢߊಽᏓߔࠆ‫ߩߘޔߒ߆ߒޕ‬ಽ
㘃‫ޔ‬ಽᏓ‫ޔ‬࿾ၞಽൻߦ㑐ߔࠆ૕♽⊛ߥㆮવቇ⺞ᩏߪ⊝ήߦ╬ߒ޿‫⎇ᧄޔߢߎߘޕ‬ⓥߢߪࠗࡦ࠼᧲ർㇱࡅࡑ࡜ࡗጊጪ
࿾Ꮺߢႎ๔ߐࠇߚᣂ⒳ࠕ࡞࠽࡯࠴ࡖ࡞ࡑࠞࠢ M. munzala ߣߩ㑐ଥ߇໧㗴ⷞߐࠇࠆࠕ࠶ࠨࡓࡕࡦࠠ࡯ߣ‫ޔ‬੝⒳ಽ㘃
ߢ⺰੎߇⛯ߊࠣ࡟ࠗ࡜ࡦࠣ࡯࡞ࠍኻ⽎ߦ‫⹜♮ޔ‬ᢱ߆ࠄߩ DNA ಽᨆᴺࠍᬌ⸛ߒߚ‫ޕ‬
♮ಽᨆᴺࠍ⹜㛎ߔࠆߚ߼‫࡞࡯ࡄࡀޔ‬࿖ౝߩ↢ᕷ࿾ 3 ࿾ὐߢណขߒߚࠕ࠶ࠨࡓࡕࡦࠠ࡯ߩ⹜ᢱ 11 ᬌ૕ࠍ᧚ᢱߦ‫ޔ‬
DNA ᛽಴ᴺ‫ޔ‬mtDNA Ⴎၮ㈩೉ߩ⸃⺒ᴺࠍᬌ⸛ߒߚ‫⹜♮ߩ࡞࡯ࠣࡦ࡜ࠗ࡟ࠣޔߚ߹ޕ‬ᢱߦߟ޿ߡ߽ห᭽ߩᣇᴺߢಽ
ᨆ߇น⢻߆⹜㛎ߒߚ‫⚿ߩߎޕ‬ᨐ‫ߪߢ࡯ࠠࡦࡕࡓࠨ࠶ࠕޔ‬቟ቯߒߡ㈩೉࠺࡯࠲ࠍᓧࠆታ㛎᧦ઙ߇⏕┙ߢ߈‫ޔ‬㕖ࠦ࡯࠼
㗔ၞߩ╙㧝นᄌၞ⚂ 580 Ⴎၮ‫ ߦ߮ࠄߥޔ‬16S rRNA ࠦ࡯࠼㗔ၞߩ⚂ 400 Ⴎၮ߇⸃⺒ߢ߈ߚ‫ઁޔߢߎߘޕ‬࿾ၞߩࠕ࠶
ࠨࡓࡕࡦࠠ࡯߿ sinica ⒳ࠣ࡞࡯ࡊߦዻߔࠆ೎⒳ߣᲧセߒ‫੍ޔ‬஻⊛ߦಽሶ♽⛔ࠍ⸃ᨆߒߚ‫⚿ߩߎޕ‬ᨐ‫⺞ޔ‬ᩏߒߚࡀࡄ
࡯࡞ߩ⹜ᢱߪࡉ࡯࠲ࡦߩࠕ࠶ࠨࡓࡕࡦࠠ࡯㧔M. a. pelops㧕߿ࠕ࡞࠽࡯࠴ࡖ࡞ࡑࠞࠢߣࠢ࡜ࠬ࠲࡯ࠍ૞ࠅ‫ࠪ࠼ࡦࠗޔ‬
࠽ࠍਛᔃߦಽᏓߔࠆࠕ࠶ࠨࡓࡕࡦࠠ࡯ߩ೎੝⒳ M. a. assamensis ߣ඙೎ߢ߈ߚ‫৻ޕ‬ᣇ‫ߩ࡞࡯ࠣࡦ࡜ࠗ࡟ࠣޔ‬ಽᨆߢ
ߪ‫ޔ‬PCR ߩ㓙ߩࡊ࡜ࠗࡑ࡯߇ਇㆡߢᡷ⦟ߩᔅⷐ߇್᣿ߒߚ‫ޕ‬
B-45 RNA ࠍၮὐߣߒߚ㔤㐳㘃ߩࠛࡇࠫࠚࡀ࠹ࠖࠢࠬ
੹᧛ᜏ਽㧔੩ㇺᄢ࡮㒮࡮ℂ㧕 ᚲౝኻᔕ⠪㧦ᄢ⍹㜞↢
ᧄ⺖㗴ߪ‫੃߶ޔ‬㘃⣖ߩࠛࡇࠥࡁࡓᒻᚑߦ㑐ࠊࠆ non-coding RNA (ncRNA) ೙ᓮࡔࠞ࠾࠭ࡓߣߘߩ⒳㑆ᄙ᭽ᕈࠍ᣿
ࠄ߆ߦߔࠆߎߣࠍ⋡⊛ߣߒߡ޿ࠆ‫ᧄޕ‬ᐕᐲߪએਅߩ 4 ὐߦߟ޿ߡ᣿ࠄ߆ߦߒߚ㧦1㧕ࡅ࠻‫ࠢࠞࡑޔ࡯ࠫࡦࡄࡦ࠴ޔ‬
ࠩ࡞‫ߩ߳࡯࠲࡯ࡕࡠࡊࠆߌ߅ߦ࠲ࡉޔࠬ࠙ࡑޔ‬னㆮવሶᝌ౉㗫ᐲ߇⒳㑆ߢ⇣ߥࠆ㧔ࡃࠗࠝࠗࡦࡈࠜࡑ࠹ࠖࠢࠬ⸃
ᨆ㧕‫ޔ‬2㧕࿕ቯߒߚᝌ౉㈩೉߆ࠄߪ RNA ォ౮߇ࠦ࡯࠼ㆮવሶߦኻߒࠕࡦ࠴࠮ࡦࠬᣇะߦ⿠ߎࠆ㧔೨ᐕᐲ߹ߢߩࡊࡠ
ࡕ࡯࠲࡯ࠕ࡟ࠗߩ⚿ᨐࠍࠗ࡞ࡒ࠽␠ᰴ਎ઍࠪ࡯ࠢࠛࡦࠬ⸃ᨆߦࠃࠅ♖ᐲࠍ㜞߼ߡᬌቯ㧕‫ޔ‬3㧕ᝌ౉㈩೉⥄૕ߪࡊࡠ
ࡕ࡯࠲࡯/ࠛࡦࡂࡦࠨ࡯ߣߒߡ‫ޔ‬DNA ࡔ࠴࡞ൻߒߚᝌ౉㈩೉ߪࠨࠗ࡟ࡦࠨ࡯ߣߒߡᯏ⢻ߒ߁ࠆ㧔ฦ⒳േ‛⚦⢩࡟ࡐ
࡯࠲࡯ࠕ࠶࠮ࠗ㧕‫ޔ‬4㧕ᝌ౉㈩೉↱᧪ promoter-associated ncRNA (pancRNA) ߣ DNA ࠣ࡝ࠦࠪ࡜࡯࠯ߩᒝ೙⊒⃻ߦࠃ
ࠅ‫ޔ‬ᝌ౉㈩೉ࠍ㈩೉․⇣⊛ߦ⣕ࡔ࠴࡞ൻ/ㆮવሶࠝࡦߦߢ߈ࠆ㧔⹜㛎▤ౝࡔ࠴࡞ൻࠕ࠶࠮ࠗ㧕‫ޕ‬એ਄ࠃࠅ‫ޔ‬னㆮવ
ሶ↱᧪㈩೉߇ㆮવሶ೙ᓮ㗔ၞߦ⒳․⇣⊛ߦᝌ౉ߐࠇࠆߎߣߢ‫ޔ‬⒳․⇣⊛ pancRNA ߇₪ᓧߐࠇ‫ޔ‬㈩೉․⇣⊛ DNA ࡔ
࠴࡞ൻࡄ࠲࡯ࡦᒻᚑࠍ੺ߒߡ‫ޔ‬⒳․⇣⊛ㆮવሶ⊒⃻ࠬࠗ࠶࠴ߩ₪ᓧߦ⥋ࠆߎߣ߇⠨߃ࠄࠇߚ‫ޕ‬
㧨ቇળ⊒⴫㧪
1) Imamura T, et al. (2011) Species-specific promoter-associated noncoding RNA mediates DNA demethylation in
macaques. Society for Molecular Biology and Evolution 2011 (2011/07/27, Kyoto)
2) ਄ဈ዁ᒄ㧘߶߆ (2011) ࡑࠞࠢࠩ࡞ߦ߅ߌࠆனㆮવሶ↱᧪ promoter-associated noncoding RNA (pancRNA) ߦࠃ
ࠆ⒳․⇣⊛ォ౮ᵴᕈൻ. ╙ 104 ࿁ᣣᧄ❥ᱺ↢‛ቇળᄢળ (2011/09/16, ⋓ጟᏒ)
3) ੹᧛ᜏ਽㧘߶߆ (2011) ࡑࠞࠢࠩ࡞ߦ߅ߌࠆனㆮવሶ↱᧪ promoter-associated noncoding RNA (pancRNA) ߦࠃ
ࠆ⒳․⇣⊛ォ౮ᵴᕈൻ. ╙ 152 ࿁ᣣᧄ₞කቇળᄢળ (2011/09/20, ႓Ꮢ)
4) Uesaka M, et al. (2011) Species-specific pseudogene insertions generate cis-acting RNA for promoter demethylation in
the macaque. ╙ 34 ࿁ᣣᧄಽሶ↢‛ቇળᐕળ (2011/12/16, ᮮᵿᏒ)
B-46 ࠾ࡎࡦࠩ࡞㓸࿅߅ࠃ߮ࠕࠞࠥࠩ࡞㓸࿅߆ࠄߩㆇേቇ⊛࠺࡯࠲ߩ෼㓸
ᣣ᥵ᵏ↵㧔ᄢ㒋ᄢ࡮㒮࡮ੱ㑆⑼ቇ㧕 ᚲౝኻᔕ⠪㧦ᐔፒ㍈⍫
⎇ⓥߩ⋡⊛ߪ㔤㐳㘃ߩ྾⿷ᱠⴕᤨߦ߅ߌࠆࡈ࠶࠻ࡈࠜ࡯࡞࡮ࡄ࠲࡯ࡦ㧔1 ᱠⴕ๟ᦼߩਛߢ‫ޔ‬Ꮐฝߩ೨⢇ߣᓟ⢇߇
ᡰᜬၮ૕ߦ⌕ߊ㗅⇟㧕ߦᓇ㗀ߔࠆⷐ࿃ࠍᛠីߔࠆߎߣߢ޽ߞߚ‫ޔߢ⊛⋡ߩߎޕ‬੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲߢ㓸࿅㘺⢒ߐ
ࠇߡ޿ࠆ࠾ࡎࡦࠩ࡞ߣࠕࠞࠥࠩ࡞ࠍኻ⽎ߦ‫ޔ‬႐ᄖߩⷰኤบ߆ࠄ 2 ⒳ߩ྾⿷ᱠⴕࠍࡆ࠺ࠝ᠟ᓇߒ‫ࠜࡈ࠻࠶ࡈߩߘޔ‬
࡯࡞࡮ࡄ࠲࡯ࡦࠍ⺞ߴࠆߣߣ߽ߦ‫ޔ‬ᡰᜬၮ૕ߩᒻ⁁߿௑ᢳ‫ޔ‬り૕ࠨࠗ࠭‫ઁޔ‬୘૕ߩㆇ៝ߩ᦭ήߣㆇ៝ᣇᴺߣ޿ߞ
ߚઁߩᄌᢙߣߩ㑐ㅪࠍಽᨆߒߚ‫ޕ‬᡼㘺႐ߦࠃߞߡ႐ౝߩ᭴ㅧ‛߇⇣ߥߞߚߎߣ߽޽ࠅ‫ޔ‬᠟ᓇߒߚᱠⴕߩᄙߊ߇࠾
ࡎࡦࠩ࡞ߪ࿾਄ߢߩ߽ߩ‫ߪ࡞ࠩࠥࠞࠕޔ‬᮸਄ⅣႺࠍᮨߒߚ᭴ㅧ‛਄ߢߩ߽ߩߢ޽ߞߚ‫ޕ‬వⴕ⎇ⓥߩߣ߅ࠅ‫ޔ‬2 ⒳ߩ
ࡈ࠶࠻ࡈࠜ࡯࡞࡮ࡄ࠲࡯ࡦߪ‫ޔ‬ᓟ⢇ߩᰴߦኻ஥ߩ೨⢇߇ᡰᜬၮ૕ߦ⌕ߊ೨ᣇ੤෷ဳ߇߶ߣࠎߤߛߞߚ‫ޕ‬ᓟ⢇ߩᰴ
ߦห஥ߩ೨⢇߇⌕ߊᓟᣇ੤෷ဳߩࡄ࠲࡯ࡦ߇ 1 ᱠⴕ๟ᦼߟߠ޿ߚ଀ߪ੹࿁ߪ⷗ࠄࠇߥ߆ߞߚ߇‫ޔ‬ਅࠅ௑ᢳߩ⒖േ
ᤨ‫ޔ‬ᲑᏅࠍਸ਼ࠅߎ߃ࠆᤨ‫ߪߚ߹ޔ‬⒖േࠍᱛ߼ࠆ⋥೨ߥߤߦߎߩࡄ࠲࡯ࡦࠍㇱಽ⊛ߦ߰ߊ߻ᱠⴕ߇ⷰኤߐࠇࠆߎߣ
㧙㧙
߇㑆‫ޕߚߞ޽ޘ‬
B-47 ㊁⨲ߩ⧰๧࡮ᷦ๧ᚑಽ฽㊂ߣ࠾ࡎࡦࠩ࡞ߩ༵ᅢᕈߣߩ㑐ㅪᕈߦߟ޿ߡ
ዊ᎑㆏ਯ㧔Ꮺᐢ⇓↥ᄢ࡮㘩⑼㧕㧘 ᦭ንᐘᴦ㧔Ꮺᐢ⇓↥ᄢ㧕 ᚲౝኻᔕ⠪㧦㋈ᧁ᮸ℂ
⧰๧࡮ᷦ๧ߩᮡຠࠍ↪޿ߚ៨㘩⹜㛎ࠍታᣉߒߚ‫ޕ‬ᮡຠߦߪ‫ߩ࠻ࡅޔ‬๧ⷡታ㛎ߦ߽ၮḰ⹜⮎ߣߒߡ૶↪ߔࠆࠠ࠾࡯
ࡀ෸߮߅⨥ߩᚑಽߢ޽ࠆࠞ࠹ࠠࡦߩ 2 ⒳㘃ࠍ↪޿ߚ‫ޕ‬೨⠪ߪࠕ࡞ࠞࡠࠗ࠼㘃‫ޔ‬ᓟ⠪ߪࡈ࡜ࡏࡁࠗ࠼㘃㧔ࡐ࡝ࡈࠚࡁ
࡯࡞㧕
ߦዻߔࠆ‫ޕ‬
ࡅ࠻ߩ⹜㛎ߢ㑣୯ߣߒߡ⏕⹺ߐࠇߡ޿ࠆỚᐲ㧧3mg/100ml ࠍ⠨ᘦߒߡ‫ޔ‬
࿕ᒻ㘺ᢱߦᦨ⚳Ớᐲ 1.875g‫ޔ‬
3.75g‫ޔ‬7.5g ߩ 3 Ბ㓏ߢᷝടߒߚੇ῎࿕ᒻ㘺ᢱࠍ 10 ୘ߠߟਈ߃ߡ༵ᅢᕈ⹜㛎㧔ඦ೨ 10:00 ߣඦᓟ 2:00 ߩ 2 ࿁㧕ታ
㛎ߒߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬3‫ޔ‬4 ᚽߩ⧯޿ࠨ࡞ߢߪ 1 㗡߇ඦ೨ߩߺ 2 ୘ᱷߒߚ߇‫ޔ‬ඦᓟߩታ㛎ߢߪỚᐲ߇㜞ߊߡ߽ߔߴߡ
㘩ߴࠆߎߣ‫ޔ‬ᐕነࠅߩࠨ࡞ 1 㗡ߪᦨૐỚᐲߢ 3 ୘‫ޔ‬ᰴߩ 3.75g ߢߪ 4 ୘ᱷߒߚ߇‫ߦࠄߐޔ‬Ớ޿Ớᐲߩ㘺ᢱࠍඦᓟ
ߦਈ߃ࠆߣߔߴߡ㘩ߴࠆߎߣ߆ࠄ‫ޔ‬ඦ೨ߩ 10 ୘ߢߪ㙄ߩ⛘ኻ㊂߇ਇ⿷ߒߡᚒᘟߢ߈ߕߦ‫ޔ‬ඦᓟߩታ㛎ߦ༵ᅢᕈߩ
೙ᓮ߇߆߆ࠄߥ޿ߣផኤߒߚ‫ޔߪߡ޿ߟߦࡦࠠ࠹ࠞޔߚ߹ޕ‬282.5g‫ޔ‬706.3g‫ޔ‬1412.5g ߩ 3 Ბ㓏ߢᷝടߒߡห᭽
ߩታ㛎ࠍታᣉߒߚ‫⚿ߩߘޕ‬ᨐ‫ߡߴߔߪ࡞ࠨ޿⧯ޔ‬㘩ߴߚ߇‫ޔ‬ᐕነࠅߩࠨ࡞ 1 㗡ߪ‫ᦨޔ‬ૐỚᐲߢߐ߃ 9 ୘㘩ߴߥ߆ߞ
ߚ‫ޔߒ߆ߒޕ‬ඦᓟߦߪ 1 ୘ᱷߒߚߛߌߢ޽ࠅ‫ޔ‬1412.5g ߩ߽ߩߪ 3 ୘ᱷߒߚ‫ޔߢߎߘޕ‬ඦ೨ߩታ㛎ᓟߦㅢᏱߩ 1/3
⒟ᐲㅢᏱߩ㙄ࠍਈ߃‫ޔ‬ඦᓟߦ 1412.5g ਈ߃ߚታ㛎ߢߪ‫ ࡞ࠨ޿⧯ޔ‬1 㗡߇ 4 ୘ᱷߒߚ߇‫ޔ‬ᐕነࠅߩࠨ࡞ 1 㗡߇ 1 ୘
ᱷߒߚ‫ޕ‬ඦ೨ߩታ㛎ߩᓟ‫ޔ‬ㅢᏱ⒟ᐲ߹ߢ㙄ࠍਈ߃ࠆߎߣߢ‫ޔ‬ඦᓟߩ๧ⷡታ㛎ߩ♖ᐲ߇⏕଻ߢ߈ࠆߣផኤߒߚ‫੹ޕ‬ᓟ‫ޔ‬
ታ㛎ⷰኤᴺߩᡷ⦟ᴺࠍ↪޿ߡ‫ޔ‬ฦỚᐲࠍߐࠄߦ㜞ߊߒߡ޿߈‫ߩࠇߙࠇߘޔ‬ᜎุ㑣୯ࠍ᳞߼ࠆ੍ቯߢ޽ࠆ‫ޕ‬
B-48 㔤㐳㘃ߩฦ⒳⚵❱ߩട㦂ᄌൻ
᧲⿥㧔ᄹ⦟⋵කᄢ࡮ක࡮⸃೬ቇ㧕 ᚲౝኻᔕ⠪㧦ᄢ⍹㜞↢
ട㦂ߦ઻߁᳇▤エ㛽ߩర⚛฽㊂ߩᄌൻࠍ᣿ࠄ߆ߦߔࠆߚ߼ߦ‫ߩ࡞ࠨޔ‬᳇▤エ㛽ߩర⚛฽㊂ߩട㦂ᄌൻࠍ⺞ߴߡ‫ޔ‬
ࡅ࠻ߩ᳇▤エ㛽ߣᲧセ⎇ⓥࠍⴕߞߚ‫ ࡞ࠩࠥࠞࠕߪ࡞ࠨߚ޿↪ޕ‬10 㗡‫ ࡞ࠩࡦࡎ࠾ޔ‬1 㗡‫ ࡞ࠩࠗࠢ࠾ࠞޔ‬3 㗡‫ޔ‬ᐕ㦂
ߪ 1 ᦬߆ࠄ 27 ᱦ‫ޔ‬㓶㓽ߪ㓶 9 㗡ߣ㓽 5 㗡ߢ޽ࠆ‫ࠅࠃ࡞ࠨޕ‬᳇▤エ㛽ࠍណขߒ‫ߣ㉄⎣ޔ‬ㆊႮ⚛㉄ࠍട߃ߡ‫ޔ‬ടᾲߒ
ߡἯൻߒ‫ޔ‬ర⚛฽㊂ࠍ㜞๟ᵄࡊ࡜࠭ࡑ⊒శಽᨆⵝ⟎㧔ICPS-7510‫ޔ‬ፉᵤ⵾㧕ߢಽᨆߒ‫ޔ‬ᰴߩࠃ߁ߥ⚿ᨐ߇ᓧࠄࠇߚ‫ޕ‬
Ԙࠨ࡞ߣࡅ࠻ߩ᳇▤エ㛽ߩࠞ࡞ࠪ࠙ࡓ฽㊂ߪ 10mg/g એ਄ߢ‫⫾ࡓ࠙ࠪ࡞ࠞޔ‬Ⓧ߇↢ߓ߿ߔ޿エ㛽ߢ޽ࠆߎߣ߇ಽ
߆ߞߚ‫ޕ‬
ԙࠨ࡞ߣࡅ࠻ߩ᳇▤エ㛽ߩࠞ࡞ࠪ࠙ࡓ‫ޔ‬ῂ฽㊂ߪᐕ㦂ߣߣ߽ߦ᦭ᗧߦჇടߒߚ‫ޕ‬
Ԛࠨ࡞ߩ᳇▤エ㛽ߩࠞ࡞ࠪ࠙ࡓ฽㊂ߪ 7 ᱦએ਄ߦߥࠆߣ㗼⪺ߦჇടߒߚ‫ߩ࠻ࡅޕ‬᳇▤エ㛽ߩࠞ࡞ࠪ࠙ࡓ฽㊂ߪ
80 ઍߦ⪺᣿ߦჇടߒߚ‫⚿ߩࠄࠇߎޕ‬ᨐ߆ࠄࠨ࡞ߣࡅ࠻ߩ᳇▤エ㛽ߦ߅޿ߡ৻ቯᐕ㦂ࠍ⿥߃ࠆߣ⍹Ἧൻ߇ᆎ߹
ࠆߎߣ߇ಽ߆ߞߚ‫ޕ‬
㧨ቇળ⊒⴫㧪
᧲⿥㧘ᄢ⍹㜞↢ (2011)ࡅ࠻ߣࠨ࡞ߩ༄㗡⬄エ㛽ߦ߅ߌࠆర⚛⫾Ⓧߩ․ᓽ㧚╙ 116 ࿁ᣣᧄ⸃೬ቇળ✚ળ࡮ో࿖ቇ
ⴚ㓸ળ㧚
B-49 㔤㐳㘃ߩㆇേⷞᯏ⢻ߦ㑐ߔࠆᲧセ⹺⍮⊒㆐⑼ቇ⊛ᬌ⸛
⊕੗ㅀ㧔ᣂẟᄢ࡮ੱᢥቇㇱ㧕 ᚲౝኻᔕ⠪㧦෹᳗㓷Ꮖ
ㆇേⷞᯏ⢻ߪⷞⷡࠍᜬߟേ‛ߦߣߞߡᦨ߽ၮ␆⊛߆ߟ㊀ⷐߥⷞᯏ⢻ߢ޽ࠅ‫ޔ‬᭽‫ߥޘ‬ㆡᔕ⊛ⴕὑߣኒធߥ߆߆ࠊࠅ
ࠍᜬߟ‫ޕ‬
ࡅ࠻߿ߘߩઁߩ㔤㐳㘃ߩㆇേⷞᯏ⢻ߦߟ޿ߡߪ‫ޔ‬ታ㛎ᔃℂቇ߿␹⚻↢ℂቇߥߤߩ⻉ಽ㊁ߦ߅޿ߡ♖ജ⊛ߦ⎇ⓥߐ
ࠇߡ߈ߚ‫ⷡⷞߩ࠻ࡅ߇ߤߥ࡞ࠩࠢࠞࡑޔࠄ߇ߥߒ߆ߒޕ‬⣖ߩᯏ⢻ࠍᬌ⸛ߔࠆ㓙ߩౖဳ⊛ߥࡕ࠺࡞േ‛ߣߐࠇࠆࠃ߁
ߦ‫࠻ࡅߣ࠻ࡅޔߪߦ⊛⥸৻ޔ‬એᄖߩ㔤㐳㘃⒳ߩㆇേⷞᯏ⢻߇㘃ૃߒߡ޿ࠆߎߣࠍ೨ឭߣߒߚ⎇ⓥ߇ᄙ൓ࠍභ߼ࠆߣ
޿߃ࠆ‫৻ߩߘޕ‬ᣇߢ‫ࠆߥ⇣ޔ‬㔤㐳㘃⒳㑆ߢㆇേⷞᯏ⢻ߩᏅ⇣ߦߟ޿ߡὶὐ߇ᒰߡࠄࠇࠆߎߣߪᲧセ⊛⒘ߢ޽ࠆ‫ߎޕ‬
ߩࠃ߁ߥ⃻⁁ߦ㐓ߺ‫⎇ᧄޔ‬ⓥ⸘↹ߪ⇣ߥࠆ㔤㐳㘃⒳ߩ㑆ߦሽ࿷ߔࠆㆇേⷞᯏ⢻ߩ౒ㅢὐߦߟ޿ߡត⚝ߔࠆߩߺߥࠄ
ߕ‫⋧ߥ߁ࠃߩߤޔ‬㆑ὐ߇ሽ࿷ߔࠆ߆ߦߟ޿ߡ߽ᬌ⸛ߔࠆߎߣࠍ⋡⊛ߣߒߚ‫ޕ‬
ౕ૕⊛ߦߪ‫ޔ‬᜛ᄢ࡮❗ዊ߿Ꮐฝᣇะߩ࿁ォߥߤߩ⋧ኻㆇേࡄ࠲ࡦߩᬌ಴ᗵᐲߦߟ޿ߡ‫ࠍ࡯ࠫࡦࡄࡦ࠴ޔ‬ኻ⽎ߦ᷹
ቯߔࠆߎߣࠍ⹜ߺߚ‫ޔߪߢ࠻ࡅޕ‬᜛ᄢ࡮❗ዊߥߤߩ᡼኿⁁ߩㆇേࡄ࠲ࡦߦኻߔࠆᬌ಴ᗵᐲ߇‫ޔ‬࿁ォㆇേߦኻߔࠆᗵ
ᐲࠃࠅ߽㜞޿ߎߣ߇ߒ߫ߒ߫ႎ๔ߐࠇࠆ‫ߚߒ߁ߎޕ‬௑ะ߇࠴ࡦࡄࡦࠫ࡯ߦ߅޿ߡ߽ⷰኤߐࠇࠆ߆ุ߆ࠍᔃℂ‛ℂቇ
⊛ታ㛎ߦࠃߞߡᬌ⸛ߒߚ‫ޕ‬ታ㛎ߢߪߘࠇߙࠇ 150 ୘ߩశὐߦࠃߞߡ᭴ᚑߐࠇࠆ⋧ኻㆇേࡄ࠲ࡦߣ࡜ࡦ࠳ࡓㆇേࡄ࠲
ࡦࠍ࠲࠶࠴ࡄࡀ࡞ᑼߩࠦࡦࡇࡘ࡯࠲ࠬࠢ࡝࡯ࡦߦኻឭ␜ߒߚ‫ޕ‬ታ㛎⺖㗴ߪࠬࠢ࡝࡯ࡦ਄ߦ⃻ࠇࠆ⋧ኻㆇേࡄ࠲ࡦߣ
࡜ࡦ࠳ࡓㆇേࡄ࠲ࡦߩ߁ߜ‫ޔ‬೨⠪ࠍᱜ⏕ߦ࠲࠶ࡇࡦࠣߔࠆߎߣߢ޽ߞߚ‫ޕ‬ฦ⹜ⴕߦ߅ߌࠆ⺖㗴ߩᱜ⺋ߦᔕߓߡ‫⋧ޔ‬
ኻㆇേࡄ࠲ࡦߩ᣿⍎ߐ߇ᄌേߒ‫߁઻ߦࠇߘޔ‬ᱜ╵‫♽ߩ╵⺋ޔ‬೉߆ࠄߘࠇߙࠇߩ⋧ኻㆇേߩᬌ಴㑣ࠍ᳞߼ߚ㧔ᄌᒻ਄
ਅᴺ㧕‫⋧߅ߥޕ‬ኻㆇേࡄ࠲ࡦߩ᣿⍎ᐲߪ‫⋧ޔ‬ኻㆇേࡄ࠲ࡦࠍ᭴ᚑߔࠆ࠼࠶࠻⟲ߩ߁ߜ‫ޔ‬છᗧߩഀวߩ࠼࠶࠻ߩㆇേ
゠㆏ࠍ࡜ࡦ࠳ࡓߥᣇะߦᄌൻߐߖࠆߎߣߦࠃߞߡᠲ૞ߐࠇߚ‫ޕ‬ታ㛎ߩ⚿ᨐ‫⋧ߩࠇߕ޿ޔ‬ኻㆇേࡄ࠲ࡦߦኻߔࠆᬌ಴
ᗵᐲ߽‫ߡ޿߅ߦ࠻ࡅޔ‬ႎ๔ߐࠇࠆ৻⥸⊛ߥᗵᐲࠍᄢ߈ߊਅ࿁ߞߚ‫⋧ࠆߥ⇣ޔߚ߹ޕ‬ኻㆇേࡄ࠲ࡦߦኻߔࠆᗵᐲ㑆ߦ
᦭ᗧߥᏅߪߥ߆ߞߚ‫ޕ‬
ߎ߁ߒߚ⚿ᨐߪ‫⋧ޔ‬ኻㆇേߩࠃ߁ߥᲧセ⊛ⶄ㔀ߥ‫ߟ߆ޔ‬ᄢၞ⊛ߥⷞㆇേࡄ࠲ࡦߩಣℂߦ߅޿ߡ‫ࡦࡄࡦ࠴ߣ࠻ࡅޔ‬
ࠫ࡯ߩ㑆ߦᄢ߈ߥᏅ⇣߇ሽ࿷ߔࠆน⢻ᕈࠍ␜ໂߔࠆ‫⺖ޔࠄ߇ߥߒ߆ߒޕ‬㗴ᢎ␜ߩലᨐ߿‫⺖ޔ‬㗴ߦኻߔࠆᘠࠇߩᓇ㗀
ߥߤ߇‫ߥ⊛⥸৻ࠆߌ߅ߦ࠻ࡅޔ‬ᗵᐲߩ㜞ߐߦ⚿߮ߟ޿ߡ޿ࠆน⢻ᕈ߇޽ࠆ‫੹ޔߡߞ߇ߚߒޕ‬ᓟߘߩࠃ߁ߥน⢻ᕈߦ
㧙㧙
ߟ޿ߡታ⸽⊛ߦᬌ⸛ߒߡ޿ߊ૞ᬺ߇ᔅⷐߢ޽ࠈ߁‫ޕ‬
B-50 㔤㐳㘃ߦ߅ߌࠆ㜑㕽ᒻᚑߩ⹏ଔ⎇ⓥ
ਃ਄┨మ㧔ਛㇱቇ㒮ᄢ࡮࡝ࡂࡆ࡝࠹࡯࡚ࠪࡦቇㇱ࡮ℂቇ≮ᴺቇ⑼㧕 ᚲౝኻᔕ⠪㧦ች࿾㊀ᒄ
࠴ࡦࡄࡦࠫ࡯ߩ⣖ߩ⊒㆐ࠍߺࠆ⋡⊛ߢ MRI ߩ T1 ᒝ⺞↹௝ߩ㜞ାภ㗔ၞࠍ⊕⾰ߣ⹏ଔߔࠆ⎇ⓥ߇ⴕࠊࠇߡ޿ࠆ‫ޕ‬
␹⚻✢⛽ߩ߹ࠊࠅߦ޽ࠆ⛘✼‛⾰ߢ޽ࠆ㜑㕽ߦߪ⢽⾰߇ᄙߊ฽߹ࠇ‫ޔ‬MRI ߩ T1 ↹௝ߢߪ㜞ାภߣߒߡ⸥㍳ߐࠇࠆ‫ޕ‬
ߘߩߚ߼‫ޔ‬㜞ାภ㗔ၞߩ⊒㆐ᄌൻߪ㜑㕽ᒻᚑߩ⚻ㆊࠍߺࠆ᦭ജߥᚻᲑߣߐࠇߡ޿ࠆ‫ޔࠄ߇ߥߒ߆ߒޕ‬MRI ߩ㜞ାภ
㗔ၞߣ⚵❱ᮡᧄߢ⹏ଔߒߚ㜑㕽ᒻᚑ߇ߤߩ⒟ᐲኻᔕߔࠆ߆ߪචಽᬌ⸛ߐࠇߡ޿ߥ޿‫ࠞࠞࡑޔߢߎߘޕ‬ዻߩࠨ࡞ߩ⊒
㆐ㆊ⒟ߢ‫ޔ‬MRI ߦࠃࠆ᠟௝ߣ⚵❱ᮡᧄߦࠃࠆ㜑㕽ᒻᚑߩ್ቯࠍหᤨߦⴕ޿‫⋧ߩߘޔ‬㑐ࠍ⹏ଔߔࠆ⎇ⓥࠍⴕߞߚ‫੹ޕ‬
ᐕᐲߪ‫ ࠻࡞࠳ࠕߩ࡞ࠩࠥࠞࠕޔ‬1 㗡㧔16 ᱦ 6 ࡨ᦬㧕ߣ੃ఽ 2 㗡㧔6 ㅳ㦂‫ޔ‬9 ㅳ㦂㧕ߩ⣖ᮡᧄࠍࡈࠔࠬ࠻࡮ࡉ࡞࡯ᨴ
⦡ߒ⊕⾰‫ޔ‬Ἧ⊕⾰㗔ၞߩᲧセࠍⴕ޿‫⾰⊹ޔ‬㗔ၞ߇੃ఽᦼߦᐢ޿ߎߣࠍ⏕⹺ߒߚ‫ߣࠇߎޕ‬ਗⴕߒߡ‫࡯ࠫࡦࡄࡦ࠴ޔ‬⣖
ߩ MRI ⸘᷹ࠍ⛮⛯ߒߚ‫ޕ‬
B-51 ࠾ࡎࡦࠩ࡞ߩ૕ᐙߣਅ⢇ᏪߩႺ⇇㗔ၞߦ߅ߌࠆ⣄㜑␹⚻೨ᨑߩᒻᘒ⊛․ᓽ
ᤨ↰ᐘਯテ㧔ၯ₹ක⑼ᄢ࡮଻ஜක≮ቇㇱ࡮ℂቇ≮ᴺ㧕 ᚲౝኻᔕ⠪㧦Ძ೑ବ㓶
࠾ࡎࡦࠩ࡞ߩ૕ᐙߣਅ⢇ᏪߩႺ⇇㗔ၞߦ߅ߌࠆ⣄㜑␹⚻೨ᨑߩᒻᘒ⊛․ᓽࠍ᣿ࠄ߆ߦߔࠆ⋡⊛ߢ‫ޔ‬⣶␹⚻ฌߣਅ
ㇱ⡻㑆␹⚻ߩⷰኤࠍⴕߞߚ‫ޔߦ․ޕ‬⣶␹⚻ฌߣ઄㛽␹⚻ฌߩႺ⇇ߢ޽ࠆಽጘ␹⚻㧔Jhering㧕ߩ⿠ᆎಽ▵߿⣶␹⚻ฌ
↱᧪ߩฦ␹⚻ߩ⿠ᆎಽ▵‫ޔ〝⚻ⴕ⿛ޔ‬ಽᏓߦᵈᗧߒߡⷰኤߒߚ‫ޔߚ߹ޕ‬ਅㇱ⡻㛽ߩᒻᘒ߽޽ࠊߖߡⷰኤߒߚ‫ޕ‬
ಽጘ␹⚻ࠍ⿠ᆎಽ▵ߩ㜞ߐ߆ࠄ L5 ⟲‫ޔ‬L5+L6 ⟲‫ޔ‬L6 ⟲ߩ 3 ⟲ߦಽߌߚ‫ޕ‬ಽጘ␹⚻⿠ᆎಽ▵ߪ‫ޔ‬਄ᣇ߆ࠄ L5 ⟲‫ޔ‬
L5+L6 ⟲‫ޔ‬L6 ⟲ߩ㗅ߢየ஥߳࠭࡟ࠆߣ⸒߃ࠆ‫ޕ‬
ᦨਅ┵ߩ⡻㑆␹⚻ᄖ஥⊹ᨑ㧔RcL㧕ߩ⿠ᆎಽ▵ߪ L5 ⟲ߢ L2‫ޔ‬L5+L6 ⟲ߢ L2+L3‫ޔ‬L6 ⟲ߢ L3 ߢ޽ߞߚ‫ᦨޕ‬ਅ┵
ߩᮡḰ⊛ߥ⡻㑆␹⚻೨⊹ᨑ㧔Rcap㧕ߩ⿠ᆎಽ▵ߪ L5 ⟲ߢ L2‫ޔ‬L5+L6 ⟲ߢ L2+L3‫ޔ‬L6 ⟲ߢ L3 ߢ޽ߞߚ‫ޔߚ߹ޕ‬L6
⟲ߦ߅޿ߡߪ╙ 1 ⣶ᬁߩ⡻㛽⓭⿠߇⡻㛽㧔⣶⡻㧕ߣߥߞߡ޿ࠆ଀߽޽ߞߚ‫ޕ‬
એ਄ࠃࠅ‫ޔ‬ಽጘ␹⚻ࠍਛᔃߣߒߚਅ⢇߳ߩ␹⚻ߩ⿠ᆎಽ▵߇የ஥߳ߕࠇࠆߣ‫ޔ‬⢵૕㧔⢷ㇱ㧕ߦ․ᓽ⊛ߥ␹⚻ߢ޽
ࠆ Rcap‫ޔ‬Rcl ߩ⿠ᆎಽ▵߽የ஥߳ߕࠇ‫ߦࠄߐޔ‬የ஥߳ߕࠇࠆߣ⣶⡻߇ᒻᚑߐࠇࠆ㧔⣶ᬁߩ⢷ᬁൻ㧕ߣ⸒߃ࠆ‫ޕ‬
╩⠪ߪ‫ޔ‬⣻ო߆ࠄਅ⢇߳ߩ⒖ⴕ㗔ၞߦ⌕⋡ߒ‫࠻ࡅޔ‬⣶␹⚻ฌ߅ࠃ߮ਅㇱ⡻㑆␹⚻ߩⷰኤࠍⴕߞߡ߈ߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬
ਅ⢇߳ಽᏓߔࠆ␹⚻㧔⣶␹⚻ฌ㧕ߩ⿠ᆎಽ▵(᭴ᚑಽ▵)߇የ஥߳ߕࠇࠆᄌ⇣߇ሽ࿷ߔࠆߎߣ‫ߩߎޔ‬ᄌ⇣ߦߣ߽ߥ޿
ᦨਅ┵ߩ⢵૕㧔⢷⣻ㇱ㧕ߦ․ᓽ⊛ߥ␹⚻ߩ⿠ᆎಽ▵߽የ஥߳ߕࠇࠆߎߣ߇᣿ࠄ߆ߦߥߞߚ‫ߩࠄࠇߎޔߚ߹ޕ‬ᄌ⇣ߦ
઻޿ᦨਅ┵ߩ⡻㛽ߩ㐳ߐߩᑧ㐳߿⡻㛽ߩᢙߩჇട㧔⣶ᬁ⡻㛽⓭⿠ߩ⡻㛽ൻ‫ޔ‬⣶⡻㧕ࠍⷰኤߒߡ޿ࠆ㧔2010‫ޔ‬2009‫ޔ‬
2008㧕
‫ޕ‬
੹࿁ⷰኤߐࠇߚ࠾ࡎࡦࠩ࡞⣶␹⚻ฌߩᒻᘒ⊛․ᓽߪ‫࠻ࡅޔ‬⣶␹⚻ฌߢⷰኤߐࠇߚ․ᓽߣห᭽ߥ߽ߩߢ޽ࠅ‫ߕ޿ޔ‬
ࠇ߽⢵૕㧔⢷ㇱ㧕ߩᑧ㐳ߦ㑐ㅪߒߚᄌ⇣ߢ޽ࠆߣ⠨߃ߚ޿‫ޕ‬
ᧄ⎇ⓥߩᚑᨐߪ╙ 28 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળߦߡ⊒⴫੍ቯߢ޽ࠆ‫ޕ‬
㧨⊒⴫᭎ⷐ㧪
╙ 27 ࿁ᣣᧄ㔤㐳㘃ቇળᄢળ 2011.7.16㨪18
ᐢ㥦₎㘃⣶␹⚻ฌߩⷰኤ ᤨ↰ᐘਯテ㧔ၯ₹ක⑼ᄢ࡮଻ஜක≮㧕
2007㨪2009 ᐕߦࠞ࠾ࠢࠗࠩ࡞‫࡯ࠫࡦࡄࡦ࠴ޔ࡞ࠩࡦࡎ࠾ޔ‬⣶␹⚻ฌߩⷰኤࠍⴕߞߚ‫੹ޕ‬࿁‫ޔ‬ᐢ㥦₎㘃⣶␹⚻ฌ
ߩⷰኤߣߒߡ‫ⷰߩࡦ࡝ࡑ࠲࠹ࠞࠕߣ࡞ࠩࠬ࡝ޔ‬ኤࠍⴕߞߚ‫ޕ‬એਅߦⷰኤ⚿ᨐߩ᭎ⷐࠍ⸥ߔ‫ޕ‬Th13㧦⣻ოߦㅴ౉ߒ
ᄖ஥⊹ᨑ㧔RcL㧕ࠍಽᨑߒ‫ޔ‬஥⣻ოߩౝ⣻ᢳ╭㧔Oi㧕ߣ⣻ᮮ╭㧔Ta㧕ߩ㑆㧔╙ 2-3 ጀ㑆㧕ࠍ⿛ⴕߒ‫ޔ‬⣻⋥╭㕽ߦ౉
ࠆ‫ޕ‬⣻⋥╭㧔R㧕ߩᓟ㕙߆ࠄ╭ᨑࠍਈ߃‫ߡ޿⽾ࠍ╭ޔ‬೨⊹ᨑ㧔Rca㧕ࠍಽᨑߔࠆ‫ߪࠇߎޕ‬⢵૕ߦ․ᓽ⊛ߥᮡḰ⊛ߥ
⡻㑆␹⚻ߩ⚻〝ߣ޿߃ࠆ‫ޕ‬L1㧦⣻ოߦㅴ౉ߒ RcL ࠍಽᨑ‫ޔ‬஥⣻ოߩ╙ 2-3 ጀ㑆ࠍ⿛ⴕߒ‫ޔ‬⣻⋥╭㕽ߦ౉ࠅ‫ޔ‬R ࠍ
⽾޿ߡ Rca ࠍಽᨑߔࠆ‫߽〝⚻ߩߎޕ‬ᮡḰ⊛ߥ⡻㑆␹⚻ߩ⚻〝ߣ޿߃ࠆ‫ޕ‬
L2㧦L3 ߳ߩ੤ㅢᨑࠍಽᨑߒߚᓟ‫ޔ‬⣻ოߦㅴ౉ߒ RcL ࠍಽᨑ‫ߩߘޕ‬ᓟ‫ޔ‬஥⣻ოߩ╙ 2-3 ጀ㑆ࠍ⿛ⴕߒ‫ޔ‬⣻⋥╭㕽
ߦ౉ࠅ‫ޔ‬Rca ࠍಽᨑߔࠆߣ޿߁ᮡḰ⊛ߥ⡻㑆␹⚻ߩ⚻〝ࠍߣࠆ‫ޕ‬L3㧦2 ᨑߦಽᨑߔࠆ‫ޕ‬1 ᨑߪ L2 ߆ࠄߩ੤ㅢᨑߣ
่วߒߚᓟ RcL ࠍಽᨑߒ‫ޔ‬஥⣻ოߩ╙ 2-3 ጀ㑆ࠍ⿛ⴕߒ‫ޔ‬⣻⋥╭㕽ߦ౉ࠅ Rca ࠍಽᨑߔࠆ‫ ߁߽ޕ‬1 ᨑߪᄖ஥ᄢ⣽
⊹␹⚻㧔CFL㧕߳ߩ੤ㅢᨑࠍಽᨑߒߚᓟ‫ޔ‬㒶ㇱᄢ⣽␹⚻ߣߥࠆ‫ޕ‬L4㧦CFL ߳ߩᨑ‫ޔ‬ᄢ⣽␹⚻㧔F㧕߳ߩᨑ‫ޔ‬㐽㎮␹
⚻㧔O㧕߳ߩᨑߩ 3 ᨑߦಽጘߔࠆ‫ޕ‬L5㧦F ߳ߩᨑ‫ޔ‬O ߳ߩᨑ‫ޔ‬ထ㛽␹⚻߳ߩᨑ 3 ᨑߦಽጘߔࠆ㧔ಽጘ␹⚻㧕‫ޕ‬એ਄
ࠃࠅ‫࡞ࠩࠬ࡝ޔ‬⣶␹⚻ฌߢߪ‫ޔ‬L2+L3 ߹ߢᮡḰ⊛ߥ⡻㑆␹⚻ߣห᭽ߥ⚻〝ࠍ⿛ࠆߎߣ߇ࠊ߆ߞߚ‫ޔߪߣߎߩߎޕ‬
࡝ࠬࠩ࡞ߩ૕ᐙߩ㗔ၞߪࡅ࠻ࠃࠅ߽ਅ૏ಽ▵߹ߢᐢ߇ߞߡ޿ࠆߣ⸒߃ࠆ‫ޕ‬⣶ᬁߩᢙߩ㆑޿ߣߩ㑐ㅪ߇޽ࠆߩߢߪ
ߥ޿߆ߣ⠨߃ߡ޿ࠆ‫⎇ᧄޕ‬ⓥߪ‫ޔ‬੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲߩ౒ห೑↪⎇ⓥߣߒߡታᣉߐࠇߚ‫ޕ‬
B-52 㔤㐳㘃ߦ߅ߌࠆࡉ࠼࠙⃿⩶ߩㅴൻ↢ᘒቇ⊛⎇ⓥ
૒‫ᧁޘ‬ፏ㧔ᗵᨴ⎇࡮ࠥࡁࡓ࠮ࡦ࠲࡯㧕 ᚲౝኻᔕ⠪㧦㋈ᧁ᮸ℂ
వⴕ⎇ⓥߩ⚿ᨐ߆ࠄ‫ޔ‬Staphylococcus delphini ࠣ࡞࡯ࡊߩ⩶⒳⟲߇ࡠ࡯࡜ࠪࠕ₞㘃․⇣⊛ߦᏱ࿷ߒ‫ᧄޔ‬ዻ⩶߇ູ੃
㘃ኋਥߣ౒ㅴൻ㑐ଥߦ޽ࠆߎߣ߇␜ໂߐࠇߡ޿ߚ‫⎇ᧄޕ‬ⓥߢߪ‫࠻ࡅޔ‬એᄖߢࡉ࠼࠙⃿⩶ߩ↢ᘒ߇ਇ᣿ߢ޽ߞߚ㔤㐳
⋡േ‛⒳ߩࡉ࠼࠙⃿⩶⒳ಽᏓࠍ⺞ߴߚ‫╭ࡦࠫࡒ࠻࠺ࡔޔࡦࡒ࠲ࠤޕ‬ᵈᛩਈߦࠃࠅోり㤗㈮ࠍታᣉߒߚ୘૕ߩ㥦೨ᐸ‫ޔ‬
ᄖ㒶ㇱࠍṌ⩶✎᫔ߢ᜞޿‫ޔ‬ᬌ૕ߣߒߚ‫ᧄޕ‬ᣉ⸳߅ࠃ߮࿖ౝⶄᢙᣉ⸳ࠃࠅ‫ޔ⑼࡞ࠩࠟ࠽ࠝޔ⑼࡞ࠩࠟ࠽࠹ޔ⑼࠻ࡅޔ‬
ࠝࡑࠠࠩ࡞⑼‫ ⸘ޔ⑼ࠬ࡝ࡠޔ⑼࡞ࠩࡀ࠷ࠠޔ‬6 ⑼‫ޔ‬22 ⒳‫ޔ‬286 ୘૕߆ࠄࡉ࠼࠙⃿ࠍಽ㔌ߒߚ‫ޕ‬S. aureus ߪ㔤㐳㘃ో
㧙㧙
⥸ߦ‫ޔ‬S. warneri, S. pasteuri ߪ⌀₎㘃ߦ߅޿ߡᐢߊಽᏓߒߡ޿ߚ‫࠻ࡅޕ‬਄⑼ߢߪ S. epidermidis, S. capitis, S. caprae㧘
ᣂ਎⇇ࠩ࡞ߢߪ S. simiae ߇ߘࠇߙࠇ․⇣⊛ߦ଻⩶ߐࠇߡ޿ߚ‫⚿ߩࠄࠇߎޕ‬ᨐ߆ࠄ‫ޔ‬㔤㐳㘃ߣ S. aureus ࠣ࡞࡯ࡊ㧔਄
⸥ 7 ⩶⒳㧕ߪ౒ㅴൻ㑐ଥߦ޽ࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬
ൻ⍹⸽᜚ߣಽሶᤨ⸘ࠍၮߦߒߚಽጘᐕઍផቯߪ‫ߪߢ⩶⚦ޔ‬ൻ⍹ᖱႎ߇ߥ޿ߚ߼࿎㔍ߢ޽ߞߚ‫⎇ᧄޕ‬ⓥߩࡉ࠼࠙⃿
⩶ో 44 ⒳ߩ↢ᘒቇ⊛ᖱႎߪ‫੃ູޔ‬㘃ฦ⒳ߩᐕઍផቯ୯ߣ✚วߔࠆߎߣߢ‫⩶⃿࠙࠼ࡉޔ‬ዻߩൻ⍹⸽᜚ߩઍ↪ߣߥࠅ
߁ࠆ‫⃻ޕ‬࿷ࡉ࠼࠙⃿⩶ዻో⩶⒳ోࠥࡁࡓႮၮ㈩೉ࠍ࠺࡯࠲࠮࠶࠻ߦ↪޿ߚᐕઍផቯࠍ⹜ߺߡ޿ࠆ‫ޕ‬
B-53 ࠴ࡦࡄࡦࠫ࡯߆ࠄߩࡅ࠻↱᧪∛ේ૕ߩಽ㔌
㇭ጊዏ♿㧔ᣣᧄࡕࡦࠠ࡯࠮ࡦ࠲࡯㧕 ᚲౝኻᔕ⠪㧦ችㇱ⾆ሶ
㔤㐳㘃⎇ⓥᚲߩ࠴ࡦࡄࡦࠫ࡯6 㗡ߦߟ޿ߡ‫ޔ‬ຜ㗡෸߮㥦⣧ߩ᜞޿ᶧ߆ࠄࡅ࠻↱᧪∛ේ૕ߩಽ㔌ࠍ⹜ߺߚ‫ࡅߦ․ޕ‬
࠻ࡔ࠲࠾ࡘ࡯ࡕ࠙ࠗ࡞ࠬ㧔hMPV㧕‫ޔ‬RS ࠙ࠗ࡞ࠬ㧔RSV㧕ߦߟ޿ߡ࠙ࠗ࡞ࠬಽ㔌ߣ∛ේ૕ㆮવሶߩᬌ಴ࠍⴕߥߞߚ‫ޕ‬
ߘߩ⚿ᨐ‫ߪࠬ࡞ࠗ࠙ޔ‬ᬌ಴ߐࠇߕ‫⩶⚦ޔ‬ၭ㙃ߢ߽Ᏹ࿷⩶ߦߣߤ߹ߞߚ‫ޔߚ߹ޕ‬એ೨߆ࠄⴕߥߞߡ߈ߚⴊᷡቇ⊛⸃ᨆ
ࠍ↪޿ߚੱ↱᧪∛ේ૕߳ߩᗵᨴ⁁ᴫࠍ⺞ߴߚ⚿ᨐ‫ߣ࠲࡯࠺ߩߢ߹ࠇߎޔ‬Ყセߒߡ․ߦᗵᨴ₸ߩ㜞߆ߞߚ∛ේ૕ߩਛ
ߢ㔤㐳⎇ߩ࠴ࡦࡄࡦࠫ࡯ߦ߅ߌࠆ௑ะࠍߟ߆߻ߎߣ߇ߢ߈ߚ‫ߩߘޕ‬ਛߢ๭ๆེ♽ᗵᨴ∝ࠍᒁ߈⿠ߎߔ⊖ᣣຕ⩶‫ࡄޔ‬
࡜ࠗࡦࡈ࡞ࠛࡦࠩ࠙ࠗ࡞ࠬ III‫ޔ‬RSV ߪ㔤㐳⎇↢߹ࠇߩ୘૕㧔A㧕ߣ↢߹ࠇߡ߆ࠄዉ౉ߐࠇߚ୘૕㧔B㧕ߩ᛫૕ଔߦ
ఝ૏ߥᏅߪ⷗ࠄࠇߥ߆ߞߚ‫ޔߒ߆ߒޕ‬hMPV ߪ A ⟲ࠃࠅ߽ B ⟲ߦ߅޿ߡఝ૏ߦ᛫૕ଔ߇㜞ߊ‫ߣ࠻ࡅޔ‬ห᭽ߦ㜞ᐕ
㦂ߦ߅޿ߡ㜞᛫૕ଔ߇⏕⹺ߐࠇߚ‫ߣ࠻ࡅ߽ߡ޿߅ߦ࡯ࠫࡦࡄࡦ࠴ߪࠬ࡞ࠗ࠙ߩࠄࠇߎޔߚ߹ޕ‬ห᭽ߦ࠙ࠗ࡞ࠬ߇વ
᠞ߒߚߩߜ‫ࠍ∉఺ޔ‬₪ᓧߢ߈ߚน⢻ᕈ߽␜ߒߡ޿ࠆ‫ޕ‬
B-54 㔤㐳㘃ࡋ࡞ࡍࠬ࠙ࠗ࡞ࠬߦ㑐ߔࠆ⎇ⓥ
శ᳗✚ሶ㧘ਛ᧛િ㧔NPO ࡊ࡜ࠗࡔࠗ࠻࡮ࠕࠧ࡜࡮ࡃࠗࠝࡔ࠺ࠖࠞ࡞⎇ⓥᚲ㧕 ᚲౝኻᔕ⠪㧦᣿㉿ብᢥ
ࠨ࡞ B ࠙ࠗ࡞ࠬ㧔BV㧕ߪࡑࠞࠢࠩ࡞ࠍ⥄ὼኋਥߣߔࠆࠕ࡞ࡈࠔࡋ࡞ࡍࠬ࠙ࠗ࡞ࠬߢ‫ߦ࠻ࡅޔ‬ᗵᨴߒߚ႐วߪ
᛫࠙ࠗ࡞ࠬᴦ≮ࠍᣉߐߥ޿ߣ㊀◊ߥਛᨔ␹⚻♽㓚ኂࠍᒁ߈⿠ߎߔߎߣ߇޽ࠅ߹ߔ‫ޕ‬BV ࠍ฽߻㔤㐳㘃ࠕ࡞ࡈࠔࡋ
࡞ࡍࠬ࠙ࠗ࡞ࠬߪ᛫ේ੤Ꮕᕈ߇޽ࠆߚ߼‫ࠆ޿↪ࠍ࡞ࠩࠢࠞࡑޔ‬ታ㛎േ‛ᣉ⸳ߢߪ BV ․⇣⊛᛫૕ᬌᩏ߇ਇนᰳߣ
ߥࠅ߹ߔ‫ޔߪߜߚ⑳ޕ‬BV ․⇣⊛ࡍࡊ࠴࠼㧔BV-gD CP㧕ࠍ Covalink ߦࠞ࠶ࡊ࡝ࡦࠣߐߖ‫ޔ‬ലᨐ⊛ߥࡉࡠ࠶ࠠࡦ
ࠣࠍᬌ⸛ߔࠆߥߤߒߡ‫ޔ‬㜞ᗵᐲߥ BV ․⇣⊛ࡍࡊ࠴࠼ ELISA ᴺࠍ⏕┙ߒ‫ߩߘޔ‬ᔕ↪ࠍ࿑ߞߡ޿߹ߔ‫ޕ‬
੹࿁‫ޔ‬੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲࠃࠅଏਈߐࠇߚࡑࠞࠢࠩ࡞ⴊᶧࠨࡦࡊ࡞ࠍ฽߻ BV 㓁ᕈ୘૕ࠨࡦࡊ࡞ߦߟ޿ߡ‫ޔ‬
᛫ BV-gD CP ᛫૕ᬌ಴㗫ᐲࠍᬌ⸛ߒ߹ߒߚ‫࠻ࡅޔߚ߹ޕ‬ᗵᨴߢߩᬌ಴ࠍᗐቯߒ‫࠻ࡅޔ‬න⚐ࡋ࡞ࡍࠬ࠙ࠗ࡞ࠬ㧔HSV㧕
㓁ᕈࡅ࠻ⴊẏࠨࡦࡊ࡞ߦ‫ޔ‬ᓸ㊂ߩ BV 㓁ᕈࡑࠞࠢࠩ࡞ⴊẏࠨࡦࡊ࡞ࠍട߃‫ޔ‬᛫ BV-gD CP ᛫૕ᬌ಴ࠍ⹜ߺ߹ߒߚ‫ޕ‬
BV 㓁ᕈࡑࠞࠢⴊᶧࠨࡦࡊ࡞ߦ߅ߌࠆ᛫ BV-gD CP ᛫૕㓁ᕈ₸ߪ⒳‫↥ޔ‬࿾ߦࠃߞߡ⇣ߥࠅ 65%߆ࠄ 93%ߣ޿߁
⚿ᨐ߇ᓧࠄࠇ߹ߒߚ‫ޔߚ߹ޕ‬HSV 㓁ᕈࡅ࠻ⴊẏਛߦ 0.5%ߩഀวߢ BV 㓁ᕈࡑࠞࠢࠩ࡞ⴊẏ߇฽߹ࠇࠇ߫‫ޔ‬᛫
BV-gD CP ᛫૕߇ᬌ಴ߢ߈ࠆߎߣ߇᣿ࠄ߆ߦߥࠅ߹ߒߚ‫ޕ‬
B-56 ࠧ࡝࡜ߦ߅ߌࠆ╭㛽ᩰ♽ߦ㑐ߔࠆ⎇ⓥ
ᄢ⍹రᴦ㧔ᣣ₞࡮₞ක㧕㧘⩆ේ⋥㆏㧔ᘮᙥᄢ࡮ℂᎿ㧕㧘⩵ᳰᵏᒄ㧔૒⾐ᄢ࡮ක㧕㧘ዊ⮐ᄢテ㧔੩ᄢ࡮ඳ‛㙚㧕 ᚲౝ
ኻᔕ⠪㧦ᳯᧁ⋥ሶ
ᄢဳ㘃ੱ₎ߦ߅ߌࠆ྾⢇ㆇേᯏ⢻ࠍ⎇ⓥߔࠆ৻Ⅳߣߒߡ‫࡜࡝ࠧ࠼ࡦ࡜࡯ࡠࠪ࠾ޔ‬㧔1 ୘૕‫ޔ‬㓶㧕ߩ೨⢇╭ߩ⊒㆐
㧔㧩╭ߩ⾰㊂㧕ߣ‫⊒ޔ‬ើ╭ജ㧔㧩╭ߩ↢ℂቇ⊛ᢿ㕙Ⓧ PCSA㧕ߦ⌕⋡ߒ‫ޔ‬ዱ૕ߩ⸃೬ࠍⴕߞߚ‫ࠕࠫࠕޔߪ࡜࡝ࠧޕ‬
ߩᄢဳ㘃ੱ₎ߢ޽ࠆࠝ࡜ࡦ࠙࡯࠲ࡦߦᲧߴ࿾਄௑ะ߇ᒝߊ‫ࠞ࡝ࡈࠕߩઁߩߤߥࡏࡁࡏ߿࡯ࠫࡦࡄࡦ࠴ޔ‬㘃ੱ₎ߩࡠ
ࠦࡕ࡯࡚ࠪࡦߦ㘃ૃߒߡ޿ࠆ‫ߩࡦ࡚ࠪ࡯ࡕࠦࡠߩࠄࠇߎޕ‬Ꮕ⇣ߪ‫ޔ‬೨⢇╭ߩ⊒㆐߿⊒ើ╭ജߦᓇ㗀ࠍਈ߃ࠆߎߣ߇
੍᷹ߐࠇࠆ‫⎇ᧄޕ‬ⓥߢߪ‫ޔ‬ฦ╭ߩ╭⾰㊂ߣ PCSA ࠍ‫ߩࠇߙࠇߘޔ‬೨⢇╭ߩ✚๺ߢഀࠆߎߣߦࠃࠅ‫ޔ‬ฦ୯ߩᲧ₸ࠍ▚
಴ߒ‫ߦߢ߹ࠇߎޔ‬ᓧࠄࠇߡ޿ࠆࠝ࡜ࡦ࠙࡯࠲ࡦߩ࠺࡯࠲ߣᲧセࠍⴕߞߚ‫ޕ‬⢀㑐▵ߦ㑐ଥߔࠆ╭⟲ߩ߁ߜ‫ޔ‬િ╭⟲㧔਄
⣨ਃ㗡╭‫ޔ‬⢀╭‫ޔ‬⢛Ṗゞ਄╭㧕ߪ࿾਄ᕈࡠࠦࡕ࡯࡚ࠪࡦᤨߦ૕㊀ࠍᡰᜬߒ‫ޔ‬ዮ╭⟲㧔⣨ᯂ㛽╭‫ޔ‬਄⣨╭‫ޔ‬਄⣨ੑ㗡
╭㧕ߪ᮸਄↢ࡠࠦࡕ࡯࡚ࠪࡦᤨߦផㅴജࠍ↢߻ߩߦ㊀ⷐߥᓎഀࠍᨐߚߒߡ޿ࠆ‫⎇ᧄޕ‬ⓥߦ߅ߌࠆ⚿ᨐߪ‫ࡠߣ⟲╭ޔ‬
ࠦࡕ࡯࡚ࠪࡦߩ㆑޿ࠍ෻ᤋߒߡ߅ࠅ‫ߪߢ࡜࡝ࠧޔ‬⢀㑐▵ߩિ╭⟲߇‫ߪߢࡦ࠲࡯࠙ࡦ࡜ࠝޔ‬⢀㑐▵ߩዮ╭⟲߇ߘࠇߙ
ࠇࠃࠅ⊒㆐ߒߡ߅ࠅ‫ޔ‬ᄢ߈ߥ╭ജࠍ⊒ើߢ߈ࠆ߽ߩߣផ᷹ߐࠇߚ‫ޔߒ߆ߒޕ‬ዮ╭⟲ߩߥ߆ߢ߽ੑ㑐▵╭ߢ޽ࠆ਄⣨
ੑ㗡╭߇ࠝ࡜ࡦ࠙࡯࠲ࡦࠃࠅ߽ࠧ࡝࡜ߦ߅޿ߡ⋧ኻ⊛ߦᄢ߈ߥ╭⾰㊂ߣ PCSA ࠍᜬߞߡ޿ߚ‫੹ޕ‬ᓟ‫ޔ‬㘃ੱ₎ࠍ⸃೬
ߔࠆᯏળ߇޽ࠇ߫‫ޔ‬ᮡᧄᢙࠍჇ߿ߒ‫੹ޔ‬࿁⹺߼ࠄࠇߚᏅ⇣߇‫ߩࡦ࡚ࠪ࡯ࡕࠦࡠޔ‬Ꮕ⇣ࠍ෻ᤋߒߡ޿ࠆ߆ࠍߐࠄߦᬌ
⸛ߒߡ޿߈ߚ޿‫ޕ‬
B-57 ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆࠨ࡞ T ⚦⢩⊕ⴊ∛࠙ࠗ࡞ࠬߩേᘒߩ⸃ᨆ࡮఺∉ᴦ≮
᧻ጟ㓷㓶㧘቟᳗⚐৻ᦶ㧘ਃᶆᧂ⍮㧔੩ㇺᄢ࡮࠙ࠗ࡞ࠬ⎇㧕 ᚲౝኻᔕ⠪㧦᣿㉿ብᢥ
ᦨೋߦ㔤㐳㘃⎇ⓥᚲߩ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ STLV-1 ᛫૕㓁ᕈ₸ࠍᬌ⸛ߒߚ‫ޕ‬ᬌ૕ណขߪ‫౉ޔ‬ᚲਛ࠾ࡎࡦࠩ࡞ߩቯ
ᦼណⴊߩ㓙ߦ‫ޔ‬หᤨߦ STLV-1 ᛫૕ᬌᩏ↪ߩណⴊࠍᣉⴕߒ‫ⴊޔ‬ẏߣනᩭ⃿㧔PBMC㧕ࠍಽ㔌ߒߚ‫ޔߦࠄߐޕ‬એ೨ߦ
ಽ㔌ߐࠇಓ⚿଻ሽߐࠇߡ޿ߚⴊẏ߽᛫૕ᬌᩏߦଏߒߚ‫ޕ‬PA ᴺߦࠃࠆࠬࠢ࡝࡯࠾ࡦࠣߩ⚿ᨐ‫ޔ‬㔤㐳㘃⎇ⓥᚲౝߩ࠾
ࡎࡦࠩ࡞ 374 㗡ਛ 178 㗡㧔48%㧕ߦ STLV-1 ᗵᨴ߇್᣿ߒߚ‫ޕ‬ኻᾖߣߒߡ⸃ᨆߒߚࠕࠞࠥࠩ࡞ߦ߅޿ߡߪ‫ޔ‬132 㗡
ਛ 1 㗡㧔0.8%㧕ߣૐ୯ߢ޽ࠅ‫ޔ‬⒳ߦࠃࠅᗵᨴ₸ߦᄢ߈ߥᏅ߇ሽ࿷ߔࠆ੐߇᣿ࠄ߆ߣߥߞߚ‫ࠬ࡞ࠗ࠙ࡠࡊޕ‬㊂ࠍ
Real-time PCR ᴺߦߡቯ㊂ߒߚߣߎࠈ‫ޔ‬ᗵᨴ⚦⢩₸ߪ 0.001%߆ࠄ 17%ߣᄢ߈ߥ୘૕Ꮕࠍ⹺߼ߚ‫࠻ࡔ࠻ࠗࠨ࡯ࡠࡈޕ‬
࡝࡯ߦࠃࠆ⸃ᨆߢߪ‫ޔ‬CD4 㓁ᕈ T ࡝ࡦࡄ⃿ఝ૏ߦ STLV-1 ߇ᗵᨴߒߡ޿ࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬STLV-1 ↱᧪ߩ Tax
㧙㧙
߅ࠃ߮ HBZ ㆮવሶࠍࠨࡉࠢࡠ࡯࠾ࡦࠣߒ‫ߩࠄࠇߎޔ‬ᵴᕈߦ㑐ߒߡಽሶ↢‛ቇ⊛ߥ⸃ᨆࠍ㐿ᆎߒߚ‫ޕ‬STLV-1 ↱᧪ߩ
Tax ߪ AP-1‫ޔ‬ฎౖ⊛ NF-+B‫ޔ‬Wnt ⚻〝ࠍᵴᕈൻߒ‫ޔ‬HBZ ߪฦ‫ࠍޘ‬ᛥ೙ߒߚ‫৻ޕ‬ᣇ‫ޔ‬TGF-+ࠪࠣ࠽࡞ߦ㑐ߒߡߪ STLV-1
Tax ߪᛥ೙ߒ‫ޔ‬HBZ ߪᵴᕈൻߒߚ‫ߩࠄࠇߎޕ‬ᚲ⷗ߪ HTLV-1 ߣᭂ߼ߡ㘃ૃߒߡ޿ࠆ‫ࠬ࡞ࠗ࠙ޔߪ࡞ࠩࡦࡎ࠾ޕ‬േᘒ
ߩ⹦⚦ߥ⸃ᨆߦ㕖Ᏹߦ᦭ᦸߥ㔤㐳㘃ࡕ࠺࡞ߢ޽ࠆߣ⸒߃ࠆ‫ޕ‬
B-58 ࡅ࠻ᚑੱ࡮⢝ఽ߅ࠃ߮ࠨ ࡞஥㗡㛽ߦ߅ߌࠆᒻᘒቇ⊛ᬌ⸛ߣ਄ඨⷙ▤ⵚ㓗∝୥⟲ߩ∛࿃ߦ㑐ߔࠆ઒⺑ߩᬌ⸽
㜞ᯅ⋥ੱ㧘ⷺ↰◊ା㧘༑ᄙ᧛ஜ㧔᧲੩ක⑼ᱤ⑼ᄢ࡮⡊㥦ຜ༄⑼㧕 ᚲౝኻᔕ⠪㧦⷏᧛೰
਄ඨⷙ▤ⵚ㓗∝୥⟲ߩ∛࿃ࠍតࠆߚ߼‫࠻ࡅޔ‬⢝ఽ࡮ᚑੱ߅ࠃ߮ࠨ࡞㧔࠴ࡦࡄࡦࠫ࡯‫࡞ࠩࡦࡎ࠾ޔ࡞ࠩࠟ࠽࠹ޔ‬㧕
ߩ஥㗡㛽ᒻᘒߦߟ߈‫ޔ‬CT ↹௝ߦࠃࠆᲧセࠍⴕߞߚ‫࠻ࡅޕ‬ᚑੱߢߪਛ㗡⬄ᐩߣ਄ඨⷙ▤ߪ߶߷ធߒߡ߅ࠅ‫ޔ‬⢝ఽᦼ
ߦߪ਄ඨⷙ▤ߪ㗡⬄ౝߦᄢ߈ߊ⓭಴ߒߡ޿ߚ‫৻ޕ‬ᣇ‫੹ޔ‬࿁᠟ᓇࠍⴕߞߚࠨ࡞ߢߪ‫ޔ‬਄ඨⷙ▤਄ᣇߦ฽᳇ⱎᎽߩ⊒㆐
߇⷗ࠄࠇ‫ޔ‬ਛ㗡⬄ᐩߣඨⷙ▤ߪ㔌ࠇߡ޿ߚ‫ࠆߌ߅ߦ࠻ࡅޕ‬⣖ߣౝ⡊ߩㄭធ߇‫ޔ‬਄ඨⷙ▤ⵚ㓗ߩᚑ࿃ߦ㑐ਈߔࠆน⢻
ᕈ߇␜ໂߐࠇߚ‫ޕ‬
㧨ቇળ⊒⴫㧪
Collegium Oto-Rhino-Laryngologicum Amicitiae Sacrum (September 2011, Belgium)
㧨⊒⴫᭎ⷐ㧪
਄ඨⷙ▤ⵚ㓗∝୥⟲ߪᲧセ⊛ᣂߒ޿∔ᖚ᭎ᔨߢ޽ࠅ‫ޔ‬ㄭᐕ‫∛ޔ‬ᘒ↢ℂቇ⊛ߥࠕࡊࡠ࡯࠴ߢߩ⸃᣿ߪㅴࠎߢ޿ࠆ߇‫ޔ‬
∛࿃ߦ㑐ߒߡߪᧂߛቯ⺑ߪ⏕┙ߒߡ޿ߥ޿‫ޕ‬⢝↢ᣧᦼߦ㛽ൻ߇ቢੌߔࠆౝ⡊ߦ߅޿ߡ㛽ᰳ៊߇↢ߓࠆࡔࠞ࠾࠭ࡓࠍ
⸃᣿ߔࠆߚ߼‫࠻ࡅޔ‬⢝ఽ࡮ᚑੱ߅ࠃ߮ࠨ࡞ߦߟ޿ߡ஥㗡㛽 CT ࠍ᠟ᓇߒ‫ޔ‬ᒻᘒቇ⊛ᬌ⸛ࠍⴕ޿⠨ኤࠍട߃ߚ‫࠻ࡅޕ‬
⢝ఽ㧔⢝↢ 16 ㅳ‫ޔ‬26 ㅳ‫ޔ‬26 ㅳ‫ޔ‬28 ㅳ㧕ߢߪ‫ޔ‬ਛ㗡⬄ᐩߦ߅޿ߡ᣿ࠄ߆ߥౝ⡊ߩ਄ᣇ⓭಴߇⷗ࠄࠇ‫ޔ‬CT ᢿ㕙ߢߪ
਄ඨⷙ▤߇ᦨ߽਄ᣇ߳૏⟎ߒ⎬⤑ߣធߔࠆࠃ߁ߦሽ࿷ߒߡ޿ߚ‫࠻ࡅޕ‬ᚑੱߦ߅޿ߡ߽਄ඨⷙ▤਄ㇱߣਛ㗡⬄ᐩߪធ
ߔࠆࠃ߁ߥ૏⟎㑐ଥߢ޽ߞߚ‫৻ޕ‬ᣇ‫࡯ࠫࡦࡄࡦ࠴ޔ‬4 ૕‫ ࡞ࠩࠟ࠽࠹ޔ‬4 ૕‫ ࡞ࠩࡦࡎ࠾ޔ‬4 ૕ߦߟ޿ߡห᭽ߩᬌ⸛ࠍ
ⴕߞߚߣߎࠈ‫ޔ‬ඨⷙ▤਄ᣇߦߪ฽᳇ⱎᎽߩ⊒⢒߇⷗ࠄࠇ‫ޔ‬ਛ㗡⬄ᐩߣౝ⡊ߪⓨ㑆⊛ߦ㔌ࠇߚ૏⟎㑐ଥߦ޽ߞߚ‫߹ޕ‬
ߚ‫ߩ࡞ࠨߣ࠻ࡅޔ‬౰⁁ᢿࠍᲧセߒߚߣߎࠈ‫ߩ࠻ࡅޔ‬ᣇ߇ਛ㗡⬄ᐩߩ஥ᣇ߳ߩ௑ᢳ߇ዊߐߊ‫ޔ‬஥㗡⪲߇ਛ㗡⬄ᐩࠍ᛼
ߒਅߍࠆࠃ߁ߥᒻ⁁ࠍߒߡ޿ߚ‫ߩࠄࠇߎࠆߌ߅ߦ࠻ࡅޕ‬ᒻᘒቇ⊛․ᓽߦࠃߞߡ⢝↢ᦼߦౝ⡊⤑ㅅ〝ߣ⎬⤑ߩធ⸅߇
↢ߓ‫ޔ‬㛽ൻ߇ᅹߍࠄࠇࠆߎߣߦࠃࠅඨⷙ▤ⵚ㓗߇ᒻᚑߐࠇࠆน⢻ᕈ߇޽ࠆߣ⠨߃ߚ‫ߡ޿߅ߦ࠻ࡅޔߚ߹ޕ‬ᒻᘒቇ⊛
Ꮕ⇣߇↢ߓࠆⷐ࿃ߣߒߡ‫ޔ‬ᄢ⣖ኈⓍ₸ߩჇട‫ࡦࠪࠝࡒޔ‬ᄌ⇣ߦࠃࠆ஥㗡╭㊂ߩᷫዋ‫ޔ‬਄ඨⷙ▤ᓘߩ⋧ኻ⊛ߥჇᄢ߇
⠨߃ࠄࠇࠆߣߩ⠨ኤࠍട߃ߚ‫ޕ‬
B-59 ࡑࠞࠢࠍ↪޿ߚᣂⷙᱤ㜑ౣ↢≮ᴺߩ⏕┙
╴੗ஜᄦ㧘㠽ዬᄢ␭㧔ᣣᧄᱤᄢ࡮↢๮ᱤቇㇱ࡮⮎ℂ㧕 ᚲౝኻᔕ⠪㧦㋈ᧁ᮸ℂ
ᐔᚑ23ᐕᐲߪ‫࡞ࠩࡦࡎ࠾ޔ‬1଀㧔10ᱦ㧕ߣࠕࠞࠥࠩ࡞1଀㧔11ᱦ㧕ߩਅ㗶㛽ߩណขᓟ‫ߪߡ޿߅ߦ࡞ࠩࡦࡎ࠾ޔ‬ਅ㗶
ฝ஥╙ੑዊ⥓ᱤߣਅ㗶Ꮐ஥╙৻ᄢ⥓ᱤߩ⭯ಾಾ ࠍ૞ᚑߒࡋࡑ࠻ࠠࠪ࡝ࡦ࡮ࠛࠝࠫࡦᨴ⦡㧔H-Eᨴ⦡㧕ࠍⴕ޿‫ࠕޔ‬
ࠞࠥࠩ࡞ߪਅ㗶ฝ஥╙ੑዊ⥓ᱤߣਅ㗶ฝ஥╙৻ᄢ⥓ᱤߦ߅ߌࠆH-Eᨴ⦡ࠍⴕߞߚ‫ߚ߹ޕ‬ᷙวᱤ೉ᦼߢ޽ࠆࠕࠞࠥࠩ
࡞1଀㧔3ᱦ㧕ࠃࠅ਄㗶ฝ஥ਛಾᱤ‫ޔ‬਄㗶ฝ஥஥ಾᱤ‫ޔ‬਄㗶ฝ஥ၒફ›ᱤ‫ޔ‬਄㗶ฝ஥੃›ᱤ‫ޔ‬਄㗶ฝ஥╙৻ᄢ⥓ᱤ‫ޔ‬
ਅ㗶ฝ஥ਛಾᱤ‫ޔ‬ਅ㗶ฝ஥╙ੑ੃⥓ᱤ‫ޔ‬ਅ㗶ฝ஥╙৻ᄢ⥓ᱤߩᱤ㜑⚦⢩ࠍណขߒೋઍၭ㙃ࠍⴕߞߚ‫ޕ‬H-Eᨴ⦡ࠍⴕ
ߞߚ࠾ࡎࡦࠩ࡞ߣࠕࠞࠥࠩ࡞ߩోߡߩ⥓ᱤߢ‫⽎ޔ‬‎⾰‫ޔ‬ᱤ㜑‫⽎ޔ‬‎⧘⚦⢩ጀ‫ޕߚ߈ߢ߇ߣߎࠆߔ⹺⏕ࠍ▤ⴊޔ⚻␹ޔ‬
߹ߚ‫⚿ߩߎޔ‬ᨐࠃࠅ࠾ࡎࡦࠩ࡞ߣࠕࠞࠥࠩ࡞ߩᱤ㜑⚵❱ߪࡅ࠻ߣ㘃ૃߒߡ޿ࠆߎߣ߇ⷰኤߐࠇߚ‫ޕ‬ೋઍၭ㙃ߪ‫ޔ‬ᷙ
วᱤ೉ᦼߩࠕࠞࠥࠩ࡞ߩ8ᧄߩᱤࠃࠅᱤ㜑⚦⢩ࠍណขߒⴕߞߚ‫ߚ߹ޕ‬਄㗶ฝ஥╙৻ᄢ⥓ᱤ↱᧪⚦⢩ߢߪࠦࡠ࠾࡯ᒻ
ᚑ₸ߩ⸃ᨆࠍⴕ޿‫ޔ‬1 x 104୘ߩ⚦⢩ࠍ100 mm ࠪࡖ࡯࡟4ᨎߦ᠞⒳ߒᐔဋ3ߟߩࠦࡠ࠾࡯߇ᒻᚑߐࠇߚ‫੹ޕ‬ᓟߪߎࠇ
ࠄ⚦⢩ࠍ↪޿ߡ‫ޔ‬in vitro⸃ᨆߢߪ⚦⢩Ⴧᱺࠍ‫ޔ‬in vivo⸃ᨆߢߪ⊹ਅ⒖ᬀߥߤࠍⴕ޿‫ߩ᧪↱ࠢࠞࡑޔ‬ᱤ㜑⚦⢩ߩ⚦⢩
․ᕈࠍಽሶ↢‛ቇ⊛ߦ⸃ᨆߔࠆ‫ޕ‬
B-60 㔤㐳㘃ߩ✂⤑㤛᢬ߦ․⇣⊛ߦ⊒⃻ߔࠆㆮવሶ⟲ߩหቯ
ฎᎹ⾆ਭ㧘૒⽾ℂ૫ሶ㧘⨹ᧁ┨ਯ㧔㧔⽷㧕ᄢ㒋ࡃࠗࠝࠨࠗࠛࡦࠬ⎇ⓥᚲ㧕 ᚲౝኻᔕ⠪㧦ᄢ⍹㜞↢
ࡅ࠻ࠍ฽߼ߚ㔤㐳㘃ߩ✂⤑ߪਛᔃㇱߦ㤛᢬ߣ޿߁㍙૕⚦⢩ߩኒᐲ߇㜞ߊ‫ⷞޔ‬ജߦ㊀ⷐߥ᭴ㅧࠍᜬߟ‫ޕ‬ᚒ‫ޔߪޘ‬㤛
᢬⊒↢ߦ㑐ࠊࠆㆮવሶ⟲ߩหቯࠍ⋡⊛ߣߒߡ‫ޔ‬๟↥ᦼࠕࠞࠥࠩ࡞ߩ✂⤑ࠍ㤛᢬ㇱߣ๟ㄝㇱߦಽߌߡណขߒ‫ߙࠇߘޔ‬
ࠇߩ✚RNAߦߟ޿ߡࡑࠗࠢࡠࠕ࡟ࠗࠍ↪޿ߡㆮવሶ⊒⃻ࠍᲧセߒߚ‫ߢߎߘޕ‬ᓧࠄࠇߚ୥⵬ㆮવሶߩਛߢ߽․ߦ
SREBP2ߦ⌕⋡ߒߡ޿ࠆ‫ޕ‬SREBP2ߪ⢽⾰ઍ⻢ߦ㑐ࠊࠆㆮવሶ⟲ߩ⊒⃻ࠍ೙ᓮߔࠆォ౮࿃ሶߢ޽ࠅ‫ޔ‬in situࡂࠗࡉ࡝
࠳ࠗ࠯࡯࡚ࠪࡦߦࠃߞߡࡑ࠙ࠬ✂⤑ߦ߅޿ߡ߽⊒↢ᦼⷞ⚦⢩ߦ⊒⃻ࠍ⹺߼ࠆ‫ޕ‬ᤓᐕߦᒁ߈⛯߈‫ޔ‬SREBP2ߩⷞ⚦⢩
ߦ߅ߌࠆ࠼ࡒ࠽ࡦ࠻ࡀࠟ࠹ࠖࡉᄌ⇣૕ߦߟ߈⸃ᨆਛߢ޽ࠆ‫ޕ‬
B-61 ࠨ࡞㘃ߦ߅ߌࠆ⣢⚿⍹ߩ∉ቇ⎇ⓥߣ⥄ὼ⊒∝ࡕ࠺࡞ߩត⚝
ỳᧄ๟ㅧ㧘㇭ஜੑ㇢㧘ᚭỈ໪৻㧘቟੗ቁ๟㧘ጟ↰ᷕᔒ㧘↰ญ๺Ꮖ㧘ᑝἑᵏᒾ㧔ฬᏒᄢ࡮⣢ᴲዩེ⑼ቇ㧕 ᚲౝኻᔕ
⠪㧦㋈ᧁ᮸ℂ
ᤓᐕߪ‫᧲ޔ‬ᣣᧄᄢ㔡ἴߦ߆߆ࠊࠆᓳ⥝ක≮ᡰេ⠪ߣߒߡ⃻࿾ߢᵴേߒߡ޿ߚߎߣ߽޽ࠅ‫⎇ޔ‬ⓥㅴ᝞߇ㆃࠇߡ޿ࠆ‫ޕ‬
หᤨߦᐔⴕߒߡⴕߞߚࡑ࠙ࠬߢߩᚑᨐႎ๔ࠍⴕ߁‫ޕ‬
⚿⍹ߩ᭴ᚑᚑಽߩ㧝ߟߢ޽ࠆࠝࠬ࠹ࠝࡐࡦ࠴ࡦ㧔OPN㧕ߪ‫ߡߦࡦࡆࡦࡠ࠻ޔ‬ಾᢿߐࠇࠆᯏ⢻⊛ࠕࡒࡁ㉄㈩೉߇޽
ࠆ‫⎇ᧄޕ‬ⓥߢߪหㇱ૏ߩࠕࡒࡁ㉄㈩೉㧔SLAYGLR㧕ߦኻߔࠆਛ๺᛫૕ࠍ૞ᚑߒ‫ޔ‬OPN ᛫૕ߩ⣢⚿⍹ᒻᚑߦਈ߃ࠆ
ᓇ㗀ࠍᬌ⸛ߒߚ‫ޕ‬OPN ߩ SLAYGLR ㈩೉ࠍ฽߻ࡍࡊ࠴࠼ࠍ↪޿‫࡞࠽࡯ࡠࠢࡁࡕޔ‬᛫૕㧔35B6 ᛫૕㧕ࠍ૞ᚑߒ‫ޔ‬8
㧙㧙
ㅳ㦂 C57BL/6 ࡑ࠙ࠬߦࠣ࡝ࠝࠠࠪ࡞㉄ࠍ⣻⣧ౝㅪᣣᛩਈߔࠆߣߣ߽ߦ‫ޔ‬35B6 ᛫૕ࠍᛩਈ‫⚿ޔ‬᥏ᒻᚑࠍ⹏ଔߒߚ‫ޕ‬
⚿᥏ᒻᚑ㊂ߪ‫ޔ‬᛫૕ᛩਈߦࠃࠅኈ㊂ଐሽᕈߦૐਅߒߚ‫ޕ‬㔚ሶ㗼ᓸ㏜ߦࠃࠆⷰኤߪ‫ޔ‬Control ⟲ߢߪ᡼኿⁁ߩ⚿᥏߇
ዩ⚦▤⚦⢩ߦขࠅㄟ߹ࠇߡ޿ߚ߇‫ޔ‬᛫૕ᛩਈ⟲ߦ߅޿ߡߪ‫ޔ‬ዩ⚦▤⣧ౝߦ⣕⪭⚵❱ߪ⹺߼ࠆߩߺߢ⚿᥏ᒻᚑߪ⹺߼
ߥ߆ߞߚ‫ޕ‬
એ਄ࠃࠅ‫ޔ‬ಾᢿဳ OPN ߦኻߔࠆਛ๺᛫૕ߪ‫⚿ޔ‬⍹ᒻᚑࡑ࠙ࠬߦ߅޿ߡ⚿᥏ߩዩ⚦▤਄⊹߳ߩធ⌕ࠍᛥ೙ߔࠆߎ
ߣߢ‫⚿ޔ‬⍹ᒻᚑࠍ੍㒐ߔࠆߣ␜ໂߐࠇߚ‫⚿ߩߎޕ‬ᨐࠍ J Bone Miner Reserch ߳ᛩⓂߒ accept ߐࠇߚ‫ޕ‬
B-62 ࠨ࡞⢝੿⢖ૐᒻᚑߩሶችౝ࿁ᓳ㧙⟠᳓ㆊዋߦࠃࠆ⢖ૐᒻᚑࡕ࠺࡞૞ᚑߣᚑ㐳࿃ሶ⸃ᨆ
ජ⪲ᢅ㓶㧘૒⮮ᵗ᣿㧘ᩑᧄ㓉ᔒ㧔࿖┙ᚑ⢒ක≮⎇ⓥ࠮ࡦ࠲࡯࡮⥃ᐥ⎇ⓥ࠮ࡦ࠲࡯㧕 ᚲౝኻᔕ⠪㧦㋈ᧁ᮸ℂ
ᧄ⎇ⓥߩ⋡⊛ߪ‫ޔ‬⢝ఽᦼߩ⢖ૐᒻᚑߦኻߒ᦭ലᕈ߇ႎ๔ߐࠇߡ޿ࠆ⃻ⴕߩሶችౝᄖ⑼ᚻⴚ㧔⢝ఽ᳇▤㐽Ⴇⴚ㧕ߩ
૞↪ᯏᐨࠍ‫⺃ޔ‬ዉߐࠇࠆ⚵❱ᚑ㐳࿃ሶߩหቯߦࠃࠅ⸃᣿ߔࠆߎߣߢ޽ࠆ‫ޕ‬૬ߖߡ‫❱⚵ߩߘޔ‬ᚑ㐳࿃ሶࠍૐᒻᚑ⢝ఽ
⢖ߦዪᚲ⊛ߦ૞↪ߐߖࠆߎߣߢ‫ޔ‬⢝ఽ᳇▤㐽Ⴇⴚߩ㧔⢖ౣ↢㧕ലᨐჇᒝߥ޿ߒߘߩઍᦧૐଚⷅᴦ≮ᚻᛛࠍ⏕┙ߔࠆ
ߎߣߦ޽ࠆ‫ޕ‬
ߎߩᄖ⑼⊛ᴦ≮ᴺߪ‫⥃ߛ߹޿ޔ‬ᐥ⊛ߦ⏕┙ߐࠇߡߪߥ޿‫ޕ‬
ߔߥࠊߜ‫ߩߎޔ‬ᚻᛛߪ⃻࿷‫ޔ‬experimental ߥ޿ߒ innovative
therapy ߩ▸⇵ߦ౉ࠆ߽ߩߢ޽ࠅ‫ޔ‬዁᧪ߩᮡḰൻࠍ⋡ᜰߒߡචಽߥታ㛎⊛⹏ଔࠍᔅ㗇ߣߔࠆ‫ޔߪࠇߘޕ‬ᖚఽߩᢇ๮
ᚻᲑߣߒߡߪ‫ߥ߁ࠃߩߎޔ‬ᚻᛛએᄖߦ‫੹ޔ‬ᓟน⢻ᕈߩ޽ࠆ߽ߩ߇⃻ᤨὐߢߪోߊሽ࿷ߒߥ޿ߚ߼ߢ޽ࠆ‫ޕ‬
ߎߩࡕ࠺࡞૞ᚑߩߚ߼ߦߪ‫⟠ޔ‬᳓ߩ৻ㇱ㒰෰ߦࠃࠆᜬ⛯⊛ߥ⟠᳓ㆊዋ⁁ᘒࠍ૞ࠅ಴ߔᔅⷐ߇޽ࠆ‫ޕ‬
৻ᤓᐕᐲߪᒰೋ⸘↹ߒߡ޿ߚᚻⴚᚻᛛߩ⏕┙߅ࠃ߮ᚑ㐳࿃ሶࠍ․ቯߔࠆታ㛎ࠍⴕ߁೨ߦ‫ޔ‬ᘕ㊀ࠍᦼߒߡ੍஻ታ㛎
ࠍⴕߞߚ‫ౕޕ‬૕⊛ߦߪᤓᐕᐲⴕߞߚၮ␆⊛ታ㛎㧔⿥㖸ᵄࠛࠦ࡯ࠍ૶ߞߡᲣ૕ᄖ߆ࠄ⢝ఽ⢷⣧ౝߦ↢ℂ⊛㘩Ⴎ᳓ࠍᵈ
౉㧕ߢߪ‫ޔ‬ታ㛎ᓟߦ߽ᅧᆼ߇ᱜᏱߦ⛮⛯ߐࠇ‫ޔ‬ᱜᏱߦᣂ↢ఽ߇↢߹ࠇߚ‫ߩߘޕ‬ᓟߘߩߎߤ߽ߪᱜᏱߦ⊒⢒ߒߡ޿ࠆ‫ޕ‬
੹ᐕᐲߪ⛮⛯ߒߡ⢝ఽ᳇▤㐽Ⴇⴚߩ੍஻ታ㛎ࠍⴕߞߚ‫ޕ‬⢝੿᳇▤㐽Ⴇⴚߦߪ⢝੿ߩ᳇▤㐽Ⴇߩߚ߼వ┵ߦࡃ࡞࡯
ࡦߩߟ޿ߚ⢝ఽ↪ߩౝⷞ㏜ࠍ↪޿ࠆ‫ߩߎޕ‬ౝⷞ㏜ߩ૏⟎ߪᠲ૞ਛߦሶችౝߢᲣ૕࡮⢝੿߳࠳ࡔ࡯ࠫࠍਈ߃ߥ޿ߚ߼
ߦᄢᄌ㊀ⷐߢ޽ࠆ‫ޕ‬
ߘߎߢᚒ‫ߪޘ‬ሶችౝߢߩౝⷞ㏜ߩ૏⟎ࠍ⿥㖸ᵄ↹௝਄ߢ⋡⊛ㇱ૏߳࠽ࡆࠥ࡯࠻ߔࠆࠪࠬ࠹ࡓࠍ㐿⊒ߒߚ‫ޕ‬
ߎߩࠪࠬ࠹ࡓߣታ㓙ߩⴚᑼߦ↪޿ࠆౝⷞ㏜ࠍ૶↪ߒߡᅧᆼࠨ࡞߳ߩౝⷞ㏜૏⟎⏕⹺ߩ੍஻ታ㛎ࠍⴕߞߚ‫ޕ‬
ᰴᐕᐲߪሶችౝౝⷞ㏜࠽ࡆࠥ࡯࡚ࠪࡦࠪࠬ࠹ࡓቢᚑߣ᳇▤ౝ㐽Ⴇⴚߩታᣉࠍ੍ቯߒߡ޿ࠆ‫ޕ‬
B-63 ࠴ࡦࡄࡦࠫ࡯ߩญ⣧ౝ⁁ᘒߩ⺞ᩏ㧦߁ⲁ࡮ᱤߩ៺⠻࡮ᱤ๟Ἳ࡮ིߺวࠊߖߩ⹏ଔࠍਛᔃߦ
᩶੗଻ሶ㧘⧎↰ାᒄ㧘ዊᎹඅ㧘੗Ꮉ⍮ሶ㧘㥲⮮ᷤ㧔㢬⷗ᄢ࡮ᱤ㧕 ᚲౝኻᔕ⠪㧦ችㇱ⾆ሶ
࠴ࡦࡄࡦࠫ࡯11 ୘૕ 342 ᱤߦኻߒߡᱤ⑼ᬌ⸻ࠍታᣉߒߚ‫ߩߘޕ‬ౝ߁ⲁᱤߪ 16 ᱤ‫ޔ‬༚ᄬᱤߪ 3 ᱤߢ޽ߞߚ‫ߞࠃޕ‬
ߡ‫⚻ⲁ߁ޔ‬㛎ᱤࠍᜰߔ DMF ᱤߪ 19 ᱤ‫ޔ‬DMF ᜰᢙߪ 1.45 ߢ޽ߞߚ‫ޕ‬ᱤ⡺Ḵߩᷓߐߪ‫ޔ‬342 ᱤਛ 317 ᱤ߇ 4 mm એ
ਅߢ޽ߞߚ‫ޕ‬ᱤ๟ࡐࠤ࠶࠻᷹ቯᤨߦ಴ⴊࠍ⹺߼ߥ߆ߞߚߩߪ 6 ୘૕‫ޔ‬േំᱤࠍ⹺߼ߥ߆ߞߚߩߪ 8 ୘૕ߢ޽ߞߚ‫ޕ‬
⪺ߒ޿ࡊ࡜࡯ࠢߩ⫾Ⓧߣᱤ⍹ߩᴉ⌕߇ 9 ୘૕ߦ⹺߼ࠄࠇߚ‫ޔߚ߹ޕ‬ᐕ㦂ߦᔕߓߡో㗶⊛ߦ㗼⪺ߥຏ⠻ࠍ⹺߼ߚ‫ޕ‬
߁ⲁᱤߪߘߩ߶߷ోߡߦ⎕᛬ࠍ⹺߼ߚ‫ߜ߁ߩߘޕ‬೨ᱤߪ 11 ᱤߢ޽ࠅ‫ޔ‬⎕᛬࡮߁ⲁᱤߪ೨ᱤㇱߦ㓸ਛߒߡ޿ࠆ‫ޕ‬
ࠃߞߡ‫ߩⲁ߁ޔ‬ේ࿃ߪᄖ்ߦ⿠࿃ߔࠆߣ⠨߃ࠄࠇࠆ‫ޕ‬ᱤ⡺Ḵߩᷓߐ߇ 4 mm એਅߢ޽ࠆᱤߪో૕ߩ 92.7 㧑ߢ޽ࠅ‫ޔ‬
ߘߩ߶ߣࠎߤ߇᷹ቯᤨߩ಴ⴊࠍ⹺߼ߥ߆ߞߚ‫ޕ‬ᷓߐ 4 mm ߩᱤ⡺Ḵߪஜᐽߥᱤ⡺ߢ޽ࠆߣផኤߔࠆ‫⃻ޕ‬࿷߹ߢߦᬌ
⸻ߒߚ୘૕ߩ߁ⲁߣᱤ๟∔ᖚ߆ࠄ⷗ࠆญ⣧ஜᐽ⁁ᘒߪ‫ޔ‬ญ⣧ⴡ↢ߦ㑐ߔࠆ੺౉ߪ⊝ήߢ޽ࠅ‫ߣࠢ࡯࡜ࡊޔ‬ᱤ⍹ߩᄙ
㊂ߩᴉ⌕ࠍᢔ⷗ߔࠆߦ߽߆߆ࠊࠄߕᭂ߼ߡ⦟ᅢߣ޿߁ߎߣ߇ߢ߈ࠆ‫ߩߎޕ‬ℂ↱ߣߒߡ‫⎇ᧄޔ‬ⓥᚲߦ߅ߌࠆ࠴ࡦࡄࡦ
ࠫ࡯߇ 100 ຠ⋡ࠍ⿥߃ࠆήടᎿߩࡃ࡜ࡦࠬߩ⦟޿㘩㙄ࠍขߞߡ޿ࠆ੐ߦ⌕⋡ߒߡ޿ࠆ‫ޕ‬
߹ߚ‫ޔ‬ណขߒߚࡊ࡜࡯ࠢౝߩ⚦⩶ฌߦߟ޿ߡ DNA ⸃ᨆߦࠃࠆࡇࡠࠪ࡯ࠤࡦࠬᴺࠍ↪޿ߚ⚿ᨐ‫ᧂޔ‬⍮ߩ⚦⩶߇ሽ
࿷ߔࠆน⢻ᕈ߇␜ໂߐࠇ‫ޔ‬Streptococcus troglodytae ߣ๮ฬߒߚ‫ޕ‬
㧨⊒⴫᭎ⷐ㧪
࠴ࡦࡄࡦࠫ࡯ߩญ⣧ౝ⁁ᘒߩ⺞ᩏ㧦߁ⲁ࡮ᱤߩ៺⠻࡮ᱤ๟Ἳ࡮ིߺวࠊߖߩ⹏ଔࠍਛᔃߦ
᩶੗଻ሶ㧘⧎↰ାᒄ㧘ዊᎹඅ㧘㊁᧛⟵᣿㧘੹੗ᅑ㧘ጟᧄ౏ᓆ㧘੗Ꮉ⍮ሶ㧘㥲⮮ᷤ㧘ችਯේ⌀↱㧘═㑆ᘕᄥ㇢㧘ጊ
ญ⾆ᄩ㧘㒙଻஻ሶ㧔㢬⷗ᄢ࡮ᱤ㧕㧘ችㇱ⾆ሶ㧘෹᳗㓷Ꮖ㧔㔤㐳⎇㧕
ญṶ
1) ‫࡯ࠫࡦࡄࡦ࠴ޟ‬ญ⣧↱᧪ߩࡒࡘ࡯࠲ࡦࠬ࡟ࡦࠨ⃿⩶᭽⚦⩶ߦ㑐ߔࠆ⎇ⓥ‫ޠ‬㒙଻஻ሶ㧘߶߆㧔㢬⷗ᄢቇᱤቇળ╙
73 ࿁଀ળ㧘㢬⷗ᄢቇળ㙚ࡔࠗࡦࡎ࡯࡞㧘2011.6.25㧕
2) ‫ޟ‬੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲߩ࠴ࡦࡄࡦࠫ࡯ߩญ⣧ஜᐽ⁁ᘒߣߘߩ߁ߜ 1 ୘૕ߦኻߔࠆᱤ⑼ಣ⟎ߦߟ޿ߡ‫ޠ‬㥲⮮ᷤ㧘
ጊญ⾆ᄩ㧘᩶੗଻ሶ㧔ᣣᧄᱤ⑼଻ሽቇળ 2011 ᐕᐲᤐቄቇⴚᄢળ㧔╙ 134 ࿁㧕
㧘᧲੩ࡌࠗ⥰ᵿࡎ࠹࡞ ࠢ࡜ࡉ
࡝࠱࡯࠻㧘2011.6.10㧘p.88.㧕
3) ‫࡯ࠫࡦࡄࡦ࠴ޟ‬ญ⣧↱᧪ߩ࡟ࡦࠨ⃿⩶ߩᕈ⁁ߦ㑐ߔࠆ⎇ⓥ‫ޠ‬ችਯේ⌀↱㧘߶߆㧔╙ 60 ࿁ᣣᧄญ⣧ⴡ↢ቇળ㧘
ᣣᧄᄢቇ᧻ᚭᱤቇㇱ㧘2011.10.8-10㧘O-25㧕
㧔ญ⣧ⴡ↢ቇળ㔀⹹ ╙ 61 Ꮞ╙ 4 ภ p.435㧕
ࡐࠬ࠲࡯
1) Okamoto M, et al. Pyrosequencing Analysis of Oral Flora isolated from Chimpanzees. The 45th Meeting of The
Continental European Division of the International Association for Dental Researchthe (CED-IADR), Budapest,
Hungary, August 31- September 3,2011. (No. 469) (࿖㓙ᱤ⑼⎇ⓥቇળ࡛࡯ࡠ࠶ࡄㇱળ), Budapest, Hungary,
August 31- September 3, 2011.http://iadr.confex.com/iadr/ced11/webprogram/Paper151705.html
㧙㧙
2)
Okamoto M, et al.Preliminary Analysis of Oral Flora isolated from Chimpanzee. XIII, International Congress of
Bacteriology and Applied Microbiology㧔࿖㓙⚦⩶ቇળ㧕. 6-10 September 2011 (Sapporo, Japan).
⺰ᢥ
Okamoto M, et al. Streptococcus troglodytis sp. nov., from the Chimpanzee Oral Cavity. International Journal of Systematic
and Evolutionary Microbiology.Accepted 21 March 2012.
http://ijs.sgmjournals.org/content/early/2012/03/19/ijs.0.039388-0.abstract?cited-by=yes&legid=ijs;ijs.0.039388-0v1
(3) ৻⥸ࠣ࡞࡯ࡊ⎇ⓥ
C-1 㑆ធ⊛ᜰᮡࠍ↪޿ߚ಄᷷Ꮺᨋߦ߅ߌࠆ࠾ࡎࡦࠩ࡞↢ᕷ⁁ᴫߩផቯ
ᳯᚑᐢ᢯㧔ቝㇺችᄢ࡮ㄘ࡮㉿ጊ㧕㧘ဈ’ߪࠆ߆㧔ጤᚻᄢ࡮ㅪㄘ㧕 ᚲౝኻᔕ⠪㧦ᷰㆻ㇌ᄦ
࠾ࡎࡦࠩ࡞ߩ◲ଢߥ୘૕⟲ࡕ࠾࠲࡝ࡦࠣᚻᴺߩ㐿⊒ࠍ⋡⊛ߦ‫ޔ‬1)ࠨ࡞᮸⊹㘩ߦࠃࠆᧁᧄᬀ‛ߩ᮸ᒻᄌൻ‫ޔ‬2)ࠨ࡞
♮ࠍㆬᛯ⊛ߦ೑↪ߔࠆ♮⯻ߩ↢ᕷኒᐲ‫ ߩޔ‬2 ߟࠍࠨ࡞↢ᕷᢙߩ㑆ធᜰᮡߣߒߡߩ᦭ലᕈࠍ⹏ଔߒߚ‫ޕ‬1)ߩ⚿ᨐ‫ߎޔ‬
ࠇ߹ߢႎ๔߇޽ߞߚࡗࡑࠣࡢએᄖߦ‫ޔ‬㜞ᧁߢ޽ࠆࡎࠝࡁࠠߦ߅޿ߡ߽㗼⪺ߥ᮸ᒻᄌൻ߇⏕⹺ߐࠇߚ‫ౕޕ‬૕⊛ߦߪ‫ޔ‬
ࠪࡘ࡯࠻㐳ߩᷫዋߣ‫ᧄ࠻࡯ࡘࠪޔ‬ᢙߩჇടߢ޽ࠆ‫ߩ࡞ࠨޔߚ߹ޕ‬ㆬᛯᕈߩ㜞޿࠷࡞ᕈᧁᧄᬀ‛ߦ߅޿ߡ߽‫ޔ‬᣿⏕ߥ
᮸ᒻᄌൻߪ⏕⹺ߐࠇߚ‫ޕ‬2)ߩ⚿ᨐ‫ޔ‬㑆ធᜰᮡߣߒߡኻ⽎ߣߒߚ Aphodius spp.ߪ‫ޔ‬ᄖㇱⅣႺ㧔᳇᷷╬㧕ߩᓇ㗀߽ᄢ߈
ߊ‫↢࡞ࠨޔ‬ᕷಽᏓ࡮ኒᐲߩ⚻ᤨ⊛ߥᄌൻࠍ⷗ࠆᜰᮡߣߒߡ೑↪ߔࠆ㓙ߪ‫ߚߒ߁ߎޔ‬ᄖㇱⅣႺߦኻߔࠆචಽߥ㈩ᘦ߇
ᔅⷐߢ޽ࠆߎߣ߇⠨߃ࠄࠇߚ‫੹ޕ‬ᓟߩ⺖㗴ߣߒߡ‫ޔ‬1)ߣ 2)ߦ↪޿ߚ㑆ធᜰᮡࠍ‫࡞࡯ࠤࠬߩߤޔ‬㧔⸃௝ᐲ㧕ߩࠨ࡞ࡕ
࠾࠲࡝ࡦࠣᜰᮡߣߒߡ೑↪ߔࠆߎߣ߇วℂ⊛ߢ޽ࠆ߆ࠍ⹏ଔߔࠆߚ߼ߩ⺞ᩏ⎇ⓥ߇ᔅⷐߢ޽ࠆߎߣ߇⠨߃ࠄࠇߚ‫ޕ‬
C-2 㔤㐳㘃‫࡮ⷡⷞޔ‬๧ⷡߩ GPCR ဳฃኈ૕ߦኻߔࠆ⿒ᄖಽశ⎇ⓥ
␹ข⑲᮸㧘 ጊ⠹ᄢ㧔ฬᎿᄢ࡮㒮Ꮏ㧕㧘ฎ⼱␭⹖㧘ᧁ᧛ືዞ㧔ಽሶ⎇࡮↢๮㍲૕㧕 ᚲౝኻᔕ⠪㧦੹੗໪㓶
ࡅ࠻ࠍ฽߻㔤㐳㘃ߩ✂⤑ߦሽ࿷ߔࠆ 3 ⒳㘃㧔⿒࡮✛࡮㕍㧕ߩ⦡ⷡⷞ‛⾰ߪ⹜ᢱ⺞⵾߇࿎㔍ߥߚ߼‫ޔ‬X ✢⚿᥏᭴ㅧ
⸃ᨆࠍ฽߻᭴ㅧ↢‛ቇ⊛⸃ᨆߪㆊ෰ߦ଀߇ߥߊ‫ޔ‬ᚒ‫⁁ߥ߁ࠃߩߘޕߚߞ޽ߢ߹߹ߩ⻘ߪࡓ࠭࠾ࠞࡔ⼂⹺⦡ߩޘ‬ᴫߩ
߽ߣ‫ޔ‬ᚒ‫ߪޘ‬ᢙᐕ೨߆ࠄᧄ౒ห⎇ⓥࡊࡠࠫࠚࠢ࠻ࠍᵴ↪ߒߡ‫ޔ‬㜞♖ᐲߩૐ᷷⿒ᄖಽశࠍ↪޿ߚ㔤㐳㘃⿒࡮✛ᗵฃᕈ
ⷞ‛⾰ߩ᭴ㅧ⸃ᨆߦ᜸ᚢߒߚ‫⚦ࡦࠟ࠻ࡅޕ‬⢩↱᧪ߩၭ㙃⚦⢩ࠍ↪޿ߡ⹜ᢱࠍ⺞⵾ߔࠆ৻ᣇ‫⹜ߥ߆ߕࠊޔ‬ᢱߢ߽⸘᷹
น⢻ߥᏅࠬࡍࠢ࠻࡞᷹ቯ♽ࠍᦨㆡൻߔࠆߎߣߢ‫ޔ‬ᐔᚑ 21 ᐕᐲߦ਎⇇ೋߣߥࠆ࠺࡯࠲ࠍႎ๔ߒߚ㧔Katayama et
al.2010;NHK ࠾ࡘ࡯ࠬߣ 35 ⚕ߩᣂ⡞ߢ⚫੺㧕
‫ߦࠄߐޕ‬ᐔᚑ 22 ᐕᐲߦߪࠬࡍࠢ࠻࡞ᐲࠍ਄ߍࠆߎߣߢ‫⾰ࠢࡄࡦ࠲ޔ‬ౝ
ㇱߦ⚿วߒߚ᳓ಽሶߩหቯߦᚑഞߒ‫ޔ‬ᐔᚑ 23 ᐕᐲߦ⺰ᢥ⊒⴫ߔࠆߎߣ߇ߢ߈ߚ㧔Katayama et al.2012;1 ⚕ߩᣂ⡞ႎ
㆏㧕
‫ޕ‬
ᦨೋߩ⺰ᢥߦࠃࠇ߫‫✛࡮⿒ޔ‬ᗵฃᕈⷞ‛⾰ߩ᭴ㅧߪ㛳ߊ߶ߤ㘃ૃߒߡ޿ߚ߇‫ޔ‬᳓ಽሶߩ♖ኒ᷹ቯߩ⚿ᨐ‫ޔ‬ౝㇱ⚿
ว᳓ߩ᳓⚛⚿วⅣႺߪ⿒࡮✛ᗵฃᕈⷞ‛⾰ߢ⇣ߥࠆߎߣ߇᣿ࠄ߆ߦߥߞߚ‫⥝ޕ‬๧ᷓ޿ߎߣߦ‫ⷰޔ‬᷹ߐࠇߚ᳓ಽሶߩ
ᝄേᢙߪⷞ‛⾰ߩๆ෼ᵄ㐳ߣ⋧㑐ࠍ␜ߒ‫ޔ‬ౝㇱ⚿ว᳓߇⦡⼂೎ߩࡔࠞ࠾࠭ࡓߦ㑐ࠊࠆߎߣ߇␜ໂߐࠇߚᐔᚑ 23 ᐕ
ᐲߦߪ⦡ⷡⷞ‛⾰ߩᄌ⇣૕ታ㛎߽㐿ᆎߔࠆߎߣ߇ߢ߈ߚߚ߼‫⸃ࡓ࠭࠾ࠞࡔߥ⚦⹦ࠅࠃޔ‬ᨆ߇ᦼᓙߐࠇࠆ‫ޕ‬
߹ߚᐔᚑ 23 ᐕᐲߦߪⷞ‛⾰ߣหߓ G Ⱞ⊕⾰౒ᓎဳฃኈ૕ߢ޽ࠆ๧ⷡฃኈ૕ߩ᭴ㅧ⸃ᨆࠍ⸘↹ߒ‫ోޔ‬෻኿⿒ᄖಽ
శ⸘᷹ߦࠃࠆ᷹ቯ♽ߩ᭴▽ࠍ㐿ᆎߒߡ޿ࠆ‫ޕ‬ኾ㐷ᕈߩ޽ࠆ⎇ⓥᚲߦ߅޿ߡ߽⇣ಽ㊁Ⲣว߇㊀ⷞߐࠇࠆᤓ੹‫౒ᧄޔ‬ห
⎇ⓥߪ‛ℂൻቇ/ಽሶ⑼ቇಽ㊁ߣߩℂᗐ⊛ߥ⇣ಽ㊁Ⲣวߢ޽ࠆߣ૏⟎ઃߌࠆߎߣ߇ߢ߈ࠆ‫ޕ‬ᡰេ޿ߚߛ޿ߡ޿ࠆ㔤
㐳⎇ߦᡷ߼ߡ⻢ᗧࠍ⴫ߒߚ޿‫ޕ‬
㧨ේ⪺⺰ᢥ㧪
Katayama K, et al. (2012) Protein-Bound Water Molecules in Primate Red- and Green-Sensitive Visual Pigments.
Bicochemistry 51:1126-113.
C-3 ࠾ࡎࡦࠩ࡞ߩੱᎿ❥ᱺࠍ⋡ᜰߒߚᛛⴚ㐿⊒
㜞ᳯᵮ᣹㧔ᧅᏻᏒ౞ጊേ‛࿦㧕㧘᳗㊁᣽ᔒ㧘ർ಴ᵏਯ㧘ဈญ⻞৻㇢㧔ർᶏ㆏ᄢ࡮₞ක㧕㧘દ⮮⌀ノ㧘⍹ᯅ૓ⷙ㧘ᦺୖ
ථ਽㧘ዊᨋ⌀਽㧔ᧅᏻᏒ౞ጊേ‛࿦㧕 ᚲౝኻᔕ⠪㧦੹੗໪㓶
ᧅᏻᏒ౞ጊേ‛࿦ߢ㘺⢒ߔࠆ࠾ࡎࡦࠩ࡞ߪ‫ߦࠬࠝߩߡోޔ‬ኻߒߡ♖▤⚿⚤ಾ㒰ಣ⟎ࠍⴕ޿‫❥ޔ‬ᱺ೙㒢ࠍታᣉߒߡ
޿ࠆ‫ޔࠄ߇ߥߒ߆ߒޕ‬㘺⢒୘૕ߩ㜞㦂ൻ߇ㅴߺ‫ޔ‬዁᧪⊛ߥ㘺⢒⟲⛽ᜬߩߚ߼‫ޔ‬ᣂߚߥ❥ᱺ▤ℂᣇᴺߣߒߡੱᎿ❥ᱺ
ࠍᬌ⸛ߒߡ޿ࠆ‫ޕ‬
ᧄ࿦㘺⢒࠾ࡎࡦࠩ࡞ߩࠝࠬ 4 㗡߆ࠄ♖ሶណขࠍ⹜ߺߚ‫♖ޕ‬Ꮍ਄૕የㇱ߆ࠄᵈ኿㊎ࠍ↪޿ߡ⓫ೝๆᒁߒ‫ޔ‬4 㗡ߔߴ
ߡ߆ࠄ♖ᶧࠍណขߒߚ‫ޕ‬4 㗡ਛ 3 㗡ߩ♖ᶧਛߩ♖ሶߪోߡᱫṌߒߡ߅ࠅ‫ޔ‬ᱷࠆ 1 㗡ߢߪ♖ሶ߇⏕⹺ߢ߈ߥ߆ߞߚ‫ޕ‬
ᤓᐕᐲߩห᭽ߩಣ⟎ߢߪᵴ⊒ߥ↢߈ߚ♖ሶࠍណขߒߡ޿ࠆߚ߼‫ޔ‬ណขᚻᛛ‫ޔ‬ណขᤨᦼ෸߮ណข୘૕ߦߟ޿ߡᬌ⸛߇
ᔅⷐߢ޽ࠆߣ⠨߃ࠄࠇࠆ‫ޕ‬
㔤㐳㘃⎇ⓥᚲ㘺⢒࠾ࡎࡦࠩ࡞ߩࠝࠬ 3 㗡߆ࠄߪ‫ޔ‬⢂㐷ߦ㔚ᭂ᫔ࠍᝌ౉ߒ‫ޔ‬㔚᳇ೝỗࠍⴕ߁ᣇᴺߢ♖ᶧណขࠍ⹜ߺ‫ޔ‬
߁ߜ 1 㗡߆ࠄᵴᕈߩ޽ࠆ♖ሶࠍណขߒߚ‫ޕ‬ណขߒߚ♖ሶࠍಓ⚿଻ሽߒ‫ޔ‬ᤓᐕᐲ㔤㐳㘃⎇ⓥᚲߦߡಓ⚿଻ሽߒߚ 1 ୘
૕ಽߩ♖ሶߣ૬ߖⲢ⸃ߒ‫ޔ‬ಓ⚿ᓟߩ♖ሶᕈ⁁ࠍᲧセߒߚߣߎࠈ‫ޔ‬ᤓᐕᐲߦᲧߴ‫੹ޔ‬ᐕᐲណขߒߚ♖ሶߩ↢ሽ₸
㧔2010㧦22㧑‫ޔ‬2011㧦1.5㧑㧕෸߮ㆇേᕈ㧔2010㧦+++15‫ޔ‬++15‫ޔ‬+10‫ޔ‬±10‫ޔ‬-50 2011㧦+++2.5‫ޔ‬++2.5‫ޔ‬+2.5‫ޔ‬±2.5‫ޔ‬
-90㧕߇ૐ޿⚿ᨐߣߥߞߚ‫⸃Ⲣޔߚ߹ޕ‬ᓟߦ߅޿ߡߘߩ↢ᱫߦ㑐ࠊࠄߕ‫ޔ‬ᄙߊߩ♖ሶ߇వ૕ࠍ༚ᄬߒߡ޿ߚ㧔2010㧦
36.5㧑‫ޔ‬2011㧦86㧑㧕‫ޕ‬
㧙㧙
ᧄᐕᐲߪᧄ࿦㘺⢒࠾ࡎࡦࠩ࡞߆ࠄ↢߈ߚ♖ሶ߇ណขߢ߈ߥ߆ߞߚߚ߼ੱᎿ᝼♖ታᣉߦ⥋ࠄߥ߆ߞߚ‫ޔߚ߹ޕ‬ಓ⚿
Ⲣ⸃♖ሶߩవ૕༚ᄬ߇⏕⹺ߐࠇ‫ޔ‬ಓ⚿♖ሶࠍ↪޿ߚੱᎿ᝼♖ᚑഞߩߚ߼ߦߪ♖ᶧណข߅ࠃ߮ಓ⚿ᚻᛛߩะ਄߇ᔅ㗇
ߢ޽ࠆߣ⠨߃ࠄࠇࠆ‫ޕ‬
C-4 ੑෆᕈ߰ߚߏ࠴ࡦࡄࡦࠫ࡯ߩⴕേ⊒㆐ߦ㑐ߔࠆᲧセ⹺⍮⊒㆐⎇ⓥ
቟⮮ኼᐽ㧔ᘮᙥ⟵Ⴖᄢ࡮ᢥ㧕㧘ጯᧄஜ㧔⡛ᔃᅚሶᄢ㧕㧘਄㊁᦭ℂ㧔ṑ⾐⋵┙ᄢ࡮ੱ㑆ᢥൻቇㇱ㧕㧘Ꮉ਄ᢥੱ㧔᧲੩Ꮏ
ᬺᄢ㧕
㧘⛚↰ବ๺㧘⑔቞ᦶ㧔㜞⍮⋵┙ߩ޿ߜേ‛౏࿦㧕 ᚲౝኻᔕ⠪㧦෹᳗㓷Ꮖ
㜞⍮⋵┙ߩ޿ߜേ‛౏࿦ߦ 2009 ᐕ 4 ᦬ 1 ߦ⺀↢ߒߚੑෆᕈߩ߰ߚߏ࠴ࡦࡄࡦࠫ࡯ߩⴕേ⊒㆐ߩㆊ⒟ࠍ‫ ᦬ޔ‬1 ࿁
❑ᢿ⊛ߦⷰኤߒ‫⊒ߩߏߚ߰ߩ࠻ࡅޔ‬㆐ㆊ⒟ߣᲧセߒߚ‫ߪߏߚ߰ޕ‬Უⷫߩߺߥࠄߕ‫․޿ߥߩ✼ⴊޔ‬ቯߩ㓽ᚑ૕ߣߘࠇ
ߙࠇⷫኒߥ㑐ଥࠍ▽߈‫ޔ‬Უⷫߩㆊᐲߥሶ⢒ߡ⽶ᜂߪ࿁ㆱߐࠇߡ޿ߚ‫ⴕߥ┙⁛ߚ߹ޕ‬േ߿ਗⴕㆆ߮ࠍߔࠆᤨ㑆߇ᄙ޿
߇‫ࠆࠇࠄߺߦ࠻ࡅޔ‬㘩ߴ‛ߩ⿅ਈߥߤߩ೑ઁ⊛ⴕേߪ⷗ࠄࠇߥ߆ߞߚ‫⋧ޕ‬੕ߩߓ߾ࠇว޿ߪᤨ‫ޔߒ↢⊒ޘ‬り૕ㆇേ
⢻ജ߇♖✺ߦߥࠆߦߟࠇߡ‫ߩߘޔ‬㗫ᐲࠍჇߒߡ޿ࠆࠃ߁ߦᕁࠊࠇߚ‫ߚ߹ޕ‬୘೎ߦㆆࠎߢ޿ࠆߣ߈ߢ߽‫ઁޔ‬ᣇߦኻߔ
ࠆᵈᗧ߇ቢోߦߥߊߥߞߡ޿ࠆࠊߌߢߪߥߊ‫৻ޔ‬ᣇ߇⒖േߔࠆߣㆃࠇߡઁᣇ߽หᣇะߦ⒖േߔࠆߎߣ߇ߒ߫ߒ߫ⷰ
ኤߐࠇߚ‫ޕ‬
㧨ቇળ⊒⴫㧪
1) ቟⮮ኼᐽ㧘߶߆ Ყセ෺↢ఽቇߩ⹜ߺ㧙ᐜఽᦼߩࡅ࠻ߣ࠴ࡦࡄࡦࠫ࡯ߩേ↹௝߆ࠄ㧚╙ 25 ࿁ᣣᧄ෺↢ఽ⎇ⓥ
ቇળቇⴚ⻠Ṷળ㧚ᐔᚑ 23 ᐕ 1 ᦬ 29 ᣣ㧘᧲੩㧚
2) ቟⮮ኼᐽ‫޿ߛ߁ࠂ߈ߩ࡯ࠫࡦࡄࡦ࠴ ߆߶ޔ‬㑐ଥ㧙Ყセ෺↢ఽቇߩ⹜ߺ(2)㧚╙ 26 ࿁ᣣᧄ෺↢ఽ⎇ⓥቇળቇⴚ
⻠Ṷળ㧚ᐔᚑ 24 ᐕ 1 ᦬ 28 ᣣ㧘᧲੩㧚
C-5 ࠴ࡦࡄࡦࠫ࡯ߩⷞⷡ࡮ᵈᗧߦ㑐ߔࠆᲧセ⹺⍮⎇ⓥ
‐⼱ᥓ৻㧔ජ⪲ᄢ࡮ᢥቇㇱ㧕㧘ᓟ⮮๺ብ㧔੩ㇺᄢ࡮↢๮⑼ቇ♽ࠠࡖ࡝ࠕࡄࠬᒻᚑ࡙࠾࠶࠻㧕 ᚲౝኻᔕ⠪㧦෹᳗㓷
Ꮖ
ᧄ⎇ⓥߪ‫ⷡⷞߩ࡯ࠫࡦࡄࡦ࠴ޔ‬ಣℂࠍࡅ࠻ߩߘࠇߣᲧセߔࠆߎߣߢ‫ޔ‬ਔ⠪ߦ߅ߌࠆ౒ㅢὐߣ⋧㆑ὐ߆ࠄ‫߅ⷡⷞޔ‬
ࠃ߮ⷞⷡ⊛ᵈᗧߩㅴൻ⊛ⷐ࿃ࠍ⸃᣿ߔࠆߎߣࠍ⋡⊛ߣߒߡ޿ࠆ‫ߦⷡⷞޕ‬㑐ߔࠆ⎇ⓥߢߪ‫ޔ‬ೝỗ↹௝ߦ฽߹ࠇࠆࡅ࠻
߹ߚߪ࠴ࡦࡄࡦࠫ࡯ߩⷞ✢ᣇะߩᑯ೎ࠍߔࠆ႐ว‫✢ⷞޔ‬వߩ‛૕ߩ᦭ήߢ㔍ᤃᐲ߇ߤ߁ᄌൻߔࠆ߆ࠍᬌ⸛ߒߚ‫ࡅޕ‬
࠻ᚑੱⵍ㛎⠪ࠍ↪޿ߚ੍஻ታ㛎ࠍ߅ߎߥߞߚ߇‫‛ޔ‬૕ߩ᦭ήߦࠃࠆⷞ✢ᣇะߩᑯ೎㔍ᤃᐲߦ㑐ߔࠆ᦭ᗧߥᏅ߇⷗ࠄ
ࠇߥ߆ߞߚ‫੹ޕ‬ᓟ‫࠻ࡅޔ‬ᚑੱߢೝỗ߅ࠃ߮⺖㗴ᚻ⛯߈ࠍౣᬌ⸛ߒߚ਄ߢ‫ⵍࠍ࡯ࠫࡦࡄࡦ࠴ޔ‬㛎૕ߣߔࠆᬌ⸛ࠍ߅ߎ
ߥ߁‫ޕ‬
ᵈᗧߦ㑐ߒߡߪ‫ߩߢ߹ࠇߎޔ‬ታ㛎ߦࠃࠅ‫↹ޔ‬㕙਄ߩ‛૕ߣ޿ߞߚ‫ࠍޠࠅ߹ߣ߹ޟ‬න૏ߦ⾮ᵴߔࠆࠃ߁ߥⷞⷡ⊛ᵈ
ᗧㆊ⒟㧔ࠝࡉࠫࠚࠢ࠻ࡌ࡯ࠬߩᵈᗧ㧕߇࠴ࡦࡄࡦࠫ࡯ߦ߽޽ࠆߎߣ߇᣿ࠄ߆ߦߥߞߚߎߣ߆ࠄ‫੹ޔ‬࿁ߪ‫‛ޔ‬૕ߩᒻ
⁁߇ᵈᗧߩ⾮ᵴߦߤ߁ᓇ㗀ߔࠆ߆ࠍᬌ⸛ߒߚ‫‛ޕ‬૕ߩᒻ⁁ࠍᠲ૞ߒ‫ޔ‬ᗐቯߐࠇࠆᵈᗧߩ⒖േ⚻〝ߪ⇣ߥࠆ߇‫ޔ‬⒖േ
రߣ⒖േవߩ⋥✢〒㔌ࠍ⛔೙ߒߚೝỗࠍ↪ᗧߒߡ‫⃻ޔ‬࿷࠹ࠬ࠻ࠍታᣉਛߢ޽ࠆ‫ޕ‬ᒁ߈⛯߈‛૕ߩᒻ⁁ߩᓇ㗀ࠍ⺞ߴ‫ޔ‬
੹ᓟߪࠃࠅⶄ㔀ߥⷞⷡ㘑᥊਄ߩೝỗዻᕈ߇ߤߩࠃ߁ߦ࠴ࡦࡄࡦࠫ࡯ߩⷞⷡ⊛ᵈᗧࠍ᝝ᝒߔࠆ߆⸃᣿ߒߡ޿ߊ੍ቯ
ߢ޽ࠆ‫ޕ‬
C-㧣 㔤㐳㘃ߦ߅ߌࠆ⢝ఽᦼ߆ࠄᕁᤐᦼ߹ߢߩ⣖ᒻᘒߩ⊒㆐⊛ᄌൻߦ㑐ߔࠆᲧセ⎇ⓥ
㈬੗᦮ሶ㧔੩ㇺᄢ࡮ℂ࡮↢‛⑼ቇ㧕㧘ਛോ⌀ੱ㧘࿖᧻⼾㧘Ꮞፉ⟤ᐘ㧔੩ㇺᄢ࡮ℂ㧕㧘ጊ↰㊀ੱ㧔੩ㇺᄢ࡮ක㧕㧘⮮Ỉ
㆏ሶ㧔੩ㇺᄢ࡮㊁↢േ‛㧕㧘㡻Ლବผ㧔࠴ࡦࡄࡦࠫ࡯࡮ࠨࡦࠢ࠴ࡘࠕ࡝࡮ቝ࿯㧕 ᚲౝኻᔕ⠪㧦෹᳗㓷Ꮖ
ᐔᚑ 23 ᐕᐲߩ౒ห೑↪⎇ⓥߢߪ‫ޔ‬ᒰೋߩ⸘↹ㅢࠅ‫ޔ‬3 ᰴర⏛᳇౒㡆↹௝ᴺ㧔MRI㧕ࠍ↪޿ߡ‫⧯ޔ‬ᚑ૕ᦼࠍㄫ߃ߚ
㔤㐳㘃⎇ⓥᚲߩ࠴ࡦࡄࡦࠫ࡯㧔11 ᱦ㧕ߩ⣖ᒻᘒߦ㑐ߔࠆ❑ᢿ⊛⸘᷹ࠍⴕߞߚ‫ޕ‬
߹ߚ‫ޔ‬㔤㐳㘃⎇ⓥᚲߩ࠴ࡦࡄࡦࠫ࡯ߩ↢ᓟ 6 ࡩ᦬߆ࠄ 6 ᱦߦ߅ߌࠆ೨㗡೨㊁ߩ⊒㆐ߦᵈ⋡ߒߚ⎇ⓥ߇‫࠻ࡦ࡟ࠞޔ‬
ࡃࠗࠝࡠࠫ࡯⹹㧔2011 ᐕ‫ޔ‬21 Ꮞ‫ޔ‬1397-1402 㗁㧕ߩේ⪺⺰ᢥߣߒߡ⸥タߐࠇߚ‫ߩ࡯ࠫࡦࡄࡦ࠴ޕ‬೨㗡೨㊁ߩ⊕⾰ኈ
Ⓧߪ‫ߣ࠻ࡅޔ‬ห᭽ߦ‫⊒ޔ‬㆐ᦼ㑆߇ઁߩ㗔ၞߦᲧߴ‫ߦ․ޔ‬ᑧ㐳ߐࠇߡ޿ߚ‫ߩ࡯ࠫࡦࡄࡦ࠴ޔࠄ߇ߥߒ߆ߒޕ‬೨㗡೨㊁
ߩ⊕⾰ኈⓍߩჇട₸ߪ‫ߊߒ⪺߽ࠅࠃ࠻ࡅޔ‬ૐߊ‫ޔ‬೨㗡೨㊁ߩ␹⚻ㅪ⚿ߩᒝൻ߇ࡅ࠻ߦᲧߴᒙ޿ߎߣ߇␜ໂߐࠇߚ‫ޕ‬
⸒޿឵߃ࠇ߫‫ߩ࠻ࡅޔ‬೨㗡೨㊁ߩ␹⚻ㅪ⚿ߩ⪺ߒ޿ᒝൻ߇‫࠻ࡅޔ‬࿕᦭ߩࠃࠅⶄ㔀ߥ␠ળᕈߩ⊒㆐߿⚻㛎ߦၮߠߊ⍮
⼂࡮ᛛⴚߩ₪ᓧߦነਈߒߡ޿ࠆߎߣ߇␜ໂߐࠇߚ‫ޕ‬
ࡅ࠻ߢߪ‫ޔ‬೨㗡೨㊁ߥߤߩ㜞ᰴߩ⣖㗔ၞߦ߅޿ߡᕁᤐᦼߦ઻߁േ⊛ߥ⊒㆐⊛ᄌൻ߇␜ߐࠇࠆߎߣ߇ႎ๔ߐࠇߡ޿
ࠆ‫ޔ߽ߡ޿߅ߦ࡯ࠫࡦࡄࡦ࠴ޕ‬ᕁᤐᦼߦ઻߁ᚑ㐳ࠬࡄ࡯࠻߇␜ߐࠇࠆߩߛࠈ߁߆‫⊒ߥ߁ࠃߩߘޔߪ޿ࠆ޽ޕ‬㆐᭽ᑼ
ߪࡅ࠻࿕᦭ߩ߽ߩߢ޽ࠅ‫ߥ⊛⇣․࠻ࡅޔ‬⣖᭴ㅧࠍᒻᚑߔࠆⷐ࿃ߩ৻ߟߣߥࠆߩߢ޽ࠈ߁߆‫੹ޕ‬ᓟ߽‫ޔ‬ᕁᤐᦼ‫࠻ࠝޔ‬
࠽ᦼ߳ߣ⛯ߊ࠴ࡦࡄࡦࠫ࡯ߩ⣖ᒻᘒߩ⊒㆐⊛ᄌൻࠍㅢߒߡ‫ߩ࠻ࡅޔ‬⣖ߩㅴൻ⊛↱᧪ࠍߐࠄߦ⸃᣿ߒߡ޿ߊ੍ቯߢ޽
ࠆ‫ޕ‬
C-10 ደਭፉૐ࿾ᨋߩ࠾ࡎࡦࠩ࡞㊁↢⟲ߦ߅ߌࠆ㧞⟲หᤨㅊ〔ߦࠃࠆ⟲㑆㑐ଥߩᬌ⸛
㋈ᧁṑ㧔㦖⼱ᄢ࡮࿖㓙ᢥൻ㧕㧘⮮↰ᔒᱠ㧔㣮ఽፉᄢ࡮ㄘ㧕㧘ਅጟࠁ߈ሶ㧔Ꮲ੩⑼ቇᄢ࡮↢๮ⅣႺ㧕㧘᧖ᶆ⑲᮸㧔੩ㇺ
ᄢ࡮㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯㧕 ᚲౝኻᔕ⠪㧦ඨ⼱๋㇢
⟲㑆㑐ଥ߇┹ว⊛ߢ޽ࠆደਭፉૐ࿾ߩ࠾ࡎࡦࠩ࡞୘૕⟲ߦ߅ߌࠆ‫ޔ‬㓞ធ⟲ߩᣣᏱ⊛ߥᓇ㗀ࠍ‫ޔ‬ㆆേၞ೑↪‫ޔ‬㖸ჿ
ࠦࡒࡘ࠾ࠤ࡯࡚ࠪࡦ‫␠ߩࠬࡔࠬࠝ ޔ‬ળ㑐ଥ߆ࠄᬌ⸛ߔࠆ‫ޔ߼ߚߩߘޕ‬㓞ធ⟲ߣߩ౒᦭ၞ߿⟲ࠇ㑆ߩⓨ㑆⊛〒㔌‫ޔ‬
㧙㧙
౒᦭㘩‛ࡄ࠶࠴ߥߤࠍ߼ߋߞߡ‫ޔ‬⒖േㅦᐲ߿‫ޔ‬ណ㘩ㅦᐲ‫ޔ‬㖸ჿ㗫ᐲ‫ޔ‬୘૕㑆〒㔌ߥߤ߇‫┹ߩߣ⟲ઁޔ‬วࠍᗧ⼂ߒߚ
߽ߩߦߥߞߡ޿ࠆ߆ߤ߁߆ࠍ⺞ߴߚ‫⺞ޕ‬ᩏᦼ㑆ߪ 2011 ᐕ 8 ᦬ 20 ᣣ߆ࠄㅪ⛯ 7 ᣣ㑆ߦ‫ޔ‬ደਭፉ⷏ㇱၞߩඨጊ࿾ၞߩ
㓞ធ 2 ⟲㧔AT ⟲ߣ E ⟲㧕ࠍ‫ޔ‬㧠ੱߩ⺞ᩏ⠪ߦࠃߞߡ‫ ࠇ⟲ߩࠇߙࠇߘޔ‬2 㗡ߕߟ㧔 ࠝࠬߣ ࡔࠬ㧕ߩหᤨ୘૕ㅊ
〔ࠍⴕߞߚ‫ޕ‬୘૕㑆〒㔌߿⒖േㅦᐲߪ‫⺞ߩࠇߙࠇߘޔ‬ᩏ⠪߇ GPS ࠍ៤Ꮺߒߡ୘૕ߩⓨ㑆⊛૏⟎ࠍ⸥㍳ߒߡಽᨆߒ
ߚ‫ⴕޕ‬േ㧔⒖േ࡮ណ㘩࡮ભᕷ࡮Ძߠߊࠈ޿ߥߤ␠ળ੤ᷤ㧕‫ᦨޔ‬ㄭធ୘૕‫ޔ‬ណ㘩ኻ⽎‫ޔ‬㖸ჿ╬ࠍ⸥㍳ߒߚ‫⺞ޕ‬ᩏᦼ㑆
ਛ㧔߭ߣߟߩ⟲ࠇߩⷰኤ߅ࠃߘ 58 ᤨ㑆⋧ᒰ㧕ߦ‫ࠇ⟲ޔ‬ห჻ߩ಴ળ޿߇৻࿁ⷰኤߐࠇ‫ޔ‬಴ળ߁೨ߩኻጺ⊛ߥ⁁ᴫߣ‫ޔ‬
৻ᣇߩ⟲ࠇߩᕆធㄭߣઁᣇߩ⟲ࠇߩㅌළ߇ߺࠄࠇ‫⺞ޔ‬ᩏኻ⽎⟲㑆ߩᢜኻ⊛㧔┹ว⊛㧕㑐ଥࠍ⏕⹺ߒߚ‫ޔߪߢ↹⸘ޕ‬
ߎߩᦼ㑆ߪ㕖⊒ᖱᦼߣߒߡ޿ߚ߇‫ޔ‬ታ㓙ߦߪᦼ㑆ਛߦ ᣇߩኻ⽎⟲ߩ ࡔࠬ߇⊒ᖱߒ‫ߣࠬࠝ ޔ‬㗫❥ߦ੤የࠍⴕߞ
ߚߚ߼‫ޔ‬੤የⴕേߦ⟲ࠇߩㆆേ߇ᓇ㗀ࠍ߁ߌߡ޿ߚ‫ޔߚ߹ޕ‬ណ㘩ኻ⽎ߣߒߡߪ‫ࠇ⟲ޔ‬㑆ߩ┹วߩⷐ࿃ߣߥࠆࠗ࠴ࠫ
ࠢዻࠕࠦ࠙ߩ⚿ታ᮸ࠍ㗫❥ߦ૶߁ߎߣ߇ᦼᓙߐࠇߡ޿ߚ߇‫ᦼ੹ޔ‬㑆ਛߪ⚿ታ߇߶ߣࠎߤߺࠄࠇߕ‫ࠇ⟲ޔ‬㑆߇಴ળ߁
㗫ᐲࠍਅߍߡ޿ߚน⢻ᕈ߇㜞޿‫ߚߒ߁ߎޕ‬ὐࠍ⠨ᘦߩ߁߃ߢ‫ߩ⚦⹦ߩ࠲࡯࠺ޔ‬ಽᨆߪ⛮⛯ਛߢ޽ࠆ‫ޕ‬
C-12 㔤㐳㘃ߦ߅ߌࠆឃෆߩ೙ᓮᯏ᭴ߦ㑐ߔࠆ⎇ⓥ
᧤᧛ඳሶ㧘೨ᄙᢘ৻㇢㧘ᄢ⬿⡡㧘਄㊁ጊ⾐ਭ㧔ฬᄢ࡮㒮࡮↢๮ㄘ㧕 ᚲౝኻᔕ⠪㧦㋈ᧁ᮸ℂ
ߍߞᱤ㘃ߣߪ⇣ߥࠅ‫ޔ‬㔤㐳㘃ߢߪࠛࠬ࠻ࡠࠫࠚࡦߦࠃࠆࠧ࠽࠼࠻ࡠࡇࡦߩᄢ㊂᡼಴߇㓽㓶ਔᣇߦ⷗ࠄࠇࠆߎߣࠍ
᣿ࠄ߆ߦߒߡ߈ߚ‫ߪߣߎߩߎޕ‬㔤㐳㘃ߦ߅޿ߡࠛࠬ࠻ࡠࠫࠚࡦߩࡐࠫ࠹ࠖࡉࡈࠖ࡯࠼ࡃ࠶ࠢ૞↪ࠍખ੺ߔࠆ␹⚻ᯏ
᭴߇㓽㓶ߣ߽ߦሽ࿷ߔࠆߎߣࠍ␜ໂߒߡ޿ࠆ‫⎇ᧄޕ‬ⓥߪ‫ࡦࡇࡠ࠻࠼࠽ࠧޔ‬᡼಴ࡎ࡞ࡕࡦ㧔GnRH㧕ಽᴲଦㅴ࿃ሶߢ
޽ࠆ␹⚻ࡍࡊ࠴࠼‫ߩߘ߮ࠃ߅⃻⊒ߩࡦ࠴ࡊࡍࠬࠠޔ‬ᵴᕈൻࠍ⚵❱ቇ⊛ߦ⸃ᨆߔࠆߎߣߦࠃߞߡ‫ޔ‬㔤㐳㘃ߦ߅޿ߡឃ
ෆࠍ⺃⿠ߔࠆࠧ࠽࠼࠻ࡠࡇࡦߩᄢ㊂᡼಴ࠍ೙ᓮߔࠆ⣖ౝࡔࠞ࠾࠭ࡓߩ⸃᣿ࠍ⋡⊛ߣߒߚ‫࡞ࠩࡦࡎ࠾ޕ‬㓽㓶⸘ 11 㗡
ࠍ↪޿‫❥ߩ࡞ࠩࡦࡎ࠾ޔ‬ᱺ㧔੤የ㧕ᦼߦ৻ㅪߩታ㛎ࠍⴕߞߚ‫ޕ‬వߕᕈ⣼㒰෰ࠍⴕ޿‫ޔ‬ᰴߦߘߩඨᢙߩ୘૕ߦࠛࠬ࠻
ࡠࠫࠚࡦࠍᛩਈߒߚ‫ߩߘޕ‬ᓟ‫ޔ‬ἠᵹ࿕ቯࠍⴕ޿‫ⷞޔ‬ᐥਅㇱࠍណขߒߚ‫ޕ‬Kiss1 ㆮવሶ⊒⃻⚦⢩ࠍ in situ hybridazation
ߦࠃࠅᬌ಴ߔࠆߣߣ߽ߦ‫ࡦࠚࠫࡠ࠻ࠬࠛޔ‬ಣℂߦࠃࠅ Kiss1 ⊒⃻⚦⢩ߦ߅ߌࠆᦨೋᦼㆮવሶ cFos ࠲ࡦࡄࠢߩ⊒⃻ࠍ
఺∉⚵❱ൻቇߦࠃࠅᬌ⸛ߒߚ‫⚿ߩߘޕ‬ᨐ‫⃻ޔ‬࿷߹ߢߦ‫ޔ‬㓽ߩ Kiss1 ⊒⃻⚦⢩ߦ߅޿ߡ‫ࡦࠚࠫࡠ࠻ࠬࠛޔ‬ଐሽᕈߩ cFos
࠲ࡦࡄࠢ⊒⃻ࠍ⷗޿ߛߒߚ‫⃻ޕ‬࿷ห᭽ߩ⸃ᨆࠍណขߒߚ㓶୘૕ߦߟ޿ߡᆎ߼ߡ߅ࠅ‫ޔ‬Kiss1 ⚦⢩ߢߩ cFos ࠲ࡦࡄࠢ
⊒⃻ࠍᬌ⸛ߒ‫ޔ‬㓽㓶ߩ⚿ᨐࠍ✚ว⊛ߦ♖ᩏߒߡ㔤㐳㘃ߦ߅ߌࠆឃෆߩ೙ᓮᯏ᭴ߩ⸃᣿ࠍ⋡ᜰߔ‫ޕ‬
(4) 㓐ᤨ൐㓸⎇ⓥ
D-1 ࠾ࡎࡦࠩ࡞ߩ଻ోቇผߦ㑐ߔࠆ⎇ⓥ (2)
๺↰৻㓶 ᚲౝኻᔕ⠪㧦ᷰㆻ㇌ᄦ
₎ኂߪ 1970 ᐕઍᓟඨ߆ࠄỗൻߒ‫ߣࠇߘޔ‬ਗⴕߒߡ࠾ࡎࡦࠩ࡞ߩ㚟㒰ᢙߪỗჇߒߚ‫ޕ‬2000 ᐕઍߦߪ㚟㒰ᢙ߇ᐕ㑆
1 ਁ㗡ࠍ⓭⎕ߒߚ‫ࠇߘޕ‬᡿‫ో଻࡞ࠨޔ‬ቇߩ␠ળ⊛ᓎഀ߇㊀ⷐᕈࠍჇߒߚ‫ޕ‬੩ᄢ㔤㐳⎇ߣᣣᧄࡕࡦࠠ࡯࠮ࡦ࠲࡯ߩ๟
ㄝ⁁ᴫࠍ⏕⹺ߔࠆ‫ޕ‬㔤㐳⎇ߩ౒ห೑↪⎇ⓥߩേะߪᣣᧄߩߘࠇࠍ෻ᤋߔࠆߣ⠨߃ࠆ‫ޕ‬㊁ᄖ⎇ⓥࠍᛮ߈಴ߔߣ‫ޔ‬1969
ᐕߩ 4 ઙ߆ࠄ 1974 ᐕߩᦨᄙ 24 ઙߦߥࠅ‫ߩߘޔ‬ᓟẋᷫߒߚ‫⺞ߩߘޕ‬ᩏ࿾ߪ‫ޔ‬ೋᦼߦߪో࿖ฦ࿾ߦಽᏓߒߡ޿ߚ߇‫ޔ‬
ᰴ╙ߦደਭፉߣ㊄⪇ጊߦ㓸ਛ‫❗ޔ‬ዊߒߡ߈ߚ‫ޕ‬
౒ห⎇ⓥߩਛߢ‫⃻ޔ‬ᴫ⎇ⓥળߪࠨ࡞ߩ଻ోࠍ⋡⊛ߦߒߚ㊀ⷐߥ㓸ળߢ޽ࠆ‫ ߪࠇߘޕ‬1972-77 ᐕ‫ޔ‬1990-91 ᐕ‫ޔ‬1993
ᐕ‫ޔ‬1995-97 ᐕ‫⺞ޕߚࠇࠊⴕߦޔ‬ᩏ࡮⎇ⓥ߇ᵴ⊒ߢ޽ߞߚ 1970 ᐕઍߦߪ 6 ᐕ㑆⛮⛯ߒߡⴕࠊࠇߚ߇‫ޔ‬1978 ᐕ߆ࠄ
1989 ᐕ߹ߢ㐿௅ߐࠇߕ‫ޔ‬ౣ㐿ߒߡ߽ᢿ⛯⊛ߢ‫ޔ‬1998 ᐕએ㒠㐿௅ߐࠇߥ߆ߞߚ‫ޕ‬
1956 ᐕߦ⸳┙ߐࠇߚᣣᧄࡕࡦࠠ࡯࠮ࡦ࠲࡯ߪ‫ޔ‬ታ㛎േ‛ଏ⛎ࠍ 1 ߟߩ⋡ᮡߦߒߚ߇‫ޔ‬㊁₎౏⧞ߩᓎഀࠍඳ‛㙚
ᵴേߦ⟎޿ߚ⧯ᚻ⎇ⓥ⠪ߩᗧ⷗ߦࠃߞߡ‫ޔ‬㊁₎ᗲ⼔ㅪ⋖ߪ⸃ᢔ‫ޔ‬ᯏ㑐⚕‫ޟ‬㊁₎‫ߪޠ‬ᑄೀߐࠇߚ‫ޔߚ߹ޕ‬᥉෸ᵴേߣ
ߒߡ㜞ߊ⹏ଔߐࠇߡ޿ߚ‫࡯ࠠࡦࡕޟ‬෹ߩળ‫⸃߇ޠ‬ᢔߒ‫ߩߘޔ‬ᯏ㑐⚕ߩ‫ߪޠ࡯ࠠࡦࡕޟ‬ᑄೀߦߥߞߚ‫ޕ‬㊁₎߽ࡕࡦࠠ
࡯߽଻ోߢߪ৻ᓎ⾈ߞߡ޿ߚߩߢ޽ࠆ‫ޕ‬
㔤㐳⎇ߩ࠾ࡎࡦࠩ࡞㊁ᄖⷰኤᣉ⸳ߪ 2008 ᐕߦ㐽㎮ߐࠇ‫ߩో଻࡞ࠨޔ‬᜚ὐ߇㔤㐳⎇߆ࠄᶖᄬߒߚ‫࠾ޔ߽ࠇߕ޿ޕ‬
ࡎࡦࠩ࡞଻ో⎇ⓥߩផㅴߦᰳߊߴ߆ࠄߑࠆⷐ⚛ߢ޽ߞߚ‫ޕ‬
D-2 㔤㐳㘃㓸࿅ߦ߅ߌࠆᗧᔒ᳿ቯߩ࡝ࠬࠢߣㆃᑧߩ㑐ଥ
ࡑ࡝ࠛ࡮ࡍ࡟㧔ࠬ࠻࡜ࠬࡉ࡞ࠣᄢቇ㧕 ᚲౝኻᔕ⠪㧦ᷰㆻ㇌ᄦ
㓸࿅↢ᵴࠍ༡߻⒳ߦߣߞߡ‫ߩߡోޔ‬୘૕ߩⴕേ߇⺞๺ߒ‫ⴕߡߞ߹ߣ߹ߟ߆ޔ‬േߔࠆߎߣ߇㊀ⷐߢ޽ࠆ‫ߩߘޕ‬႐ว‫ޔ‬
ᅤ૗ߦᣧߊᱜ⏕ߦ‫ޔ‬㓸࿅ߣߒߡߩᗧᔒ᳿ቯ߇ⴕࠊࠇࠆߩ߆ߣ޿߁ߎߣ߇໧㗴ߦߥࠆ߇‫ޔ‬ᗧᔒ᳿ቯߩᣧߐߣᱜ⏕ߐߩ
㑆ߦߪ‫࠼࡯࡟࠻ޔ‬㨯ࠝࡈߩ㑐ଥ߇޽ࠆ‫ߚ߹ޕ‬㓸࿅ߩᄢ߈ߐ߿᭴ᚑ߽‫ޔ‬㓸࿅ߣߒߡߩᗧᔒ᳿ቯߦᓇ㗀ߔࠆ‫ߚߒ߁ߎޕ‬
㓸࿅ߣߒߡߩᗧᔒ᳿ቯㆊ⒟ࠍ᣿ࠄ߆ߦߒ‫ߚ߹ޔ‬ᗧᔒ᳿ቯߩᣧߐߣ୘૕㑆ߢߩⴕേ਄ߢߩ㥿㦉߇߅ߎࠆ႐วߪ‫߁ߤޔ‬
޿߁ⷐ࿃߇㑐ଥߒߡ޿ࠆߩ߆ࠍ᣿ࠄ߆ߦߔࠆߎߣࠍ⋡ᮡߦ‫ ╙ޔ‬2 ࠠࡖࡦࡄࠬߩ᡼㘺⟲ࠍኻ⽎ߦⷰኤࠍⴕߞߚ‫߆ߒޕ‬
ߒ‫ޔ‬ታ㓙ߦ୘૕⼂೎ࠍߒ⟲ࠇ୘૕㑆ߩ␠ળ㑐ଥߦ߅ߌࠆၮᧄ⊛ᖱႎࠍ෼㓸ߒߡ޿ߚ߇‫ߩ⥸⻉ޔ‬੐ᖱ߆ࠄᏫ࿖ࠍ૛௾
ߥߊߐࠇ‫੹ޔ‬࿁ߩṛ࿷ᦼ㑆ਛߦߘࠇએ਄ߩⷰኤߦ〯ߺㄟ߻ߎߣߪߢ߈ߥ߆ߞߚ‫ⷰޔߒ߆ߒޕ‬ኤᣇᴺߩౕ૕⊛ߥ⚵ߺ
┙ߡ߿‫ޔ‬ታ㛎⊛ᚻᴺߩ⚵ߺวࠊߖ╬‫ޔ‬ౣ⸰ߒߚ႐วߦߪߔߋ⎇ⓥࠍ㐿ᆎߔࠆߎߣ߇ߢ߈ࠆߛߌߩ‫⷗ޔ‬ㅢߒࠍᓧࠆߎ
ߣ߇ߢ߈ߚ‫੹ޕ‬ᓟ‫ޔ‬ౣ᧪ᣣߒߡߎߩ⺖㗴ߢߩ⎇ⓥࠍౣ㐿ߔࠆߎߣࠍ⸘↹ߒߡ޿ࠆ‫ޕ‬
D-3 ࠨ࡞ߩ⴫ᖱવᨴߦ㑐ߔࠆ⎇ⓥ
㧙㧙
Ꮉวિᐘ㧔ฬᄢ࡮㒮࡮ᖱႎ⑼ቇ㧕 ᚲౝኻᔕ⠪㧦㚅↰໪⾆
ઁ⠪߇޽ࠆ⴫ᖱࠍߒߚߣ߈ߦ‫ⷰࠍࠇߘޔ‬ኤߒߡ޿ࠆࡅ࠻ߪߟ޿หߓࠃ߁ߥ⴫ᖱࠍߔࠆߎߣ㧔⴫ᖱવᨴ㧕߇⍮ࠄࠇ
ߡ޿ࠆ‫⴫ߥ߁ࠃߩߎޕ‬ᖱવᨴߪ‫ⷰߣࠅ߈ߞߪߦ⊛ⷡⷞޔ‬ኤߢ߈ࠆ߫޽޿߽޽ࠆ߇‫╭ߪߢ࠻ࡅޔ‬㔚ࠍ᷹ቯߒߡ‫ޔ‬㗼࿷
ൻߒߥ޿⴫ᖱࡄ࠲࡯ࡦߣߒߡ␜ߐࠇࠆߎߣ߇ᄙ޿‫ޕ‬
࠴ࡦࡄࡦࠫ࡯ߩ‫߇ޠ߮ߊ޽ޟ‬વᨴߔࠆߣߩႎ๔߇޽ࠆ߇‫⴫ߩߢ࡞ࠨޔ‬ᖱવᨴߩႎ๔ߪ߹ߛߥ޿‫ߣ࠻ࡅޕ‬หߓࠃ߁
ߦ‫ⴕޔ‬േߣߒߡⷰኤߐࠇߥߊߡ߽╭㔚ߩ࡟ࡌ࡞ߢ⴫ᖱ߇વᨴߒߡ޿ࠆน⢻ᕈ߇޽ࠆ‫ࠍߣߎߩߘޕ‬ᬌ⸛ߔࠆߚ߼ߦ‫ޔ‬
ࠨ࡞ࠍࡕࡦࠠ࡯࠴ࠚࠕߦ࿕ቯߒ‫⴫ޔ‬ᖱવᨴ߇↢ߓࠆ߆ࠍᬌ⸛ߒߚ‫ޕ‬
H23 ᐕᐲߪ੍஻⊛ߥ⎇ⓥߣ૏⟎ߠߌ‫ߦࠕࠚ࠴࡯ࠠࡦࡕࠍ࡞ࠨߕ߹ޔ‬ᐳࠄߖࠆ⸠✵ߣ‫╭ޔ‬㔚↪㔚ᭂߩ㚔⥌ࠍⴕߞ
ߚ‫ߣߣߎߩߘޕ‬ᐔⴕߒߡ‫ޔ‬ೝỗߣߒߡឭ␜ߔࠆߚ߼ߩࠨ࡞ߩ⴫ᖱࠍ᠟ᓇߒߚ‫ޔߒ߆ߒޕ‬㘺⢒ࠤ࡯ࠫౝߢߪశ㊂߇⿷
ࠄߕ㗻߇ᥧߊߥࠆߎߣ߿‫ޔ‬ᱜ㕙ࠍะ޿ߚ⴫ᖱ⴫಴ߩ႐㕙ࠍ߁߹ߊ᠟ᓇߢ߈ߥ޿ߎߣ߇޽߈ࠄ߆ߦߥߞߚ‫ޔߢߎߘޕ‬
޽ࠆ⴫ᖱߣਛᕈ㗻ߩࡕ࡯ࡈࠖࡦࠣߦࠃࠅ⴫ᖱ⴫಴ߩേ↹ߩ૞ᚑࠍߒߡ޿ࠆ‫⥄ࠅࠃޔߦࠄߐޕ‬ὼߥ⴫ᖱ߇⴫಴ߐࠇࠆ
㊁ᄖߩࠨ࡞ߩ⴫ᖱࠍ᠟ᓇߒ‫ޔ‬ೝỗ↪ߦടᎿࠍⴕߞߚ‫ޕ‬
ᰴᐕᐲએ㒠‫ߩࠄࠇߎޔ‬ೝỗࠍ↪޿ߡ‫ߢࠕࠚ࠴࡯ࠠࡦࡕޔ‬቟ቯߒߡࡕ࠾࠲ࠍ⷗ࠄࠇࠆ୘૕߆ࠄ‫ઁޔ‬୘૕ߩ⴫ᖱⷰኤ
ᤨߩ╭㔚ࠍ᷹ቯߔࠆ੍ቯߢ޽ࠆ‫ޕ‬
D-5 ࡑࡂ࡟ߩ࠴ࡦࡄࡦࠫ࡯ߩ⁚⁸⡺㘩ⴕേߦ㑐ߔࠆᄙⷺ⊛ಽᨆ
଻ဈ๺ᒾ㧔㎨ୖᅚሶᄢ࡮ఽ┬㧕 ᚲౝኻᔕ⠪㧦MA Huffman
੹ᐕᐲߪ‫ޔ‬᡿⷏↰೑⽵ඳ჻‫ޔ‬ਛ᧛⟤⍮ᄦඳ჻㧔੩ᄢ㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯㧕߆ࠄឭଏߐࠇߚࡑࡂ࡟ጊ႙㧔࠲ࡦࠩ
࠾ࠕ㧕ߩ࠴ࡦࡄࡦࠫ࡯ߩ⁚⁸⡺㘩ⴕേ⾗ᢱࠍᢛℂߒ‫ޔ‬Hosaka et al. (2001) એ㒠‫ᦝޔ‬ᣂߐࠇߡ޿ߥ޿ 1996 ᐕᐲ⺞ᩏએ
㒠ߩ࠺࡯࠲ߩ౒᦭૞ᬺࠍㅴ߼‫ޔ‬ᚑᨐ౏⴫ߦะߌߡᛂߜวࠊߖߚ‫ޕ‬ቢᚑ߹ߢߦ‫ⶄߦࠄߐޔ‬ᢙߩ౒ห⎇ⓥ⠪ߦ࠺࡯࠲ឭ
ଏߩଐ㗬ࠍⴕ߁ᔅⷐ߇޽ࠆ‫↰⷏ޕ‬ඳ჻߇ㅤ෰ߒߚߚ߼‫ޔ‬ឭଏߐࠇߚ࠺࡯࠲ߦߟ޿ߡᧄੱߦ໧޿วࠊߖࠆߎߣ߇ਇน
⢻ߣߥࠅ‫ޔ‬ඳ჻߇ㆮߒߚࡈࠖ࡯࡞࠼ࡁ࡯࠻ߩ৻ㇱ⺒ߺ⋥ߒࠍⴕ߁ᔅⷐ߇↢ߓߚ‫ߩࠄࠇߎޕ‬૞ᬺߪᰴᐕᐲ࠹࡯ࡑߦ߅
޿ߡ߽⛮⛯ߔࠆᔅⷐ߇޽ࠆ‫ࠈߎߣߩ੹ޕ‬᣿ࠄ߆ߦߥࠅߟߟ޽ࠆ੐ታߪએਅߩㅢࠅߢ޽ࠆ‫ޕ‬Ԙࠕࠞࠦࡠࡉࠬ߇₪‛ߣ
ߥࠆ㗫ᐲ෸߮ഀวߪ 1990 ᐕઍᓟඨ߆ࠄᄢ߈ߊᄌࠊߞߡ޿ߥ޿‫ޕ‬ԙㄭᐕ‫ߡ޿ߟߦ࡯ࠫࡦࡄࡦ࠴ߩ࡟ࡃࠠ߿ࡌࡦࠧޔ‬
⁚⁸ⴕേߩ⎇ⓥࠍ⊒⴫ߔࠆ⎇ⓥ⠪ߪ‫ߩ࡯ࠫࡦࡄࡦ࠴ޟޔ‬ᄢੱ㓶߇⡺ಽ㈩ࠍ᡽ᴦ⊛ߦ೑↪ߔࠆ‫߁޿ߣޠ‬઒⺑ߦ⇼ᔨࠍ
๒ߒߡ޿ࠆ‫ޔߒ߆ߒޕ‬1991 ᐕએ㒠‫ޔ‬8 㗡ߩࠕ࡞ࡈࠔ㓶߇಴⃻ߒߚࡑࡂ࡟ߩ⾗ᢱߪ‫ߩߎࠈߒ߻ޔ‬઒⺑ࠍᡰᜬߔࠆ௑ะ
ࠍ␜ߒߡ޿ࠆ‫⾗ޕ‬ᢱᢛℂ࡮ಽᨆࠍᕆ߉‫ߦ⴫౏ޔ‬ᜬߜㄟߺߚ޿‫⋧ߩ⸃⷗ޕ‬㆑߇࿾ၞ⊛ߥᏅ⇣ߢ޽ࠆน⢻ᕈߦߟ޿ߡ߽
వⴕ⎇ⓥࠍี๧ߒߡࠃߊ⸛⺰ߒߡ߅߈ߚ޿‫ޕ‬
D-6 The genetic basis of blue eyes in primates
M Przeworski, W Meyer, J Pickrell (University of Chicago) ᚲౝኻᔕ⠪㧦੹੗໪㓶
We have performed sequencing of the region homologous to the conserved region of HERC2 containing the well-supported
causal site for blue/brown polymorphism in humans in the 20 macaques from the PRI. We have obtained sequence data for a
subset of the conserved region of HERC2 in eight of the nine macaques from the Choshikei Monkey Park for which we
obtained DNA, and we are currently working to obtain sequence data for the remainder of the region and the ninth macaque.
So far, we find no single nucleotide polymorphism (SNP) or combination of SNPs within the region that segregate(s)
completely with eye color. We are also working with the macaque photographs in order to develop the best way to assess the
quantitative variation in eye color while accounting for technical variation among photos.
D-7 㘃ੱ₎ߩ♮ዩࠍ↪޿ߚᣂߚߥ↢ℂᜰᮡ ߩ⹏ଔᴺߩ㐿⊒
ᷡ᳓ᘮሶ㧔ጟጊℂ⑼ᄢ࡮ℂቇㇱ࡮േ‛ቇ⑼㧕 ᚲౝኻᔕ⠪㧦ᯅᧄජ⛗
♮ዩਛࡎ࡞ࡕࡦ᷹ቯᴺࠍ㘃ੱ₎߅ࠃ߮ࡑࠞࠢዻࠨ࡞ߩ㓶࡮㓽ߩ❥ᱺ⁁ᘒߩផቯ߅ࠃ߮ᕈᚑᾫᐲ߿⠧ൻߩ⒟ᐲߩ
ផቯߦᔕ↪ߔࠆߚ߼‫ޔ‬ᕈ⣼߅ࠃ߮೽⣢⊹⾰↱᧪ߩࠬ࠹ࡠࠗ࠼ࡎ࡞ࡕࡦߦߟ޿ߡ‫ޔ‬ੑ᛫૕㉂⚛఺∉᷹ቯᴺߦࠃࠆ᷹ቯ
♽ߩ㐿⊒߅ࠃ߮ߘߩᬌ⸛ࠍⴕߞߚ‫ޕ‬ട߃ߡ‫ޔ‬ታ㓙ߦ㘃ੱ₎߅ࠃ߮ࡑࠞࠢߩ♮ዩࠍ↪޿‫ߩࠄࠇߎޔ‬᷹ቯߦ߅޿ߡ⥋ㆡ
ߩ଻ሽ᧦ઙ߿᛽಴᧦ઙࠍ᳿ቯߔࠆߚ߼᭽‫ߥޘ‬ᣇᴺࠍ⹜ߺߚ‫⚿ߩࠄࠇߎޕ‬ᨐ‫⺧↳ߦߢ߹ࠇߎޔ‬⠪ࠄ߇⏕┙ߒߚᕈ⣼↱
᧪ߩࡊࡠࠥࠬ࠴ࡦ‫ߩࡦࡇࡠ࠻࠼࠽ࠧޔࡦࠥࡠ࠼ࠕޔࡦࠥࡠ࠻ࠬࠛޔ‬ੑ᛫૕ᴺ㉂⚛఺∉᷹ቯᴺߦട߃‫ޔ‬೽⣢↱᧪ߩࠕ
ࡦ࠼ࡠࠬ࠹ࡦࠫࠝࡦ‫ߩߡ޿ߟߦࡦࠝࠫࡦ࠹ࠬࡠ࠼ࡦࠕࡇࠛࡠ࠼ࡅ࠺ޔ‬᷹ቯᴺࠍ⏕┙ߔࠆߎߣ߇ߢ߈ߚ‫ޔߜࠊߥߔޕ‬
♮ዩਛࡊࡠࠥࠬ࠴ࡦ‫ࡦࡇࡠ࠻࠼࠽ࠧ߮ࠃ߅ࡦࠝࠫࡦ࠹ࠬࡠ࠼ࡦࠕޔࡦࠥࡠ࠼ࡦࠕޔࡦࠥࡠ࠻ࠬࠛޔ‬Ớᐲ᷹ቯߦࠃࠆ
ᕈ೎߅ࠃ߮ᕈᚑᾫᐲផቯ‫ޔ‬ෆ⢩⊒⢒߿㤛૕ᒻᚑ‫ޔ‬ឃෆ߿ᅧᆼߩ⏕⹺߇น⢻ߣߥߞߚ‫ޕ‬
ߐࠄߦᧄᐕᐲߪࠬ࠻࡟ࠬ㑐ㅪࡎ࡞ࡕࡦߦߟ޿ߡ߽♮ዩࠍ↪޿ߚ᷹ቯᴺࠍ⏕┙ߒߚ‫ߩࠄࠇߎޔߚ߹ޕ‬ᣇᴺࠍ↪޿
ߡ‫ޔ‬㘺⢒ਅࡑࠞࠢዻࠨ࡞ߩ♮߅ࠃ߮ዩ‫ޔ‬ኻᔕ⠪߇ណขߒߚ㊁↢࠴ࡦࡄࡦࠫ࡯ߩ♮߅ࠃ߮ዩਛߩࡎ࡞ࡕࡦ㊂ࠍ᷹ቯߔ
ࠆߎߣ߇ߢ߈ߚ‫ޔߚ߹ޕ‬㊁ᄖߦ߅ߌࠆ♮ዩߩណขᴺ‫଻ޔ‬ሽᣇᴺߩᡷ⦟‫ޔ‬᛽಴᧦ઙߩᬌ⸛ࠍ߅ߎߥ޿‫ޔ‬಄⬿‫ޔ‬಄ಓ⸳
஻ߩ⏕଻ߢ߈ߥ޿㊁ᄖߦ߅޿ߡᓧࠄࠇߚࠨࡦࡊ࡞߆ࠄ߽ࡎ࡞ࡕࡦઍ⻢↥‛᷹ቯน⢻ߥᣇᴺࠍ⠨᩺ߒߚ‫⃻ޕ‬࿷ߐࠄߦ
♖ᐲะ਄ߩߚ߼ߦᬌ⸛ࠍⴕߞߡ޿ࠆ‫ޕ‬
D-8 ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆⴊ▤ᯏ⢻ߦ㑐ߔࠆ⎇ⓥ
↰๺ᱜᔒ㧘ጟ᧛ንᄦ㧔ṑ⾐කᄢ࡮⮎ℂቇ㧕 ᚲౝኻᔕ⠪㧦ᄢ⍹㜞↢
ૐ㉄⚛޽ࠆ޿ߪౣ㉄⚛ਅߢߪ‫㉄৻ޔ‬ൻ⓸⚛㧔NO㧕ଏਈ૕ߦࠃࠆ౰⁁േ⣂᜛ᒛ૞↪߇ᷫᒙߒߡ޿ࠆ߇‫ߩࠄࠇߎޔ‬
⁁ᴫ߇ NO ⥄りߦᓇ㗀ࠍ෸߷ߒߡ޿ࠆߩ߆‫ߦ〝⚻࡞࠽ࠣࠪߩߘ߽ߣࠇߘޔ‬ᓇ㗀ࠍ෸߷ߒߡ޿ࠆߩ߆ߦߟ޿ߡߪਇ᣿
㧙㧙
ߢ޽ࠆ‫ޕ‬
੹࿁ߩ⎇ⓥߢߪ‫ޔ‬ㅢᏱ㉄⚛ਅ‫ޔ‬ૐ㉄⚛ਅ‫ޔ‬ౣ㉄⚛ਅߦ߅ߌࠆ sGC stimulator ߦࠃࠆ╬ዤᕈߩᒛജᄌൻࠍࡑࠣ࠿ࠬ
ᴺߦḰߓߡ⸥㍳ߒߚ‫ޔ߅ߥޕ‬sGC stimulator ߣߪ NO ߩ૞↪ὐߢ޽ࠆนṁᕈࠣࠕ࠾࡞㉄ࠪࠢ࡜࡯࠯㧔sGC㧕ࠍ NO
㕖ଐሽ⊛ߦᵴᕈൻߔࠆ⮎‛ߢ޽ࠆ‫⚿ߩߘޕ‬ᨐ‫ޔ‬ૐ㉄⚛޽ࠆ޿ߪౣ㉄⚛᥸㔺ߦࠃࠅ‫ޔ‬sGC stimulator ߦࠃࠆ౰⁁േ⣂
᜛ᒛ૞↪ߪ᦭ᗧߢߪߥ޿߽ߩߩᷫᒙߔࠆ௑ะࠍ␜ߒߚ‫⎇ᧄޕ‬ⓥ⚿ᨐߪ‫ޔ‬ૐ㉄⚛޽ࠆ޿ߪౣ㉄⚛ߦ᥸㔺ߐࠇߚ౰⁁േ
⣂ߢߪ‫ޔ‬sGC ߦ߽૗ࠄ߆ߩᄌൻ߇↢ߓߡ޿ࠆน⢻ᕈࠍ␜ໂߒߡ޿ࠆ‫ޕ‬
੹ᐕᐲߪ౒ห೑↪࡮౒ห⎇ⓥߩೋᐕᐲߢ޽ࠅ‫ޔ‬ᚑᨐࠍ⊒⴫ߔࠆ߹ߢߦߪ⥋ߞߡ޿ߥ޿‫ᧄޕ‬ᚑᨐߪ‫᧪ޔ‬ᐕᐲએ㒠ߦ
⊒⴫ߒߡ޿ߊ੍ቯߢ޽ࠆ‫ޕ‬
D-9 ູ੃㘃ߦ߅޿ߡߩ๰ྨ࡝࠭ࡓߣ૕㊀ߦ߅ߌࠆ⋧ኻᚑ㐳ߩ㑐ଥ
㐳ፃᐽඳ㧔ᣣᧄᱤ⑼ᄢᣂẟ࡮㒮࡮⍶ᱜ㧕㧘ኹ↰ຬੱ㧔ᣣᧄᱤ⑼ᄢᣂẟ࡮⍶ᱜ㧕㧘૒⮮⟵⧷㧔ᣣᧄᱤ⑼ᄢᣂẟ࡮↢ℂ㧕
ᚲౝኻᔕ⠪㧦ᐔፒ㍈⍫
๰ྨ࡝࠭ࡓߪ‫ޔ‬⣖ᐙߦሽ࿷ߔࠆਛᨔᕈࡄ࠲࡯ࡦ⊒↢ེߦࠃࠅᒻᚑߐࠇࠆߣ⠨߃ࠄࠇߡ޿ࠆ‫ߩߘޔࠄ߇ߥߒ߆ߒޕ‬
࡝࠭ࡓߩⷙቯ࿃ሶߪ᣿ࠄ߆ߦߥߞߡ޿ߥ޿‫ޕ‬ㆊ෰ߩ⎇ⓥ߆ࠄ‫ޔ‬๰ྨ࡝࠭ࡓߪ૕㊀ߣ㑐ଥ߇޽ࠆߎߣ߇ႎ๔ߐࠇߡ߅
ࠅ‫ޔ‬ਅ㗶ㆇേߪ㗶㑐▵ࠍਛᔃߣߒߚ࠹ࠦߩㆇേߢ޽ࠆ‫⎇ᧄߢߎߘޕ‬ⓥߪ‫ޔ‬๰ྨ࡝࠭ࡓߣ૕㊀ߩ㑐ଥࠍౣᬌ⸛ߒ‫ޔ‬๰
ྨ࡝࠭ࡓߣਅ㗶㛽ᒻᘒߩ㑐ଥࠍᬌ⚝ߒߚ‫ޕ‬
࠺ࠫ࠲࡞ࠞࡔ࡜ߢ๰ྨᤨߩ㔤㐳㘃ࠍ᠟ᓇߒߚ‫ޕ‬㙄ࠍ៨㘩ߒߡ߆ࠄྚਅߔࠆ߹ߢߩᦨ߽቟ቯߒߚ 4 ࿁ߩ๰ྨࠍ᛽಴
ߒ‫ޔ‬ᐔဋࠍ᳞߼ฦ୘૕ߩ๰ྨ࡝࠭ࡓߣߒߚ‫ޕ‬ኻ⽎േ‛ߩ૕㊀ߪᢥ₂ࠍᒁ↪ߒߚ‫ߚ߹ޕ‬ਅ㗶㛽ᮡᧄࠍࠞࡔ࡜ߢ᠟ᓇߒ‫ޔ‬
↹௝਄ߢਅ㗶㛽㐳ᓘߣਅ㗶㛽ߩ㊀ᔃߩ૏⟎ࠍ᳞߼ߚ‫ߡߒߘޕ‬ਅ㗶㗡߆ࠄਅ㗶㛽㊀ᔃߩ߹ߢߩ〒㔌ߣਅ㗶㗡߆ࠄਅ㗶
╙৻ᄢ⥓ᱤ߹ߢߩ〒㔌ࠍ⸘᷹ߒ‫ޔ‬⒳㑆ߢᲧセᬌ⸛ߒߚ‫ޕ‬
๰ྨ࡝࠭ࡓߣ૕㊀‫ޔ‬ਅ㗶㛽㐳ᓘ‫ޔ‬ਅ㗶㗡߆ࠄਅ㗶㛽㊀ᔃ߹ߢߩ〒㔌߅ࠃ߮ਅ㗶㗡߆ࠄਅ㗶╙৻ᄢ⥓ᱤ߹ߢߩ〒㔌
ߣߩ㑆ߦߪ‫ޔ‬ᱜߩ⋧㑐߇⹺߼ࠄࠇߚ‫ߩࠦ࠹ޔࠄ߇ߥߒ߆ߒޕ‬ේℂߦ㑐ଥߔࠆਅ㗶㛽ᒻᘒ㧔ਅ㗶㗡߆ࠄਅ㗶㛽㊀ᔃ߹
ߢߩ〒㔌߅ࠃ߮ਅ㗶㗡߆ࠄਅ㗶╙৻ᄢ⥓ᱤ߹ߢߩ〒㔌㧕ߪ‫ޔ‬⒳㑆ߦ߅ߌࠆ㆑޿ߪ⹺߼ࠄࠇߥ߆ߞߚ‫ޕ‬
ߎࠇࠄߩߎߣ߆ࠄ‫ޔ‬๰ྨࡄ࠲࡯ࡦ⊒↢ེߦࠃࠅᒻᚑߐࠇࠆ๰ྨ࡝࠭ࡓߪ‫ޔ‬૕㊀ߦࠃߞߡⷙቯߐࠇߡ޿ࠆน⢻ᕈ߇
޽ࠅ‫ޔ‬ਅ㗶㛽ᒻᘒߣߩ㑐ㅪߪዊߐ޿੐߇␜ໂߐࠇߚ‫ޕ‬
D-10 㘃ੱ₎ߩ␹⚻વ㆐㑐ㅪㆮવሶߩᄙ᭽ᕈ⸃ᨆ
᧛ጊ⟤Ⓞ㧔੩ㇺᄢ࡮㊁↢േ‛㧕 ᚲౝኻᔕ⠪㧦ㇹᐽᐢ
ᧄ⎇ⓥߢߪ‫ߢ࠻ࡅޔ‬ႎ๔ߐࠇߡ޿ࠆᕈᩰߦ㑐ਈߔࠆㆮવሶߩ⋧ห㗔ၞࠍ㘃ੱ₎ߢ⸃ᨆߒ‫ޔ‬⒳㑆ߩႮၮ㈩೉Ყセ
߿‫ޔ‬୘૕ߩᕈᩰ⹏ቯߣߩ㑐ㅪ⸃ᨆࠍⴕߞߡ‫ޔ‬㘺⢒߿❥ᱺߦᵴ↪ߔࠆᖱႎࠍᓧࠆߎߣࠍ⋡ᜰߒߡ޿ࠆ‫ޕ‬㑐ㅪᕈߩ⸃ᨆ
ߦߪᄙᢙߩ⹜ᢱ߇ᔅⷐߥߚ߼‫ޔ‬GAIN ࠍㅢߓߡ㘺⢒㘃ੱ₎ߩ⹜ᢱឭଏࠍଐ㗬ߒ‫ޔ‬Ყセน⢻ߥ࠺࡯࠲ߩ⫾Ⓧࠍ⋡ᜰߒ
ߡ޿ࠆ‫ޕ‬23 ᐕᐲߪ࠴ࡦࡄࡦࠫ࡯3‫ ࡜࡝ࠧ࠼ࡦ࡜࡯ࡠࠪ࠾ޔ‬2‫ ࡦ࠲࡯࠙ࡦ࡜ࠝޔ‬1‫ ࡞ࠩࠟ࠽࠹ࡠࠢࡈޔ‬1 ߩ⸘ 7 ୘૕
ߩ⹜ᢱឭଏࠍฃߌ‫ޔ‬DNA ࠍ᛽಴ߒߚ‫ߩ࠻ࡅޔߡ޿ߟߦ࡜࡝ࠧߜ߁ߩߘޕ‬ᕈᩰ‫ߦ․ޔ‬ਇ቟߿᡹᠄ᕈߣߩ㑐ਈ߇ႎ๔
ߐࠇߡ޿ࠆ␹⚻વ㆐߅ࠃ߮ࡎ࡞ࡕࡦવ㆐㑐ㅪߩ 5 ㆮવሶ㧔ࡃ࠰ࡊ࡟ࠪࡦฃኈ૕‫ ࠯࡯࠳ࠪࠠࠝࡦࡒࠕࡁࡕޔ‬A‫ࡁࡕޔ‬
ࠕࡒࡦࠝࠠࠪ࠳࡯࠯ B‫ࡦࠥࡠ࠼ࡦࠕޔ࡯࠲࡯ࡐࠬࡦ࡜࠻ࡦ࠾࠻ࡠ࠮ޔ‬ฃኈ૕㧕8 㗔ၞࠍᲧセߒߚ‫⚿ߩߘޕ‬ᨐ‫ࠪ࠾ޔ‬
ࡠ࡯࡜ࡦ࠼ࠧ࡝࡜ߩ࿖ౝ㘺⢒ 14 ୘૕ߢߪ‫ޔ‬8 㗔ၞߔߴߡߢᄙဳ߇⷗ࠄࠇߚ㧔ᐔဋኻ┙ㆮવሶᢙ 3.8‫ޔ‬ᐔဋ߳࠹ࡠធ
ว₸ 0.517㧕‫ޕ‬㊁↢ߩ࠾ࠪࡠ࡯࡜ࡦ࠼ࠧ࡝࡜߅ࠃ߮ࡑ࠙ࡦ࠹ࡦࠧ࡝࡜ߣᲧセߒߚߣߎࠈ‫ޔ‬ኻ┙ㆮવሶ㗫ᐲߩᏅ⇣߇
⷗޿ߛߐࠇߚ‫ޕ‬㘺⢒୘૕㑆ߢᏅ߇ᄢ߈޿ߎߣ߆ࠄ‫ޔ‬ᕈᩰߣߩ㑐ㅪ⸃ᨆߩࡑ࡯ࠞ࡯ߣߒߡ᦭ലߢ޽ࠆߣ⠨߃ࠄࠇࠆ‫ޕ‬
߹ߚ⒳㑆Ꮕߦߟ޿ߡߪ‫ⴕޔ‬േ߿␠ળߩ⒳Ꮕߦ߽ߣߠ޿ߡ⸃㉼ߢ߈ࠆ߆߽ߒࠇߥ޿‫੹ޕ‬ᓟߪ୘૕ᢙ‫ޔ‬୥⵬ㆮવሶᢙࠍ
Ⴧ߿ߒߡ‫ޔ‬ᕈᩰߩࡑ࡯ࠞ࡯ߣߥࠆㆮવሶࠍត⚝ߔࠆ੍ቯߢ޽ࠆ‫ޕ‬
D-11 㔤㐳㘃ߦ߅ߌࠆ⚵❱․⇣⊛ DNA ࡔ࠴࡞ൻᯏ᭴ߩ⸃᣿
⑔↰ᷧ㧘৻ᩉஜม㧘૒‫⵨ᧁޘ‬ਯ㧔਻Ꮊᄢ࡮↢ක⎇㧕 ᚲౝኻᔕ⠪㧦ㇹᐽᐢ
DNA ࡔ࠴࡞ൻߪઍ⴫⊛ߥࠛࡇࠫࠚࡀ࠹ࠖ࠶ࠢߥᯏ᭴ߩ৻ߟߢ‫ဳ⃻⴫ޔ‬ᄙ᭽ᕈߦ㑐ਈߔࠆߎߣ߇⍮ࠄࠇߡ޿ࠆ‫ޕ‬
ߘߩߚ߼‫ߩ࡯ࠫࡦࡄࡦ࠴ߣ࠻ࡅޔ‬ㅴൻߦ߅ߌࠆࠛࡇࠫࠚࡀ࠹ࠖ࠶ࠢߥᯏ᭴ߩᓎഀࠍ⸃᣿ߔࠆߦߪ⦟޿ኻ⽎ߢ޽ࠆ‫ޕ‬
ᚒ‫࡯ࠫࡦࡄࡦ࠴ߣ࠻ࡅߦߢ߹ࠇߎߪޘ‬㧔㔤㐳㘃⎇ⓥᚲߩ㘺⢒୘૕㧕ߩᧃ᪳ⴊ⊕ⴊ⃿ߦ߅ߌࠆ DNA ࡔ࠴࡞ൻᲧセ⎇
ⓥࠍⴕ޿‫ޔ‬ᨴ⦡૕ 21‫ޔ‬22 ⇟਄ߦ 16 ࠞᚲߩࡔ࠴࡞ൻᏅ⇣㗔ၞ ࠍหቯߒߡ޿ߚ‫ޕ‬
ࡔ࠴࡞ൻᏅ⇣߇⊒↢ߩߤߩᤨᦼߦᒻᚑߐࠇࠆ߆ࠍ⺞ߴࠆߎߣߢ‫࡞࠴ࡔޔ‬ൻᏅ⇣ߩᒻᚑၮ⋚ߩ৻┵߇᣿ࠄ߆ߦߥࠆ
ߣᦼᓙߒ‫⎇ᧄޔ‬ⓥߢߪᧃ᪳ⴊ⊕ⴊ⃿ߦ߅ߌࠆࡔ࠴࡞ൻᏅ⇣㗔ၞߩࡔ࠴࡞ൻࠍઁ⚵❱㧔⣛⤳‫ޔ⤳⤟ޔ‬ᄢ⣖‫♖ޔ‬Ꮍ‫♖ޔ‬
ሶ㧕ߢ߽⺞ߴߚ‫ߪ࡞ࡊࡦࠨ❱⚵ߩ࡯ࠫࡦࡄࡦ࠴ޕ‬ᄢဳ㘃ੱ㘃ᖱႎࡀ࠶࠻ࡢ࡯ࠢ㧔GAIN㧕߅ࠃ߮ࠨࡦࠢ࠴ࡘࠕ࡝ቝ
࿯߆ࠄಽਈߒߡ㗂޿ߚ߽ߩࠍ೑↪ߒߚ‫࡞࠴ࡔޕ‬ൻ⸃ᨆߩ⚿ᨐ‫࡞࠴ࡔߩߤࠎߣ߶ޔ‬ൻᏅ⇣ߪᧃ᪳ⴊ⊕ⴊ⃿․⇣⊛ߢ޽
ࠆߎߣ߇ࠊ߆ߞߚ‫࡞࠴ࡔޔߪߣߎߩߎޕ‬ൻᏅ⇣ߪਥߦ⊒↢ᓟᦼߦᒻᚑߐࠇࠆߎߣࠍ␜ໂߒߡ޿ࠆ‫ޔࠄ߇ߥߒ߆ߒޕ‬
ࡔ࠴࡞ൻࠍ⺞ߴߚ㗔ၞߪ㕖Ᏹߦ㒢ࠄࠇߡ޿ࠆߩߢ‫੹ޔ‬࿁ߩ⚿ᨐ߇ࠥࡁࡓࡢࠗ࠼ߢ߽ᒰߡߪ߹ࠆߩ߆ࠍ᣿ࠄ߆ߦߔࠆ
ߚ߼ߦߪ‫⸃ߥ⊛⟜✂ࠅࠃޔ‬ᨆ߇ᔅⷐߢ޽ࠆ‫ޕ‬
⃻࿷‫ޔ‬㜞ㅦࠪ࡯ࠢࠛࡦࠨ࡯ࠍ↪޿ߡࡅ࠻ߣ࠴ࡦࡄࡦࠫ࡯ߩᧃ᪳ⴊ⚦⢩ߩࡔ࠴࡞ൻࠍࠥࡁࡓࡢࠗ࠼ߦᲧセߒߡ޿ࠆ
ߩߢ‫੹ޔ‬ᓟߪઁ⚵❱ߩࡔ࠴࡞ൻ߽ห᭽ߦ⸃ᨆߔࠆ੍ቯߢ޽ࠆ‫ޕ‬
㧨ቇળ⊒⴫㧪
㧙㧙
⑔↰ᷧ㧘߶߆‫ޟ‬DNA methylation differences between humans and chimpanzees in chromosomes 21 and 22‫ޠ‬. ╙ 7 ࿁ ਻
Ꮊᄢቇ Global-COE Young Investigators Forum
D-12 ⢽⾰ࠍᮡ⊛ߣߒߚࠨ࡞఺∉ࠪࠬ࠹ࡓߩ⸃᣿
᧖↰᣽ᒾ㧘᫪↰ᄢテ㧔੩ㇺᄢ࡮࠙ࠗ࡞ࠬ⎇㧕 ᚲౝኻᔕ⠪㧦㋈ᧁ᮸ℂ
⚦⩶߿࠙ࠗ࡞ࠬߩᗵᨴߦ߅޿ߡ‫∛ޔ‬ේ૕߇↥↢ߔࠆ⢽⾰ಽሶ޽ࠆ޿ߪ⢽⾰ࠍ฽᦭ߒߚⶄวಽሶࠍᮡ⊛ߣߒߚᗵᨴ
㒐ᓮᔕ╵߇⺃⿠ߐࠇࠆߎߣ߇᣿ࠄ߆ߦߥࠅߟߟ޽ࠆ‫⎇ޕ‬ⓥઍ⴫⠪ࠄߪࡅ࠻∛ේ૕㧔⚿ᩭ⩶߿ࠛࠗ࠭࠙ࠗ࡞ࠬߥߤ㧕
߇ኋਥ↢૕ౝߢ↥↢ߔࠆ lipidic ߥ఺∉ᮡ⊛ಽሶߦኻߔࠆ T ࡝ࡦࡄ⃿ᔕ╵ߩ⸃ᨆࠍⴕߞߡ߈ߚ‫఺ߥ⊛⥸৻ޔߒ߆ߒޕ‬
∉⸃ᨆߦ᦭↪ߥዊേ‛ߢ޽ࠆࡑ࠙ࠬ߿࡜࠶࠻ߪߎߩ఺∉ࠪࠬ࠹ࡓࠍᰳᅤߒߡ޿ࠆߚ߼‫ߥ⚦⹦ߩߘޔ‬ಽሶ࡮⚦⢩ᯏᐨ
ߩ⸃᣿ߦߪࡅ࠻ߦ㘃ૃߒߚ఺∉ࠪࠬ࠹ࡓࠍ᦭ߔࠆേ‛߇ਇนᰳߢ޽ࠆ‫⎇ᧄߢߎߘޕ‬ⓥߢߪ‫ᧃ࡞ࠩࠥࠞࠕޔ‬᪳ⴊනᩭ
⃿ࠍ↪޿‫∉఺ߩߎޔ‬ᔕ╵ߦ㑐ਈߔࠆಽሶ࡮⚦⢩ᯏᐨࠍ᣿ࠄ߆ߦߔࠆߣߣ߽ߦ‫ߩߘޔ‬೙ᓮᴺࠍ⏕┙ߔࠆߎߣࠍ⋡⊛ߣ
ߔࠆ‫ޕ‬
߹ߕ‫ᧃ࡞ࠨޔ‬᪳ⴊࠃࠅ♖⵾ߒߚන⃿ࠍࡑ࠙ࠬ࡮࡜࠶࠻߳ߣ఺∉ߒߚᓟ‫ޔ‬B ⚦⢩ࠍขࠅ಴ߒ‫⚦ࡑ࡯ࡠࠛࡒޔ‬⢩ߣߩ
Ⲣวࠍⴕߞߚ‫ ⚂ߦߢ߹ࠇߎޕ‬4000 ࠢࡠ࡯ࡦߩࡂࠗࡉ࡝࠼࡯ࡑࠍන㔌ߒ‫৻ࠆࠃߦ࡯࡝࠻ࡔ࠻ࠗࠨ࡯ࡠࡈޔ‬ᰴࠬࠢ࡝
࡯࠾ࡦࠣ߆ࠄ 270 ࠢࡠ࡯ࡦߩࠨ࡞න⃿․⇣⊛ߥࡕࡁࠢࡠ࡯࠽࡞᛫૕ࠍㆬᛮߒߚ‫ޔߡ޿⛯ޕ‬T ࡝ࡦࡄ⃿ᔕ╵ߩ㒖ኂ᛫
૕ࠍតߔੑᰴࠬࠢ࡝࡯࠾ࡦࠣ߆ࠄ 7 ࠢࡠ࡯ࡦߩࡕࡁࠢࡠ࡯࠽࡞᛫૕ࠍ⷗಴ߒߚ‫↢ޕ‬ൻቇ⸃ᨆߦࠃࠆ⹺⼂᛫ේߩหቯ
ࠍㅴ߼ߚ⚿ᨐ‫∉఺ߩߎޔ‬ᔕ╵ߦ㑐ࠊࠆᧂ⍮ߩ఺∉ಽሶ߿ធ⌕ಽሶߩ୥⵬ࠍ⛉ࠅㄟࠎߛ‫ޕ‬
D-13 㔤㐳㘃ࠍ฽߻ູ੃㘃ߩ྾⢇㛽ᒻ⁁᭴ㅧߩ᧚ᢱജቇ⊛ᕈ⾰ߣᆫ൓ㆇേߣߩ㑐ଥ
๺↰⋥Ꮖ㧘᧼ᧄ๺ੳ㧘ᓟ⮮ᘏ㧔ጊญᄢ㧕㧘⮮↰ᔒᱠ㧔㣮ఽፉᄢ㧕 ᚲౝኻᔕ⠪㧦⷏᧛೰
྾⢇㛽‫ߦ․ޔ‬⢋↲㛽ߩᒻ⁁ߣࡠࠦࡕ࡯࡚ࠪࡦߩ㑐ଥࠍ᣿ࠄ߆ߦߒ‫੃ູޔ‬㘃ߩ↢ᕷၞߩᄙ᭽ᕈߣࡠࠦࡕ࡯࡚ࠪࡦߩ
㑐ଥ߆ࠄ‫↢ޔ‬ᕷⅣႺߩ↢૕ߦ߅ࠃ߮ᓇ㗀ࠍ᣿ࠄ߆ߦߔࠆߩ߇ᧄ⎇ⓥ⺖㗴ߩ⋡⊛ߢ޽ࠆ‫⎇ޕ‬ⓥߪ╭࡮㛽ᩰ♽ߩ⸃೬ቇ
⊛⎇ⓥ‫ߩࡦ࡚ࠪ࡯ࡕࠦࡠޔ‬᠟ᓇ࠺࡯࠲ߩㆇേቇ⊛⎇ⓥࠍਥߣߒߡⴕࠊࠇࠆ‫ޕ‬2011 ᐕᐲ߹ߢߦ㔤㐳㘃ࠍ฽߼ߡ⚂ 70
⒳ߩູ੃㘃ߩ⢋↲㛽ࠍ฽߼ߚ㛽ᩰߩ CT ᠟ᓇ‫ߚ߹ޕߚߞⴕࠍޔ‬⢋↲㛽ߩᄖᒻ⸘᷹‫ޔ‬๟ㄝߩ╭ߦߟ޿ߡߩ⺞ᩏࠍⴕߞ
ߚ‫᦭ޕ‬㒢ⷐ⚛ᴺߦࠃࠆᔕജߩ▚಴૞ᬺ߇Ꮏቇ♽ߩ⎇ⓥ⠪ߩදജࠍᓧߡᆎ߹ߞߚ‫ߩࡦ࡚ࠪ࡯ࡕࠦࡠޕ‬᠟ᓇߪਥߦേ‛
࿦ߢⴕߞߡ޿ࠆ‫ޕ‬േ‛ߩㆇേߩዷ␜ࠍ⋡ᜰߒߡ⸳ߌࠄࠇߚࠨࡈࠔ࡝࡜ࡦ࠼ߢߪ㜞ㅦ⿛ⴕߩ᠟ᓇ߽น⢻ߢ޽ࠆ‫ࡎ࠾ޕ‬
ࡦࠩ࡞‫ߪߡ޿ߟߦࠞࠪޔ‬ደਭፉⷰኤ࠮ࡦ࠲࡯ߩ೑↪ࠍ⸵นߒߡ߽ࠄ޿⥄ὼⅣႺਅߩࡠࠦࡕ࡯࡚ࠪࡦࠍ᠟ᓇߒߚ‫ູޕ‬
੃㘃ࠍℂ⸃ߔࠆߎߣࠍ⋡⊛ߒߡ޿ࠆߚ߼‫ߦࠄߐޔ‬ᄙߊߩ⸃೬‫ޔ‬ㆇേቇ⊛૞ᬺ߇ᔅⷐߢ޽ࠆ߇‫⌕ޔ‬ታߦ࠺࡯࠲ߪ⫾Ⓧ
ߐࠇߡ޿ࠆ‫ޕ‬
D-14 㔤㐳㘃✂⤑߅ࠃ߮⣖ߦ߅ߌࠆࠝࡊࠪࡦ⊒⃻ㇱ૏ߩ⸃ᨆ
৾↰⧐ೣ㧔੩ㇺᄢ࡮㒮ℂ㧕㧘ᄢౝᶻઍ㧔ᓼፉᄢ࡮㒮࠰ࠪࠝ࠹ࠢࡁࠨࠗࠛࡦࠬ㧕㧘ጊਅ㜞ᑝ㧔੩ㇺᄢ࡮㒮ℂ㧕 ᚲౝ
ኻᔕ⠪㧦ਛ᧛స᮸
ࡅ࠻ࠍ฽߻㔤㐳㘃ߩࠥࡁࡓߦߪ‫⚦ⷞߩ⤑✂ޔ‬⢩ߦ⊒⃻ߒⷞⷡߩಽሶၮ⋚ߣߥࠆశฃኈ࠲ࡦࡄࠢ⾰㧔ࠝࡊࠪࡦ㧕ㆮ
વሶએᄖߦ߽‫ࡦࠪࡊࠝߩ߆ߟߊ޿ޔ‬ㆮવሶ߇⏕⹺ߐࠇߡ޿ࠆ‫ߥ߁ࠃߩߤ߇ࠄࠇߘޔߒ߆ߒޕ‬ಽሶ⊛ᕈ⾰ࠍ᦭ߒ‫ߤޔ‬
ߎߦ⊒⃻ߒ‫ⷡⷞޔ‬એᄖߩߤߩࠃ߁ߥ↢ℂᯏ⢻ߦ㑐ࠊࠆ߆‫ᧂߪߡ޿ߟޔ‬⍮ߩㇱಽ߇ᄙ޿‫ޕ‬
ᚒ‫⎇ᧄߪޘ‬ⓥ⺖㗴ࠍ㐿ᆎߔࠆ೨ߦ‫ޔ‬㕖ⷞⷡᯏ⢻ࠍᜂ߁ࠝࡊࠪࡦ Opn5 ߦߟ޿ߡ࠾ࡢ࠻࡝ߢ⸃ᨆࠍⴕ޿‫⚡ޔ‬ᄖశᗵ
ฃᕈߢ✂⤑ߩࠕࡑࠢ࡝ࡦ⚦⢩࡮␹⚻▵⚦⢩ߦ⊒⃻ߔࠆߎߣࠍ⷗޿ߛߒߡ޿ߚ‫ᧄߢߎߘޕ‬㓐ᤨ൐㓸⎇ⓥ⺖㗴ߦ߅޿
ߡ‫ޔ‬㔤㐳㘃ߦ߅ߌࠆߎߩࠝࡊࠪࡦߩ↢ℂᯏ⢻ߦㄼࠆߚ߼‫ࠆߌ߅ߦ⤑✂ޔ‬ዪ࿷ࠍ᣿ࠄ߆ߦߔࠆߎߣࠍ⋡⊛ߣߒߚ‫ࡑޕ‬
࡯ࡕ࠮࠶࠻߅ࠃ߮ࠕࠞࠥࠩ࡞ߩ✂⤑ߦኻߒߡ․⇣⊛᛫૕ࠍ↪޿ߚ఺∉ᨴ⦡ታ㛎ࠍⴕߞߚߣߎࠈ‫⚦ⷞޔ‬⢩એᄖߩ৻ㇱ
ߩ␹⚻⚦⢩ߦࠪࠣ࠽࡞ࠍ⷗޿ߛߔߎߣߦᚑഞߒߚ‫ޕ‬
D-15 㗡ㇱ்ኂᜰᮡឭ᩺ߦะߌߚࠬࠤ࡯࡝ࡦࠣᚻᴺߩ㐿⊒
J Antona㧘ዊ㊁ฎᔒ㇢㧘ᳯፉ᤯㧔৻⥸⽷࿅ᴺੱ ᣣᧄ⥄േゞ⎇ⓥᚲ㧕 ᚲౝኻᔕ⠪㧦⷏᧛೰
A new method has been applied to develop a finite element model of the head-neck complex of a Macaque from medical
images. The skull and the brain have been validated based on tissue and component experimental data from literature. The
kinematics of the head under occipital impacts has been validated against a sub-set of head impact experiments carried out in
the past at the Japan Automobile Research Institute. The validated model has been used to simulate 19 occipital impacts
case-by-case. The correlation between mechanical parameters of the different brain organs at the simulated impacts and the
occurrence of concussion in the experiments was analyzed. Stress in the brainstem showed significant correlation to
concussion as recorded in the experimental data from the past. The developed model and the presented results constitute the
first step towards the development of a tissue level injury criterion for human that is based on experimental animal data.
D-16 ੇቄߦ߅ߌࠆ࠴ࡖࠗࡠࠠ࠷ࡀࠩ࡞ߩណ㘩ࡄ࠲࡯ࡦ㧦ᨐታ㘩ߣ⪲㘩ߩ⇣ߥࠆᯏ⢻
૒⮮ብ᮸㧔੩ㇺᄢ࡮ࠕࡈ࡝ࠞ⎇㧕 ᚲౝኻᔕ⠪㧦ඨ⼱๋㇢
ࡑ࠳ࠟࠬࠞ࡞↥㔤㐳㘃ߩ߁ߜ‫ ⑼࡞ࠩࡀ࠷ࠠޔ‬Eulemur ዻߩ㘩ᕈߪ‫ޔ‬ᄙߊߩⷰኤ⎇ⓥ߅ࠃ߮ᶖൻ▤᭴ㅧ߆ࠄᨐታ㘩
ߢ޽ࠆߎߣ߇ᜰ៰ߐࠇߡ޿ࠆ‫࡞ࠞࠬࠟ࠳ࡑޔߒ߆ߒޕ‬ർ⷏ㇱߩᾲᏪੇ῎ᨋߦ↢ᕷߔࠆ࠴ࡖࠗࡠࠠ࠷ࡀࠩ࡞ߪ‫ੇޔ‬ቄ
ᓟඨߦᣣਛߩᨐታ㘩ഀวࠍᄢ߈ߊᷫࠄߒ‫ޔ‬Lissochilus rutenbergii㧔࡜ࡦ⑼㧕ߩ⨲ᧄࠍིߺ✦߼ߡ⚵❱ᶧࠍ⥤߼ขࠆ
ⴕേߦ㐳ᤨ㑆ࠍ⾌߿ߔߎߣ߇ߎࠇ߹ߢߩⷰኤ߆ࠄ᣿ࠄ߆ߦߥߞߡ޿ࠆ‫৻ޕ‬ᣇ‫ޔ‬ᄛ㑆ߪߎߩ⪲ࠍోߊ೑↪ߖߕ‫ޔ‬ᨐታ
ࠍਛᔃߦណ㘩ߔࠆ‫⚿ߩᦼᤨߩߎޕ‬ታᧁኒᐲߪઁߩቄ▵ߣᄌࠊࠄߥ޿‫ޕ‬ᣣਛߩ⪲㘩ߣᄛ㑆ߩᨐታ㘩ߩᯏ⢻ࠍតࠆߚ
㧙㧙
߼‫ޔ‬2011 ᐕੇቄᓟඨߦ㘩‛⾗Ḯࠍណขߒ‫ޔ‬ᩕ㙃ࠍಽᨆߒߚ‫ޕ‬L. rutenbergii ߪੇ῎㊀㊂ߢ࠲ࡦࡄࠢ⾰߇ 9.2%‫ޔ‬นṁ
ή⓸⚛‛㧔NFE㧕߇ 20.8%ߢ޽ࠆߩߦኻߒ‫ੇޔ‬ቄ⚿ታᨐታ 3 ⒳ߩᨐ⡺ߪ࠲ࡦࡄࠢ⾰߇ 2.7㧙14.2%‫ޔ‬NFE ߇ 52.7㧙69.2%
ߣߥߞߚ‫ޕ‬᳓ಽ฽᦭₸ߪ‫ޔ‬L. rutenbergii ߇Ḩ㊀㊂ߢ 80.1%‫ޔ‬ᨐ⡺߇ 6.3㧙27.4%ߩ᳓ಽࠍ฽ࠎߢ޿ߚ‫ੇޕ‬ቄᓟඨߪᣣ
ਛߩ᳇᷷߇ 1 ᐕߢᦨ߽㜞ߊ‫ߪ࡞ࠩࡀ࠷ࠠࡠࠗࡖ࠴ޕࠆ޽ߢᦼᤨߚߒ῎ੇޔ‬ᥤᾲ᧦ઙਅߢ᳓ಽࠍ₪ᓧߔࠆߚ߼ߦᣣਛ
ߪ L. rutenbergii ߩ⪲ߩ೑↪ࠍఝవߒ‫ޔ‬ᄛ㑆ߪࠛࡀ࡞ࠡ࡯៨ขߩߚ߼ߦᨐታࠍណ㘩ߔࠆߣ⠨߃ࠄࠇࠆ‫⪲ߩߎޕ‬㘩ߣᨐ
ታ㘩ߩ⇣ߥࠆᯏ⢻ߣ‫ᤨޔ‬㑆ߦࠃࠆ㘩ߴಽߌ߇‫ੇޔ‬ቄߦߺࠄࠇࠆ๟ᣣⴕᕈߩㆡᔕᗧ⟵ࠍ⺑᣿ߔࠆ઒⺑ߣߥࠆ‫ޕ‬
㧨⪺ᦠ㧪
૒⮮ብ᮸㧚ᤤ߽ᄛ߽േߊࠠ࠷ࡀࠩ࡞ߩ⻘㧚㧔ਛᎹዏผ㧘෹᳗㓷Ꮖ㧘ጊᭂᄈ৻ ౒✬㧕‫ޡ‬ᣣᧄߩࠨ࡞ቇ㧙⧯ᚻ⎇ⓥ
⠪ߩᦨ೨✢‫ޢ‬੩ㇺㅢା␠㧔಴ ੍ቯ㧦2012/09㧕
D-17 ⠧㦂⣖ߦ߅ߌࠆ࠲࠙Ⱞ⊕⾰ߩ⊒⃻ಽሶ⒳߅ࠃ߮࡝ࡦ㉄ൻߦ㑐ߔࠆᲧセ∛ℂቇ⊛⎇ⓥ
ਛጊ⵨ਯ㧘ౝ↰๺ᐘ㧘࠴ࠚࡦࡃ࡯࠭ ࠫࠚ࡯ࡓ࠭㧔᧲੩ᄢ࡮㒮࡮ㄘቇ↢๮⑼ቇ㧕 ᚲౝኻᔕ⠪㧦㋈ᧁ᮸ℂ
੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲߦ଻▤ߐࠇߡ޿ࠆࠨ࡞㘃ߩ⣖ߩࡎ࡞ࡑ࡝ࡦ࿕ቯᮡᧄ‫ࡦࠖࡈ࡜ࡄޔ‬൮ၒࡉࡠ࠶ࠢ߆ࠄ⚵❱ಾ
ࠍ૞⵾ߒ‫ޔ‬ฦ⒳ᨴ⦡‫∉఺ޔ‬ᨴ⦡ߥߤࠍⴕߞߡ‫∛ޔ‬ᄌࠍ⸃ᨆߔࠆ‫↢ޕ‬೨ߩⴕേ⹏ଔ࠺࡯࠲߇޽ࠇ߫‫ߣࠄࠇߎޔ‬⣖∛
ᄌߩ⒳㘃‫ޔ‬⒟ᐲߣࠍᲧセߔࠆ‫ޕ‬
ᧄ⎇ⓥ⺖㗴߇ណᛯߐࠇߚߩ߇ 2011 ᐕ 8 ᦬ߛߞߚߚ߼‫ޔ‬ᒰᐕᐲߪ㔤㐳㘃⎇ⓥᚲߦ⵬ቢߐࠇߡ޿ࠆ਄⸥ᮡᧄߩᢛℂ
ߣᔅⷐߥᮡᧄߩ᛽಴ߩߺࠍⴕߞߚ‫ޕ‬
ᧄ⎇ⓥߪ 2012 ᐕᐲ߽⛮⛯ߒߡណᛯߐࠇߚߩߢ‫ޔ‬ᣧㅦᮡᧄࠍ૞⵾ߒ਄⸥ߩᬌ⚝ࠍⴕ߁‫ޕ‬
D-18 ᜰߩⵚ்ߩ⊒↢ෂ㒾ᕈߦ㑐ߔࠆ⹏ଔᴺ⏕┙ߩߚ߼ߩ↢૕ജቇ⊛⎇ⓥ
ဈᧄੑ㇢㧔㊄ᴛᄢ࡮ᯏ᪾Ꮏቇ♽㧕㧘ችፒ␭੺㧔᧲੩Ꮏᬺᄢ࡮ᖱႎⅣႺቇ㧕㧘ᄙ↰లᓼ㧔↥ᬺᛛⴚ✚ว⎇ⓥᚲ࡮࠺ࠫ
࠲࡞ࡅࡘ࡯ࡑࡦᎿቇ⎇ⓥ࠮ࡦ࠲࡯㧕 ᚲౝኻᔕ⠪㧦⷏᧛೰
ᣣᏱ↢ᵴߦ߅ߌࠆ੐᡿ߣߒߡᜰ᜽ߺߪᄙ޿‫ߦ․ޕ‬ሶߤ߽ߩኅᐸౝ੐᡿ߦ߅ߌࠆฃ்ㇱ૏ߩ╙ 2 ૏ߪᚻߢ޽ࠅ‫ޔ‬ਛ
ߢ߽ᜰߪߐߺ੐᡿ߦ߅ߌࠆⵚ்⊒↢ࡔࠞ࠾࠭ࡓࠍ⸃᣿ߔࠆߎߣ߇ߘߩ੍㒐ߩߚ߼ߦߪᔅⷐߢ޽ࠆ‫ߩ⤏⊹ޔߒ߆ߒޕ‬
ⵚ்⊒↢ࡔࠞ࠾࠭ࡓߪ⸃᣿ߐࠇߡ߅ࠄߕ‫⹏ߩߘޔ‬ଔᣇᴺ߽ሽ࿷ߒߥ޿‫ޕ‬
ߘߎߢ‫⎇ᧄޔ‬ⓥߪ‫੃ޔ‬ᐜఽߩᚻᜰߩࠨࠗ࠭ߣ߶߷╬ߒ޿࠾ࡎࡦࠩ࡞߅ࠃ߮ࠕࠞࠥࠩ࡞ߩዱ૕ᚻᜰߩ₂૕ࠍ↪޿
ߡ‫ߦ⤏⊹ޔ‬㑐ߔࠆታ㛎ߣߘࠇࠍౣ⃻ߒߚࠪࡒࡘ࡟࡯࡚ࠪࡦࠍታᣉߔࠆ‫ߩᤨ்ⵚ⤏⊹ޔࠅࠃߦࠇߎޕ‬ജቇ⊛᧦ઙࠍ᣿
ࠄ߆ߦߔࠆߎߣߢ‫⹏ߩ்ⵚߩ⤏⊹ޔ‬ଔᣇᴺࠍ⏕┙ߔࠆߎߣࠍ⋡⊛ߣߔࠆ‫ޕ‬
ᧄᐕᐲߪ‫ߩ࠻ࡅޔ‬ᚻߣኸᴺ߇ㄭ޿࠾ࡎࡦࠩ࡞߅ࠃ߮ࠕࠞࠥࠩ࡞ߩዱᜰߦኻߒߡ‫᧚ޔ‬ᢱ․ᕈขᓧࠍ⋡⊛ߣߒߡၮ␆
⊛ߥታ㛎ࠍታᣉߒߚ‫⿥ߩ❱⚵⤏⊹ޔߕ߹ޕ‬ᒢᕈ․ᕈߣ⎕ᢿ․ᕈࠍขᓧߔࠆߚ߼ߦዱᜰߩ᛼ߒㄟߺታ㛎ࠍⴕ޿‫⊹⴫ޔ‬
⎕ᢿ߹ߢߩ᛼ߒㄟߺ⩄㊀ߣ᛼ߒㄟߺ㊂ߩ㑐ଥࠍขᓧߒߚ‫࡞ࠨޔߦࠄߐޕ‬ዱᜰ߆ࠄಾࠅ಴ߒߚಾ ߦኻߔࠆᒁᒛ⹜㛎
ࠍታᣉߒ‫ߩ⊹⴫ޔ‬ၮ␆⊛ߥജቇ․ᕈߩขᓧ߽ⴕߞߚ‫ޕ‬
ߎࠇࠄߩታ㛎ߩ⚿ᨐ‫ޔ‬㔤㐳㘃ߩᚻᜰߩജቇ․ᕈߦ㑐ߔࠆၮ␆⊛ߥ࠺࡯࠲෼㓸ࠍⴕ߁ߎߣ߇ߢ߈ߚ‫੹ޕ‬ᓟߪߎࠇࠄ
࠺࡯࠲ࠍᵴ↪ߒ‫ੱޔ‬૕ᜰ߳ߩജቇ․ᕈߩࠬࠤ࡯࡝ࡦࠣᣇᴺߩ㐿⊒ߣࠪࡒࡘ࡟࡯࡚ࠪࡦߦࠃࠆⵚ்⊒↢ࡔࠞ࠾࠭ࡓߩ
⸃᣿ࠍታᣉߒ‫ࠅࠃޔ‬቟ోߥ⵾ຠ࡮ⅣႺߩታ⃻ߦነਈߒߚ޿‫ޕ‬
D-19 6-OHDA ᵈ౉ߦ߅ߌࠆ DA ␹⚻ᡰ㈩ߩ㒖ኂലᨐߩᬌ⸛
⦁ᯅᣂᄥ㇢㧔੩ㇺᄢ࡮ߎߎࠈߩᧂ᧪⎇ⓥ࠮ࡦ࠲࡯㧕㧘ᷡ᳓ᘮሶ㧔ጟጊℂ⑼ᄢ࡮ℂ㧕㧘ฎ↰⾆ኡ㧔੩ㇺᄢ࡮ක㧕 ᚲ
ౝኻᔕ⠪㧦ᱜ㜞ା↵
೨ᐕᐲ߹ߢߩ⎇ⓥߢ‫ޔ‬ᐜᐕࡑࠞࠢࠩ࡞ߩ೨㗡ㅪว㊁ߦᛩ኿ߔࠆ࠼ࡄࡒࡦ㧔DA㧕♽✢⛽ࠍ 6-OHDA ߦࠃࠅ⎕უߒ‫ޔ‬
ߘߩᓟߩⴕേⷰኤߦࠃࠅ ADHD ఽߦ⷗ࠄࠇࠆⴕേ․ᓽߣห᭽ߩ․ᓽ߇↢ߓࠆߎߣࠍⴕേቇ⊛ߦᬌ⸛ߔࠆߣหᤨ
ߦ‫ޔ‬⎕უߦࠃࠆ㓚ኂߩ⥃⇇ᦼߩ᦭ήࠍᬌ⸛ߒߡ߈ߚ‫੹ޕ‬ᐕᐲߪ‫⎇ޔ‬ⓥߦ↪޿ߡ߈ߚേ‛ࠍߔߴߡታ㛎Ვߒ‫ޔ‬೨㗡⪲
ߦ߅ߌࠆ 6-OHDA ߦࠃࠆ⎕უߩലᨐߩᬌ⸽ߣ‫ⴕޔ‬േታ㛎⚿ᨐߣߩ㑐ଥࠍ߽ߣߦ‫ޔ‬േ‛ࡕ࠺࡞ߣߒߡߩ᦭ലᕈࠍᬌ
⸽ߒߚ‫ޕ‬
ਔ஥ߩ೨㗡ㅪว㊁⢛ᄖ஥ㇱߦ 6-OHDA ᵈ౉ߒߚࠨ࡞‫ޔ߮ࠃ߅ޔ‬㕖ᵈ౉ߩኻᾖࠨ࡞ࠍታ㛎Ვߒ‫ޔ‬೨㗡ㅪว㊁ߦ߅
ߌࠆ 6-OHDA ߦࠃࠆ DA ✢⛽ߩ⎕უലᨐࠍ‫ޔ‬Tyrosine hydroxylase ߩ఺∉⚵❱ቇ⊛ᨴ⦡ᴺߦࠃࠅᬌ⸛ߒߚ‫⚿ߩߘޕ‬ᨐ‫ޔ‬
6-OHDA ᵈ౉ㇱ૏ߢߪ‫ޔ‬DA ✢⛽߇߶ߣࠎߤⷰኤߐࠇߕ‫ޔ‬᣿⏕ߥ⎕უലᨐ߇޽ߞߚߣหᤨߦ‫ޔ‬⎕უലᨐ߇ᢙᐕߦࠊ
ߚࠆ㐳ᦼ㑆ᜬ⛯ߒߡ޿ࠆߎߣ߽⏕⹺ߐࠇߚ‫⃻ޕ‬࿷‫ޔ‬ᓧࠄࠇߚ 6-OHDA ߦࠃࠆ⎕უലᨐߣ‫ⴕޔ‬േቇ⊛ᬌ⸛ߢᓧࠄࠇ
ߚ⚿ᨐࠍ⚵ߺวࠊߖ‫ޔ‬േ‛ࡕ࠺࡞ߣߒߡߩ᦭ലᕈ࡮ᅷᒰᕈࠍ⏕⹺ߒߡ޿ࠆ‫ޕ‬
D-20 ࠾ࡎࡦࠩ࡞ߩฦ⒳⚛᧚ߦኻߔࠆ⊓ࠅⴕേߩ⸃᣿
ᳯญ␭テ㧘ጊ↰ᓀ㧘਄↰ᒄೣ㧔ㄭਛ྾ㄘ⎇㧕㧘ၴጊቬ৻㇢㧔㤗Ꮣᄢ࡮₞ක㧕㧘ฎ⼱⋉ᦶ㧔ၯ₹ㄘᨋ✚⎇࠮ࡦ࠲࡯㧕
ᚲౝኻᔕ⠪㧦ඨ⼱๋㇢
㊁↢㠽₞ߦࠃࠆㄘ૞‛ⵍኂࠍ㒐ߋߚ߼ߦߪ‫ޔ‬ኻ⽎ߣߥࠆേ‛ߩ↢ᘒ߅ࠃ߮ⴕേࠍᛠីߒ‫ޔ‬ᣂߚߥ㒐㒰ᛛⴚߩ㐿⊒
߿✚ว⊛ኻ╷ߩዷ㐿ࠍ࿑ࠆᔅⷐ߇޽ࠆ‫ⵍߩ࡞ࠩࡦࡎ࠾ޔߒ߆ߒޕ‬ኂኻ╷ߦ⋥ធ⚿߮ߟ޿ߚㆇേ⢻ജߦ㑐ߔࠆⴕേቇ
⊛⎇ⓥߪዋߥߊ‫ޔ‬ၮ␆⊛ߥ⍮⷗ߩ⫾Ⓧ߇ᔅⷐߣߥࠆ‫ޕ‬ㄘ૞‛ⵍኂߩ㒐ᱛࠍ㔍ߒߊߒߡ޿ࠆේ࿃ߪേ‛ߩㆇേ⢻ജߣ
ቇ⠌⢻ജߩ㜞ߐߦ޽ࠅ‫⎇ߩߡ޿ߟߦࠄࠇߎޔ‬ⓥࠍㅴ߼ߡ޿߆ߥߌࠇ߫ߥࠄߥ޿‫⎇ᧄޔߢߎߘޕ‬ⓥߪ‫ߩ࡞ࠩࡦࡎ࠾ޔ‬
ㆇേ⢻ജ⎇ⓥߩ৻Ⅳߣߒߡ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆ⊓ࠅⴕേߦߟ޿ߡᄥߐߩ⇣ߥࠆᩇ߿ⷺᐲߩ⇣ߥࠆ௑ᢳࠍ↪޿ߡⴕ
㧙㧙
േߩⷰኤ࡮᷹ቯࠍⴕ޿‫ޔ‬㒐㒰ᩋ╬ߩ㐿⊒ᡷ⦟ߩၮ␆⊛⍮⷗ࠍᓧࠆߎߣࠍ⋡⊛ߣߒߚ‫ޕ‬
⋥ᓘߩ⇣ߥࠆᡰᩇߩ਄ㇱߦႎ㈽㘺ᢱࠍขࠅઃߌߡ⊓ࠅⴕേࠍ⺞ᩏߒߚ⚿ᨐ‫⋥ߪ࡞ࠨޔ‬ᓘ߇ 25cm એ਄ߩ᫔㧔Ⴎൻ
ࡆ࠾࡞⵾࡮⎬Ბࡏ࡯࡞⵾㧕ߢߪ⊓ࠅߩᚑഞ₸߇ᕆỗߦᷫዋߒߚ‫៺ޕ‬ᡂଥᢙߩૐ޿⚛᧚ࠍᡰᩇߦᏎ߈ઃߌߚ႐ว‫⚛ޔ‬
᧚ࠍᏎ߈ઃߌࠆ૏⟎ߦࠃߞߡ⊓ࠅߩᚑഞ₸ߦᄢ߈ߥᏅ߇⹺߼ࠄࠇߚ‫ޕ‬
߹ߚ‫ޔ‬㘺⢒ᣉ⸳ߩოߦ᧼ࠍᢳ߼ߦ┙ߡ߆ߌࠆࠃ߁ߦ࿕ቯߒߚ㕙ߩ਄ㇱߦႎ㈽㘺ᢱࠍขࠅઃߌߡⴕേࠍⷰኤߒߚ⺞
ᩏߢߪ‫⊓ߩ࡞ࠩࡦࡎ࠾ޔ‬ဈน⢻ⷺᐲߪ᧼㕙㧔㕒ᱛ៺ᡂଥᢙ 0.35㧕ߢ 40 ᐲߢ޽ࠅ‫ޔ‬45 ᐲ߆ࠄߪṖࠆ‫ޔ‬〡べߔࠆ‫✼ޔ‬
ߦߟ߆߹ߞߡ⊓ࠆⴕേ߇⹺߼ࠄࠇߚ‫ޕ‬
D-21 ໜᶧࠕࡒ࡜࡯࠯ㆮવሶᄙဳߣ㘩‛៨ข⁁ᴫߦ㑐ߔࠆ⎇ⓥ
㐳᎑ᵏ↢㧔ฬነᏒ┙ᄢ࡮ᩕ㙃㧕㧘㋈ᧁ⦟㓶㧘ਛ᧛ᕶሶ㧔㗅ᄤၴᄢ࡮ࠬࡐ࡯࠷⑼ቇ⑼㧕㧘ᳰ↰໪৻㧔㗅ᄤၴᄢ࡮ஜᐽ
⑼ቇ⑼㧕 ᚲౝኻᔕ⠪㧦੹੗໪㓶
ࡅ࠻ߩໜᶧࠕࡒ࡜࡯࠯ㆮવሶ AMY1 ߦߪࠦࡇ࡯ᢙᄙဳ߇ሽ࿷ߒ‫⃻⊒ࠢࡄࡦ࠲࠯࡯࡜ࡒࠕޔ‬㊂ߦᓇ㗀ࠍ෸߷ߔߎ
ߣ߇ႎ๔ߐࠇߡ޿ࠆ‫ࡦࡊࡦ࠺ޔߚ߹ޕ‬៨ข㊂ߩᄙ޿᳃ᣖߢߪ AMY1 ࠦࡇ࡯ᢙ߇⋧ኻ⊛ߦᄙ޿ߚ߼ positive selection
߇௛޿ߡ޿ࠆߣ⠨߃ࠄࠇࠆ߇‫ޔ‬ห৻㓸࿅ߦ߅ߌࠆ㘩੐ߩ࠺ࡦࡊࡦ៨ข㊂ߣ AMY1 ࠦࡇ࡯ᢙߣߩ㑐ㅪᕈߦߟ޿ߡߪ
ߎࠇ߹ߢ᣿ࠄ߆ߦߐࠇߡ޿ߥ޿‫ޕ‬೨ᐕᐲߩ⎇ⓥࠃࠅ‫ޔ‬AMY1 ࠦࡇ࡯ᢙ೎ߩ⟲㑆Ყセߢ㘩੐೨ᓟߩໜᶧࠕࡒ࡜࡯࠯ᄌ
േߦ᦭ᗧᏅ߇⷗ࠄࠇ‫࠯࡯࡜ࡒࠕޔ‬ᵴᕈߩ୘ੱᏅ߇␜ߐࠇߚߎߣ߆ࠄ‫ޔ‬AMY1 ࠦࡇ࡯ᢙ߇୘ੱߩ࠺ࡦࡊࡦᕈ㘩ຠߩ༵
ᅢߦ߽ᓇ㗀ߔࠆน⢻ᕈ߇޽ࠆ‫⎇ᧄߢߎߘޕ‬ⓥߪ㘩‛៨ข⁁ᴫߣ AMY1 ࠦࡇ࡯ᢙߣߩ㑐ㅪᕈߦߟ޿ߡᬌ⸛ߔࠆߎߣ
ࠍ⋡⊛ߣߒߚ‫⎇ޕ‬ⓥኻ⽎⠪ߪஜᏱߥᄢቇ↢ 30 ฬߢ‫ޔ‬Ԙၮᧄዻᕈߣ㘩੐ᤨ㑆ߦ㑐ߔࠆ⾰໧⚕⺞ᩏ‫ޔ‬ԙ⁁ᘒ-․ᕈਇ቟
ᬌᩏ STAI‫ޔ‬Ԛᤤ㘩೨ᓟߩໜᶧࠕࡒ࡜࡯࠯ᵴᕈߩ᷹ቯ‫ޔ‬ԛBDHQ ߦࠃࠆㆊ෰ 1 ߆᦬㑆ߩ㘩‛៨ข㗫ᐲ⺞ᩏࠍ 7 ᦬‫ޔ‬
11 ᦬ߦታᣉߒߚ‫ޕ‬AMY1 ㆮવሶࠦࡇ࡯ᢙߪ࡝ࠕ࡞࠲ࠗࡓ PCR ߩ SYBR green ᴺ߆ࠄ▚ቯߒ‫ޔ‬ㆮવሶࠦࡇ࡯ᢙߩᮡ
Ḱ⹜ᢱߣߒߡ AMY1 ࠦࡇ࡯ᢙᄙဳ߇ሽ࿷ߒߥ޿࠴ࡦࡄࡦࠫ࡯ࠥࡁࡓࠍ↪޿ߚ‫⚿ޕ‬ᨐߦߟ޿ߡߪ⃻࿷ಽᨆࠍㅴ߼ߡ
߅ࠅ‫ޔ‬ቄ▵ᄌേߥߤ߽⠨ᘦߒߥ߇ࠄ㘩੐߳ߩᓇ㗀ⷐ࿃ࠍᬌ⸛ߔࠆ੍ቯߢ޽ࠆ‫ޔߚ߹ޕ‬ໜᶧࠕࡒ࡜࡯࠯ᵴᕈߩㆇേࠬ
࠻࡟ࠬ߳ߩᔕ╵ߣ AMY1 ࠦࡇ࡯ᢙߣߩ㑐ㅪᕈߦߟ޿ߡ߽ᬌ⸛⺖㗴ߣߒߡ⸳ቯߒߡ߅ࠅ‫੹ޔ‬ᓟ⹦⚦ߥಽᨆࠍⴕ߁੍
ቯߢ޽ࠆ‫ޕ‬
D-22 ࠨ࡞⣄㜑↱᧪㑆⾰♽ᐙ⚦⢩ߩၭ㙃ߣߘߩ⒖ᬀߦࠃࠆ࡜࠶࠻⣄㜑៊்ୃᓳലᨐߩᬌ⸛
ฎᎹᤘᩕ㧘⑔శ⑲ᢥ㧘ቬችੳ⟤㧔ጘ㒂⮎⑼ᄢ㧕 ᚲౝኻᔕ⠪㧦ᄢ⍹㜞↢
⣄㜑ߪᧃ᪳⚵❱ߣ⣖ߩㆇേ‫ޔ‬⍮ⷡᖱႎߩવ㆏〝ߢ޽ࠅ்ኂߐࠇࠆߣ‫ޔ‬ㆇേ㤗∽߿⍮ⷡ㓚ኂߥߤߩ㊀◊ߥり૕㓚ኂ
ߦ㒱ࠆ‫ޕ‬ᚒ‫ޔߪޘ‬⣄㜑៊்ࡕ࠺࡞࡜࠶࠻ߩ⣄㜑ታ⾰ౝߦ㜞Ớᐲߩ FGF2 ࠍᵈ౉ߔࠆߣㆇേᯏ⢻߇㗼⪺ߦᡷༀߔࠆߎ
ߣࠍ⷗಴ߒߚ‫ޔ߈ߣߩߎޕ‬⣄㜑ౝߢჇᱺߒߡ޿ߚ✢⛽⧘⚦⢩ࠍၭ㙃ਅߢჇ߿ߒ‫்៊ޔ‬ㇱ૏ߦ⒖ᬀߒߚߣߎࠈ‫ޔ‬FGF2
ᛩਈࠃࠅ߽ᦝߦ㗼⪺ߥㆇേᯏ⢻ᡷༀ߇⹺߼ࠄࠇߚ‫ޔߢߎߘޕ‬㔤㐳㘃ߦ߅ߌࠆ㘃ૃߩ✢⛽⧘⚦⢩ߩ᦭ήࠍ⏕⹺ߔࠆߚ
߼‫ޔ‬એਅߩࠃ߁ߥ౒ห೑↪⎇ⓥࠍታᣉߒߚ‫࡞ࠨޕ‬⣄㜑ࠍ៰಴ᓟ‫ޔ⤑⎬ޔ‬㜑⤑╬ࠍ೸㔌ߒ‫ޔ‬⣄㜑ታ⾰ㇱࠍ⭯ಾߒ 0.3-1.0
mm ෘߩ⚵❱ ࠍᓧߚ‫⭯ߦࡘࠪࠖ࠺࠻࡯ࠦࡦࠥ࡯࡜ࠦޕ‬ಾ⚵❱ ࠍਗߴ‫ޔ‬10 ng/ml FGF2‫ޔ‬10% FBS ࠍ฽߻ DMEM
ၭ࿾ࠍṢਅߒ‫ޔ‬CO2 ࠗࡦࠠࡘࡌ࡯࠲ౝߦ㕒⟎ߒߚ‫ޕ‬⠉ᣣ‫ޔ‬ಾ ߇ᶐࠆ⒟ᐲߦၭ࿾ࠍㅊടߒ‫⚦ࠄ߆ ❱⚵ޔ‬⢩߇ㆆ⿛
ߒߡߊࠆ߹ߢ‫⚳ޔ‬Ớᐲ 10 ng/ml ߣߥࠆࠃ߁ߦ FGF2 ࠍᲤᣣᷝടߒ⛯ߌߚ‫᧦ᧄޕ‬ઙߦ߅޿ߡ‫࡞ࠨޔ‬⣄㜑߆ࠄ߽࡜࠶࠻
⣄㜑ߣห᭽ߦ✢⛽⧘⚦⢩߇ㆆ⿛ߒߡߊࠆߎߣ߇⏕⹺ߐࠇߚ‫⃻ޕ‬࿷‫ޔ‬ห✢⛽⧘⚦⢩ߩၭ㙃ࠍ⛮⛯ਛߢ޽ࠆ‫੹ޕ‬ᓟ‫࡜ޔ‬
࠶࠻⣄㜑៊்ࡕ࠺࡞ߦห⚦⢩ࠍ⒖ᬀߒ‫ޔ‬ㆇേᯏ⢻ߩ࿁ᓳലᨐࠍ᦭ߔࠆ߆ߤ߁߆ࠍᬌ⸛ߔࠆ੍ቯߢ޽ࠆ‫ޕ‬
(5) 㔡ἴ㑐ㅪ
E㧔㔡ἴ A-1㧕㊄⪇ጊፉߦ↢ᕷߔࠆ㊁↢࠾ࡎࡦࠩ࡞ߩ୘૕ᢙ⺞ᩏ
દᴛ⚔↢㧔ችၔߩࠨ࡞⺞ᩏળ㧕㧘ਛᎹዏผ㧔੩ㇺᄢ࡮㒮࡮ℂ㧕㧘⮮↰ᔒᱠ㧔㣮ఽፉᄢ࡮ㄘ㧕㧘㘑ᒛ༑ሶ㧔੩ㇺᄢ࡮
㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯㧕㧘Ꮉᷝ㆐ᦶ㧔੩ㇺᄢ࡮㒮࡮ℂ࡮↢‛⑼ቇ㧕㧘ቝ㊁ჽᤐ㧘㑐ஜᄥ㇢㧘ਃᧁᷡ㓷㧔(ว)࡮ችၔ㨯
㊁↢േ‛଻⼔▤ℂ࠮ࡦ࠲࡯㧕 ᚲኻᔕ⠪㧦ฎᏒ೰ผ
ᤓᐕ 11 ᦬ᓟඨߣᧄᐕ 3 ᦬ᓟඨߩ⸘ 2 ࿁‫ޔ‬2012 ᐕᐲߩ୘૕ᢙߦ㑐ߔࠆ৻ᢧ⺞ᩏࠍ‫ޔ‬ፉߦ↢ᕷߔࠆ 6 ⟲ߣࠝࠬࠣ࡞
࡯ࡊ‫ࠍ࡞ࠩ࡟࠽ࡂޔ‬ኻ⽎ߦ଀ᐕㅢࠅታᣉߒߚ‫⚿ޕ‬ᨐߪ 11 ᦬ߩ⺞ᩏߢߪ 259 㗡‫ޔ‬3 ᦬ߩ⺞ᩏߢߪ 254 㗡ߛߞߚ‫ޕ‬
ߎࠇࠄ 2 ࿁ߩ৻ᢧ⺞ᩏߣߘߩ೨ᓟߦⴕߞߚ୘೎⺞ᩏߢ‫ޔ‬3.11 ᧲ᣣᧄᄢ㔡ἴߢਛᱛࠍ૛௾ߥߊߐࠇߚᤓᐕ 3 ᦬ᤨὐ
ߢߩ 2011 ᐕᐲߩ୘૕ᢙࠍᓳరߔࠆߎߣ߇ߢ߈ߚ‫ޕ‬㗡ᢙߪ 238 㗡ߢ޽ࠆ‫ޔߚ߹ޕ‬2012 ᐕᐲߩ಴↢ᢙߣࠕࠞࡦࡏ࠙ߩ
౻ᦼᱫ੢₸߽޽ࠊߖ⺞ᩏߒߚ‫⚿ޕ‬ᨐߪ಴↢ᢙ߇ 35 㗡‫ޔ‬౻ᦼᱫ੢₸߇ 5.7%ߛߞߚ‫ޕ‬
3.11 Ꮒᄢ࿾㔡ߩࠨ࡞߳ߩᓇ㗀‫߮ࠃ߅ޔ‬Ꮒᄢ࿾㔡ߦࠃࠆ࿾⋚ߩ✭ߺߣ 9 ᦬ 21 ᣣߩ㓸ਛ⽕㔎߇ේ࿃ߢፉߓࠀ߁ߢ⿠
ߎߞߚ࿯⍹ᵹ߿ᕆᢳ㕙ߩ፣⪭ߩࠨ࡞߳ߩᓇ㗀ߦߟ޿ߡߪ‫ߩߢ⏷ޔ‬ᶏ⮺ߩ⢒ᚑ߿⑺ߩၷᨐ㧔࠽࠶࠷㘃㧕ߩ⒤ࠅߣ޿ߞ
ߚࠨ࡞ߩ㘩‛ߣࠨ࡞ߩ୘૕ᢙߩᄌൻߩਔ㕙߆ࠄ⺞ᩏߒߚ‫⚿ߩߘޕ‬ᨐߪ‫ޔ‬ਔᣇߣ߽ࠨ࡞ߦኻߔࠆ㗼⪺ߥᓇ㗀ߪ⹺߼ࠄ
ࠇߥ߆ߞߚ‫ޕ‬
ߚߛ‫ޔ‬Ꮒᄢ࿾㔡߿࿯⍹ᵹߩࠨ࡞߳ߩᓇ㗀ߪ‫ޔ‬ᒰ⹥ᐕᐲߛߌߢߪߥߊ‫ߦࠄߐޔ‬㐳޿ᤨ㑆᏷ߩਛߢ಴ߡߊࠆน⢻ᕈ߽
޽ࠅ‫੹ޔ‬ᓟߣ߽ᵈᗧᷓߊ⛮⛯ߒߡ⺞ᩏߒߡ޿ߊᔅⷐ߇޽ࠆߛࠈ߁‫ޕ‬
ߥ߅‫⎇ᧄޔ‬ⓥߢታᣉߒߚฦ⒳⺞ᩏߩ⚿ᨐߦߟ޿ߡࠃࠅ⹦ߒߊߪ 9 ᦬ᤨὐ߹ߢߩ߹ߣ߼ߪ‫ޟ‬㔤㐳㘃⎇ⓥ‫ޠ‬27 ภߢ
ႎ๔ߒ‫ޔ‬3 ᦬ᤨὐ߹ߢߩ߹ߣ߼ߪห⹹ 28 ภߦ⃻࿷ᛩⓂਛߢ޽ࠆ‫ޕ‬ෳᾖߦߐࠇߚ޿‫ޕ‬
㧙㧙
3. ᐔᚑ 23 ᐕᐲߢ⚳ੌߒߚ⸘↹⎇ⓥ
ᣥ਎⇇ࠩ࡞ߩᄌ⇣ᕈߣㅴൻߦ㑐ߔࠆᄙ㕙⊛ࠕࡊࡠ࡯࠴
ታᣉ੍ቯᦼ㑆㧦ᐔᚑ 21㨪23 ᐕᐲ
⺖㗴ផㅴ⠪㧦㜞੗ᱜᚑ㧘⷏᧛೰㧘ᳯᧁ⋥ሶ㧘ࡑࠗࠢ࡮ࡂࡈࡑࡦ㧘Ძ೑ବ㓶
ᧄ⸘↹⎇ⓥߪ‫ޔ‬ᣥ਎⇇ࠩ࡞㘃㧔ࠝ࠽ࠟࠩ࡞⑼㧕ߩᄌ⇣ᕈߣㅴൻߦ㑐ߒߡ‫ޔ‬᭽‫ߥޘ‬ಽ㊁߆ࠄߩᄙ㕙⊛ߥࠕࡊࡠ࡯࠴
ߦࠃࠆ⎇ⓥࠍ⋡ᜰߒߡⴕߞߚ‫࡞ࠩࠟ࠽ࠝߪ࡞ࠨߩ⑼࡞ࠩࠟ࠽ࠝޕ‬੝⑼ߣࠦࡠࡉࠬ੝⑼ߩੑߟߩࠣ࡞࡯ࡊ߆ࠄߥࠆ߇‫ޔ‬
ߘߩᒻᘒ࡮㘩ᕈ࡮ⴕേࡄ࠲࡯ࡦߥߤߦᄢ߈ߥᄌ⇣߇ሽ࿷ߔࠆ‫ߚߞ޿߁ߎޕ‬ᣥ਎⇇ࠩ࡞ߩᄙ᭽ᕈߣߘߩㅴൻߦ㑐ߒߡ‫ޔ‬
᭽‫⎇ߥޘ‬ⓥಽ㊁ߩᚻᴺࠍ↪޿ߡ⎇ⓥࠍផㅴߒߚ‫⎇ޕ‬ⓥኻ⽎ߪ⃻↢⒳ߣൻ⍹⒳ߩਔᣇߦ߹ߚ߇ࠅ‫ޔ‬㗡㛽࡮ਅ㗶㛽࡮ᱤ࡮
ᬁ㛽࡮⿷ᩮ㛽࡮ᄢ⣽㛽ߥߤ‫ޔ‬᭽‫ߥޘ‬ㇱ૏ߩᒻᘒߦߟ޿ߡߩ⎇ⓥ߇ⴕࠊࠇߚ‫ޕ‬વ⛔⊛ߥࡁࠡࠬࠍ૶ߞߚ⸘᷹‫ᦨޔ‬ㄭߩ
ᵹⴕߢ޽ࠆធ⸅ဳߩ 3 ᰴర⸘ེ᷹ࠍ↪޿ߚᐞ૗ቇ⊛ᒻᘒ⸘᷹‫ޔ‬X ✢ CT ࠍ↪޿ߚ㛽ౝㇱߩ᭴ㅧ⸃ᨆ‫ޔ‬ᵢ─ౝߩࠨ࡞
ߩ㛽ᩰൻ⍹ߣ♮ߩಽᨆߥߤ‫ޔ‬ᄙጘߦࠊߚࠆᚻᴺ߇↪޿ࠄࠇߚ‫⎇ߩࠄࠇߎޕ‬ⓥ⚿ᨐߩ৻ㇱߪ‫ޔ‬ᐔᚑ 23 ᐕ 3 ᦬ 7㨪8 ᣣ
ߦ㐿௅ߐࠇߚ౒ห೑↪⎇ⓥળ‫ޟ‬CT ࠍ↪޿ߚ㔤㐳㘃⎇ⓥߩᣂዷ㐿‫ޕߚࠇߐ⴫⊒ߢޠ‬
⎇ⓥታᣉ⠪
㧨ᐔᚑ 21 ᐕᐲ㧪
ዊ⮳ᄢテ 㧔᧲੩ᄢ࡮㒮࡮ℂቇ♽㧕‫߮ࠃ߅↢⃻ޟ‬ൻ⍹ࠦࡠࡉࠬ㘃ߦ߅ߌࠆㅴൻᒻᘒቇ⊛⎇ⓥ‫ޠ‬
᧲ ⪇ጪ 㧔ጘ㒂ᄢ࡮කቇ⎇ⓥ⑼㧕‫ޟ‬㔤㐳㘃ᬁ㛽ߦ߅ߌࠆਃᰴర↹௝ߩ㔚⣖⸃ᨆ‫ޠ‬
ㄭ⮮ାᄥ㇢㧔ᗲ⍮ቇ㒮ᄢ࡮ᱤ࡮⸃೬㧕‫ޟ‬ᣥ਎⇇ࠩ࡞ਅ㗶㛽ᄖ஥㕙ߦߺࠄࠇࠆ㓉⿠ߩ⒳㑆ᄌ⇣‫ޠ‬
ੑ␹ජᤐ 㧔ᗲ⍮ቇ㒮ᄢ࡮㒮࡮ᱤ㧕‫ࠆߌ߅ߦ࡞ࠩࡦࡎ࠾ޟ‬਄㗶੃⥓ᱤ‫ޔ‬ዊ⥓ᱤ‫ޔ‬ᄢ⥓ᱤߩᱤ౰ࠨࠗ࠭ߩ㑐ଥ‫ޠ‬
ᆌፒᥓሶ 㧔⟲㚍⋵ඳ㧕‫ޟ‬⠨ฎㆮ〔಴࿯࠾ࡎࡦࠩ࡞ߩ㛽ᒻᘒߩ࿾ℂ⊛ᄌ⇣ߦ㑐ߔࠆ⎇ⓥ‫ޠ‬
㧨ᐔᚑ 22 ᐕᐲ㧪
᧲ ⪇ጪ 㧔ጘ㒂ᄢ࡮කቇ⎇ⓥ⑼㧕‫ޟ‬㔤㐳㘃ᄢ⣽㛽㗪ㇱߦ߅ߌࠆਃᰴర↹௝ߩ㔚⣖⸃ᨆ‫ޠ‬
ᆌፒᥓሶ 㧔⟲㚍⋵ඳ㧕‫߮ࠃ߅↢⃻ޟ‬⠨ฎㆮ〔಴࿯࠾ࡎࡦࠩ࡞ߩ㛽ᒻᘒᄌ⇣ߦ㑐ߔࠆ⎇ⓥ‫ޠ‬
ዊ⮳ᄢテ 㧔੩ㇺᄢ࡮✚วඳ㧕‫߮ࠃ߅↢⃻ޟ‬ൻ⍹ࠦࡠࡉࠬ㘃ߦ߅ߌࠆㅴൻᒻᘒቇ⊛⎇ⓥ‫ޠ‬
ၔࡩේࠁ߁㧔ጟጊℂᄢ࡮㒮࡮✚วᖱႎ㧕‫ޟ‬㔤㐳㘃ߩあ㛽෸߮〒㛽ߦ߅ߌࠆ୘૕⊒↢‫ޠ‬
ᒛ Ⓨᄸ 㧔ਛ࿖⑼ቇ㒮࡮ฎ⣄ᬁേ‛㧛ฎੱ㘃⎇㧕‫ޟ‬ਛ࿖ᐢ⷏߆ࠄ↥಴ߒߚ೨ᦼᦝᣂ਎ࡑࠞࠢߩోり㛽ᩰൻ⍹ߩ
Ყセ⸃೬ቇߣᯏ⢻⸃೬ቇ⊛⎇ⓥ‫ޠ‬
⍫㊁ ⥶ 㧔੩ㇺᄢ࡮ℂ㧕‫࡞ࠩࠟ࠽ࠝޟ‬ᣖߩ⡬ེⷡቭߩᯏ⢻ᒻᘒቇ⊛ㅴൻߦ㑐ߔࠆ⎇ⓥ‫ޠ‬
ㄭ⮮ାᄥ㇢㧔ᗲ⍮ቇ㒮ᄢ࡮ᱤ࡮⸃೬㧕‫࡞ࠩࠟ࠽ࠝޟ‬੝⑼ߩਅ㗶㛽ᄖ஥㕙ߦߺࠄࠇࠆ㓉⿠ߩട㦂ᄌ⇣‫ޠ‬
㧨ᐔᚑ 23 ᐕᐲ㧪
ዊ⮳ᄢテ 㧔੩ㇺᄢ࡮✚วඳ㧕‫߮ࠃ߅↢⃻ޟ‬ൻ⍹ࠝ࠽ࠟࠩ࡞㘃ߦ߅ߌࠆㅴൻᒻᘒቇ⊛⎇ⓥ‫ޠ‬
㎄ᧄᱞਭ 㧔ᨋේ↢‛ൻቇ⎇࡮ฎ↢‛࠮ࡦ࠲࡯㧕‫↢⃻ޟ‬ᣥ਎⇇ࠩ࡞ߦ߅ߌࠆ〒㛽ߩᄌ⇣ߣൻ⍹߳ߩᔕ↪‫ޠ‬
ᆌፒᥓሶ 㧔⟲㚍⋵ඳ㧕‫߮ࠃ߅↢⃻ޟ‬⠨ฎㆮ〔಴࿯࠾ࡎࡦࠩ࡞ߩ㛽ᒻᘒᄌ⇣ߦ㑐ߔࠆ⎇ⓥ‫ޠ‬
᧲ ⪇ጪ 㧔ጘ㒂ᄢ࡮කቇ⎇ⓥ⑼㧕‫ޟ‬㔤㐳㘃ᬁ㛽ߩᄖㇱᒻ⁁ߣౝㇱ᭴ㅧߩ⛔ว⸃ᨆ‫ޠ‬
ᨰᧁஜม࡮㒙ㇱാᴦ࡮ἑਯญ␽ቁ㧔ንጊᄢ࡮ℂᎿቇㇱ㧕
‫ࠆߥ⇣ޟ‬᳇୥ਅߦ߅ߌࠆ࠾ࡎࡦࠩ࡞ൻ⍹ߩ㛽૕ᒻ⾰Ყセ‫ޠ‬
㧔ᢥ⽿㧦㜞੗ᱜᚑ㧕
4. ౒ห೑↪⎇ⓥળ
╙ 12 ࿁࠾ࡎࡦࠩ࡞⎇ⓥ࠮ࡒ࠽࡯
ᣣᤨ㧦2011 ᐕ 6 ᦬ 11 ᣣ࡮12 ᣣ
႐ᚲ㧦੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲᄢળ⼏ቶ
⎇ⓥળ਎⹤ੱ㧦ඨ⼱๋㇢㧘ㄞᄢ๺㧔੩ᄢ࡮㔤㐳㘃⎇ⓥᚲ㧕
࠾ࡎࡦࠩ࡞⎇ⓥ࠮ࡒ࠽࡯ߪ‫ߢ߹ࠇߎޔ‬ㆊ෰ 9 ᐕߦᷰߞߡ‫౒ޔ‬ห೑↪⎇ⓥળ߿⥄ਥ⊛ߥ㓸ળߣߒߡታᣉߒߡ߈ߚ‫ޕ‬
ߎߩ⎇ⓥળߢߪ‫ࠍ࡞ࠩࡦࡎ࠾ޔ‬ኻ⽎ߣߒߚࡈࠖ࡯࡞࠼ߩ⎇ⓥ⠪߇‫ޔ‬੤ᵹߒ⸛⺰ߢ߈ࠆ႐ࠍ૞ࠆߎߣࠍ⋡⊛ߣߒߡ޿
ࠆ‫ ╙ޕ‬12 ࿁⋡ߣߥࠆ੹࿁߽⧯ᚻ⎇ⓥ⠪ߩᣇߦୃ჻⺖⒟߿ඳ჻⺖⒟ߢߩ⎇ⓥᚑᨐࠍਛᔃߦ⊒⴫ࠍ߅㗿޿ߒ‫ޔ‬ਛၷ㨯
ࡌ࠹࡜ࡦ⎇ⓥ⠪߇‫ߦࠇߘޔ‬ኻߒߡࠦࡔࡦ࠻ߔࠆߣ޿߁ࠬ࠲ࠗ࡞ߢⴕࠊࠇߚ‫౏ࠍ⴫⊒࡯࠲ࠬࡐޔߚ߹ޕ‬൐ߒ‫ୃޔ‬჻࡮
ඳ჻⺰ᢥߩㅜਛ⚻ㆊߥߤߦߟ޿ߡ⊒⴫ߒߡ߽ࠄ߁ᯏળࠍ⸳ߌߚ‫ޕ‬48 ฬߩᣇߦෳട޿ߚߛ߈‫ޔ‬ᵴ⊒ߥ⼏⺰ࠍߔࠆߎ
ߣ߇ߢ߈ߚ‫ޕ‬
㧨ࡊࡠࠣ࡜ࡓ㧪
6 ᦬ 11 ᣣ㧔࿯㧕
12:58㨪13:00 ᜿ᜦ ඨ⼱๋㇢㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕
13:00㨪14:00 ᢪ⮮᣽ᐘ 㧔ᮮᵿ࿖┙ᄢቇᄢቇ㒮 ⅣႺᖱႎቇᐭ㧕ㇺᏒ߆ࠄ᫪ᨋߦ⥋ࠆ᥊ⷰ௑ᐲߣ㊁↢ູ੃㘃
㧙㧙
14:00㨪15:00 ਛ᧛ᄢテ 㧔ጘ㒂ᄢቇᄢቇ㒮 ㅪวㄘቇ⎇ⓥ⑼㧕ㇺᏒㄭ㇠࿾ၞߦ߅ߌࠆ࠾ࡎࡦࠩ࡞ߦࠃࠆⵍኂᗧ⼂ߩ
ታᘒ
15:00㨪15:15 ભᙑ
15:15㨪16:15 ဈ’ߪࠆ߆㧔ጤᚻᄢቇᄢቇ㒮 ㅪวㄘቇ⎇ⓥ⑼㧕಄᷷Ꮺᨋߦ߅ߌࠆ࠾ࡎࡦࠩ࡞㊁↢⟲ߩ౻ᦼ᫪ᨋ೑↪
ߦ㑐ߔࠆⓨ㑆⊛⹏ଔ
16:15㨪17:15 Andrew MacIntosh㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕Epidemiology of nematode parasite infection among wild
Japanese macaques: heterogeneity in the external and internal environments
17:15㨪18:00 ࡐࠬ࠲࡯⊒⴫
18:00㨪20:15 ᙣⷫળ
6 ᦬ 12 ᣣ㧔ᣣ㧕
9:00㨪10:00 ᄢ⼱ᵗ੺ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕࠾ࡎࡦࠩ࡞㓶ߩ㓸࿅߆ࠄߩ৻ᤨ㔌⣕ⴕേ㧦ណ㘩‫❥ޔ‬ᱺᚢ⇛ߩ
ⷰὐ߆ࠄ ࠦࡔࡦ࠹࡯࠲㧦ਛᎹዏผ㧔੩ㇺᄢቇᄢቇ㒮 ℂቇ⎇ⓥ⑼㧕
10:00㨪11:00 ㋈ᧁධ⟤ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕ㆮવሶ࡮⚦⢩࡮୘૕࡟ࡌ࡞߆ࠄߺߚ࠾ࡎࡦࠩ࡞ߩ⧰๧ᗵⷡ ࠦ
ࡔࡦ࠹࡯࠲㧦ᴡ᧛ᱜੑ㧔᧲੩ᄢቇᄢቇ㒮 ᣂ㗔ၞഃᚑ⑼ቇ⎇ⓥ⑼㧕
11:00㨪11:10 ભᙑ
11:10㨪12:10 ਄㊁዁ᢘ 㧔ᄢ㒋ᄢቇᄢቇ㒮 ੱ㑆⑼ቇ⎇ⓥ⑼㧕ൎጊ࠾ࡎࡦࠩ࡞㓸࿅ߦ߅ߌࠆᲫߠߊࠈ޿ߩ㚟ߌᒁ߈
ߣ੕ᕺᕈ ࠦࡔࡦ࠹࡯࠲㧦ዊᎹ⑲ม㧔ਛ੩ᄢቇ㧕
ࡐࠬ࠲࡯⊒⴫
P-1 ᦸ᦬⠍ᄥ㧔ᣂẟᄢቇᄢቇ㒮࡮⥄ὼ⑼ቇ⎇ⓥ⑼㧕᥊ⷰ᭴ㅧߩ㆑޿߇࠾ࡎࡦࠩ࡞ߩㄘ࿾ଚ౉ߦਈ߃ࠆᓇ㗀
P-2 ㊄⼱⠍ᄥ㧔ᣂẟᄢቇᄢቇ㒮࡮⥄ὼ⑼ቇ⎇ⓥ⑼㧕࠾ࡎࡦࠩ࡞ߩ▤ℂࠍ⋡⊛ߣߒߚၷᨐ㘃ߩࡑ࠶ࡇࡦࠣ
P-3 Ⴧ㑆ᜏ਽㧔㐳ጟᛛⴚ⑼ቇᄢቇ࡮↢‛ᯏ⢻Ꮏቇኾ᡹㧕േ‛ኾ↪๟ᵄᢙᏪࠍ೑↪ߒߚ࠾ࡎࡦࠩ࡞ߩ࡝ࠕ࡞࠲ࠗࡓࡕ
࠾࠲࡝ࡦࠣࠪࠬ࠹ࡓ
P-4 ൎ㊁ฮሶ㧔ᄢ㒋ᄢቇᄢቇ㒮࡮ੱ㑆⑼ቇ⎇ⓥ⑼㧕ࡢࠞࡕࡁᦼ࠾ࡎࡦࠩ࡞ߦ߅ߌࠆᲣᆷ㑐ଥߣᚑ૕ߣߩᲫߠߊࠈ޿
㑐ଥߩᒻᚑ
P-5 ᶏ⠧ේኡ࡮㜞ᮎᚑ♿㧔㤗Ꮣᄢቇᄢቇ㒮㧕ㄘ࿾ࠍ೑↪ߔࠆࠃ߁ߦߥߞߚ࠾ࡎࡦࠩ࡞⟲ߩ⟲⪭೑↪
㧔ᢥ⽿㧦ඨ⼱๋㇢㧕
‫␠ޟ‬ળⴕേߣ⣖ ̆ࠗࡔ࡯ࠫࡦࠣߣಽሶ̆‫ޠ‬
ᣣᤨ㧦2012 ᐕ 2 ᦬ 2 ᣣ㧔ᧁ㧕
࡮3 ᣣ㧔㊄㧕
႐ᚲ㧦੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲᄢળ⼏ቶ
⎇ⓥળ਎⹤ੱ㧦ਛ᧛స᮸㧘㜞↰᣽ᒾ㧘ች࿾㊀ᒄ㧘ᄢ⍹㜞↢
ࡅ࠻ࠍ㗂ὐߣߒߚⶄ㔀ߥ␠ળⴕേߪ㔤㐳㘃ߩⴕേ․ᕈߢ޽ࠆ‫␠ߚߒ߁ߎޕ‬ળⴕേߩၮߦߥࠆ␹⚻ᯏᐨ߇‫ޔ‬ᓢ‫ߦޘ‬
⑼ቇ⊛ߦ᣿ࠄ߆ߦߐࠇߡ߈ߡ޿ࠆ‫ޕ‬ᄢ߈ߊੑߟߩ⎇ⓥಽ㊁߇޽ࠆ‫ߪߟ৻ߩߘޕ‬㨒MRIࠍਛᔃߣߒߚࡅ࠻ߩ⣖ᯏ⢻ࠗ
ࡔ࡯ࠫࡦࠣ⎇ⓥߢ޽ࠆ‫ࠍࡦࠪ࠻ࠪࠠࠝޔߪߟ৻߁߽ޕ‬ਛᔃߣߒߚࡎ࡞ࡕࡦߩⴕേ߳ߩᓇ㗀ࠍ⺞ߴࠆ⎇ⓥߢ޽ࠆ‫੹ޕ‬
ᓟߩ㔤㐳㘃⎇ⓥߩᄢ߈ߥ࠹࡯ࡑߣߥࠆߢ޽ࠈ߁␠ળⴕേߣ⣖ᯏ⢻ߩ㑐ଥࠍ‫ޔ‬਎⇇ߩ࠻࠶ࡊ࡟ࡌ࡞ߢ⎇ⓥߒߡ޿ࠆ⎇
ⓥ⠪ߦෳട޿ߚߛ߈‫੹ޔ‬ᓟ․ߦࠨ࡞㘃ߢⴕߞߡ޿ߊߴ߈⎇ⓥߩᣇ㊎ࠍᬌ⸛ߔࠆ⋡⊛ߢ㐿௅ߒߚ‫ߞߍޕ‬ᱤ㘃߆ࠄࡅ࠻
߳ߩᯅᷰߒߣߒߡߩᓎഀ‫ⴕࠆࠇࠄߺߦ࠻ࡅޔ‬േߩ␹⚻ᯏᐨࠍ⺞ߴࠆኻ⽎ߣߒߡߩᓎഀ‫ޔ‬ഃ⮎⎇ⓥߦ߅ߌࠆᓎഀ╬‫ޔ‬
ࠨ࡞㘃ߩᓎഀ߇᣿ࠄ߆ߦߢ߈ߚ‫ߘࠃ߅߅ޕ‬40ฬߩᣇߦෳട޿ߚߛ߈‫ޔ‬ᵴ⊒ߥᗧ⷗੤឵߿⼏⺰߇ߢ߈ߚ‫ޕ‬
㧨ࡊࡠࠣ࡜ࡓ㧪
2᦬2ᣣ
13:00㨪13:05 ᜿ᜦ ਛ᧛స᮸㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕
1ㇱ ࠝࠠࠪ࠻ࠪࡦ ᐳ㐳㧦ਛ᧛స᮸㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕
13:05㨪13:50 ᧻੗⑲᮸㧔ጟጊᄢቇᄢቇ㒮 කᱤ⮎ቇ✚ว⎇ⓥ⑼㧕‫ޟ‬⣖ౝࡎ࡞ࡕࡦ࡮ࠝࠠࠪ࠻ࠪࡦߦࠃࠆਇ቟ᖱേ࡮
߁ߟ∝⁁ߩ೙ᓮ‫ޠ‬
13:50㨪14:35 ንỈ৻ੳ㧔ᾢᧄᄢቇᄢቇ㒮 ↢๮⑼ቇ⎇ⓥㇱ㧕‫ࠆࠃߦࡦࠪ࠻ࠪࠠࠝޟ‬ઍ⻢೙ᓮ‫ޠ‬
14:35㨪15:20 ⷏᫪సᒾ㧔᧲ർᄢቇᄢቇ㒮 ㄘቇ⎇ⓥ⑼㧕‫ޟ‬ㆮવሶᄌ឵ࡑ࠙ࠬࡕ࠺࡞ࠍ↪޿ߚਛᨔᕈࠝࠠࠪ࠻ࠪࡦฃ
ኈ૕ߩᯏ⢻⸃ᨆ‫ޠ‬
15:35㨪16:05 ਛ᧛స᮸㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕‫ⴕⷫߩ࠻࠶࠮ࡕ࡯ࡑޟ‬േߦ߅ߌࠆࠝࠠࠪ࠻ࠪࡦߩᓇ㗀‫ޠ‬
16:05㨪16:50 ᧲↰㓁ඳ㧔㊄ᴛᄢቇᄢቇ㒮 කቇ♽⎇ⓥ⑼㧕‫ޟ‬CD38ߦࠃࠆࠝࠠࠪ࠻ࠪࡦಽᴲ೙ᓮ‫␠ޔ‬ળᕈⴕേߣ⥄
㐽∝‫ޠ‬
2᦬3ᣣ
2 ㇱ ࠗࡔ࡯ࠫࡦࠣ ᐳ㐳㧦ች࿾㊀ᒄ㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕
9:30㨪10:15 የ਄ᶈ㓉㧔⁛┙ᴺੱ ℂൻቇ⎇ⓥᚲ ಽሶࠗࡔ࡯ࠫࡦࠣ⑼ቇ⎇ⓥ࠮ࡦ࠲࡯㧕
‫⹺࡞ࠨޟ‬⍮࡮␠ળⴕേߦ߅
㧙㧙
ߌࠆ⣖ᯏ⢻ߩ PET ࠗࡔ࡯ࠫࡦࠣ⸃ᨆ‫ޠ‬
10:15㨪11:00 ᵏ⟜㓷⊓㧔᧲੩ක⑼ᱤ⑼ᄢቇ㧕‫ޟ‬⣖ᵴേ߆ࠄᔃࠍ⺒ߺขࠆ‫ޠ‬
11:00㨪11:45 㘵㜞ື਽㧔ฬฎደᄢቇᄢቇ㒮 කቇ♽⎇ⓥ⑼㧕‫␠ޟ‬ળ⣖ߦ߅ߌࠆᚳ᩶૕ߩᓎഀ‫ޠ‬
11:45㨪12:30 ᧻రஜੑ㧔₹Ꮉᄢቇ ⣖⑼ቇ⎇ⓥᚲ㧕‫ߩ࠻ࡅޟ‬േᯏߠߌߩ⣖ᯏ⢻ࠗࡔ࡯ࠫࡦࠣ‫ޠ‬
㧔ᢥ⽿㧦ਛ᧛స᮸㧕
‫ⴕޟ‬േ․ᕈࠍᡰ㈩ߔࠆࠥࡁࡓၮ⋚ߣ⣖ᯏ⢻ߩ⸃᣿‫ޠ‬
ᣣᤨ㧦2012 ᐕ 3 ᦬ 2 ᣣ㧔㊄㧕13:30㨪3 ᦬ 3 ᣣ㧔࿯㧕15:00
႐ᚲ㧦੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲᄢળ⼏ቶ
⎇ⓥળ਎⹤ੱ㧦㜞↰᣽ᒾ
ⴕേ․ᕈߩ㕙߆ࠄࡅ࠻ߣࠨ࡞ࠍᲧセߔࠆߣ‫∔ߢ࠻ࡅޔ‬ᖚဳߩⴕേ߇ࠨ࡞ߢߪ৻⥸⊛ߥⴕേߣᝒ߃ࠄࠇ‫ޔ‬ㅒߦࡅ࠻
ߢ৻⥸⊛ߥⴕേ߇ࠨ࡞ߢߪ⇣Ᏹߥⴕേߣᝒ߃ࠄࠇࠆ‫⥝߁޿ߣޔ‬๧ᷓ޿੐⽎߇޽ࠆ‫ޕ‬㘃ૃߩ੐⽎ߪࠥࡁࡓ࡟ࡌ࡞ߦ߅
޿ߡ߽⏕⹺ߐࠇߡ޿ࠆ‫ⴕޕ‬േߣࠥࡁࡓࠍࠠ࡯ࡢ࡯࠼ߦߒߡߎߩࠃ߁ߥࡅ࠻ߣࠨ࡞ߩ⋧੕㑐ଥࠍ⸃ᨆߔࠆߎߣߪ‫ࡅޔ‬
࠻ߩⴕേ․ᕈ‫∔⚻␹࡮␹♖ޔ‬ᖚߩࡔࠞ࠾࠭ࡓ‫ੱޔߦࠄߐޔ‬㘃ߩㅴൻߩਛߢ‫ߪߣ࠻ࡅޟ‬૗߆‫⸃ࠍޠ‬᣿ߔࠆߚ߼ߩ㊀ⷐ
ߥ♻ญߣߥࠆน⢻ᕈࠍ⒁߼ߡ޿ࠆ‫ޕ‬ᐔᚑ 23 ᐕᐲ߆ࠄ㐿ᆎߐࠇߚ౒ห೑↪㨯౒ห⎇ⓥࡊࡠࠫࠚࠢ࠻ߩ⸘↹⎇ⓥ‫ⴕޟ‬േ
․ᕈࠍᡰ㈩ߔࠆࠥࡁࡓၮ⋚ߣ⣖ᯏ⢻ߩ⸃᣿‫(ޔߪߢޠ‬1)ਥߦࡑࠞࠢ㘃ࠍ↪޿ߚࠥࡁࡓߩ✂⟜⊛⸃ᨆࠍታᣉߒ‫ޔ‬ᄙ᭽ߥ
ⴕേ․ᕈࠍ␜ߔ⥄ὼ⊒↢⊛ㆮવሶᄌ⇣ࡕ࠺࡞ࠍࠥࡁࡓ࡟ࡌ࡞ߣ⣖ᯏ⢻࡟ࡌ࡞ߢ⸃ᨆߔࠆ‫(ޔ‬2)♖␹⑼㗔ၞߣㅪ៤ߒߡ‫ޔ‬
⣖∛ᘒࠥࡁࡓᄙဳߣਛ㑆⴫⃻ဳߦ㑐ߔࠆ࠺࡯࠲ࡌ࡯ࠬࠍᵴ↪ߒ‫∔␹♖ޔ‬ᖚߦ㑐ㅪߔࠆ࡝ࠬࠢㆮવሶࠍࡑࠞࠢ㘃ߦ߅
޿ߡ✂⟜⊛ߦᬌ⚝ߔࠆ‫(ޔ‬3)ࠥࡁࡓᖱႎߦၮߠ޿ߡੱὑ⊛ㆮવሶᡷᄌࡕ࠺࡞ࠍ૞಴ߒ‫ⴕߩߘޔ‬േ․ᕈߣ⣖ᵴേࠍ⸃ᨆ
ߔࠆ‫⎇ߚߞ޿ߣޔ‬ⓥࠍዷ㐿ߒߡ޿ࠆ‫౒ᧄޕ‬ห೑↪⎇ⓥળߪ‫⎇↹⸘ߥ߁ࠃߩߎޔ‬ⓥߩ⎇ⓥࠣ࡞࡯ࡊࠍਛᔃߦߒߡ‫ⴕޔ‬
േ․ᕈࠍ᳿ቯߔࠆࠥࡁࡓ‫߇ࡓࡁࠥޔ‬೙ᓮߔࠆ⣖ᯏ⢻‫ޔ‬⣖ᯏ⢻߇ⷙቯߔࠆⴕേ․ᕈߣ޿߁↢‛ቇ⊛࠻࡜ࠗࠕࡦࠣ࡞ߩ
ታ૕ࠍ᣿ࠄ߆ߦߔࠆߎߣࠍ⋡⊛ߣߒߡ㐿௅ߐࠇߚ‫⎇ޕ‬ⓥᚲౝᄖ߆ࠄ⚂ 50 ฬߩෳട⠪ࠍᓧߡ‫ޔ‬ᵴ⊒ߥᖱႎ੤឵‫ޔ‬ᗧ
⷗੤឵‫߮ࠃ߅ޔ‬୘‫⎇ߩޘ‬ⓥ⸘↹߿⎇ⓥᚑᨐߦ㑐ߔࠆ⼏⺰ࠍⴕߞߚ‫ޕ‬
㧨ࡊࡠࠣ࡜ࡓ㧪
3 ᦬ 2 ᣣ㧔㊄㧕
13:30㨪13:45 㜞↰᣽ᒾ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕᜿ᜦ
13:45㨪14:15 ᐔ੗໪ਭ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕ᣧ⠧∝ࡕ࠺࡞࠾ࡎࡦࠩ࡞ߩ⚦⢩ಽⵚ㧔❫⛽⧘⚦⢩㧕
14:15㨪14:45 ࠕ࡟ࠢࠪ࠴࡮ࡉ࡜ࡦࠦ㧔ฬฎደᄢቇᄢቇ㒮කቇ♽⎇ⓥ⑼㧕Copy number variation in schizophrenia in the
Japanese population
14:45㨪15:15 ㇹ ᐽᐢ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕ࠨ࡞ⴕേࠥࡁࡓᄙဳߩᬌ⚝ߣ⸃ᨆޯోࠥࡁࡓ࡮ࠛࠠ࠰࡯ࡓ࡮
࠻࡜ࡦࠬࠢ࡝ࡊ࠻࡯ࡓ⸃ᨆߦࠃࠆᄙဳᖱႎᢛ஻ޯ
15:15㨪15:45 ᴡ᧛ᱜੑ 㧔᧲੩ᄢቇᄢቇ㒮 ᣂ㗔ၞഃᚑ⑼ቇ⎇ⓥ⑼㧕㔤㐳㘃⥄ὼ㓸࿅ߦኻߔࠆᗵⷡ㑐ㅪㆮવሶߩᄙ
᭽ᕈߣㆡᔕㅴൻߩᬌ⸽
15:45㨪16:00
— ࡉ࡟ࠗࠢ —
16:00㨪16:30 ᄢ⍹㜞↢ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕ࡑࠞࠢߩ⣖ᯏ⢻߳ߩ᜸ᚢ㧦ㆮવሶ⊒⃻ߩട㦂ᄌൻߣⴕേ࠹ࠬ࠻
16:30㨪17:00 ਛ᧛స᮸ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕ࡑ࡯ࡕ࠮࠶࠻ߩ␠ળⴕേ
17:00㨪17:30 ጟᧄቬ⵨ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕㔤㐳⎇ߩࡑࠞࠢ㘃ߦ⷗ࠄࠇࠆᄌ⇣࡮ⴕേ⇣Ᏹߣࡕ࠺࡞♽⛔ߩ
૞⵾
17:30㨪18:00 ੹੗໪㓶 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕㔤㐳㘃⧰๧ฃኈ૕ߩಽሶ↢‛ቇ⊛⸃ᨆ
3 ᦬ 3 ᣣ㧔࿯㧕
9:30㨪10:00 የፒ♿ᄦ 㧔ฬฎደᄢቇᄢቇ㒮 කቇ♽⎇ⓥ⑼㧕♖␹⑼⥃ᐥߩ࠾࡯࠭ߦᔕ߃ࠆ⎇ⓥᚑᨐ㧦㔤㐳㘃ࠍ↪
޿ߚၮ␆⎇ⓥߣ♖␹⑼⥃ᐥ⎇ⓥߩㅪ៤ߦࠃࠆ The Valley of Death ߩస᦯ࠍ⋡ᜰߒߡ
10:00㨪10:30 ᯅᧄ੫ᄥ 㧔ᄢ㒋ᄢቇᄢቇ㒮 ㅪวዊఽ⊒㆐ቇ⎇ⓥ⑼㧕ಽሶ࡮⣖ᯏ⢻࡮♖␹∔ᖚࠍ⚿߱ᣂߒ޿ࠕࡊࡠ
࡯࠴㧙ࡅ࠻⣖⴫⃻ဳࠦࡦ࠰࡯ࠪࠕࡓ
10:30㨪11:00 ጤ↰ખ↢ 㧔⮮↰଻ஜⴡ↢ᄢቇ කቇㇱ㧕♖␹∔ᖚ⎇ⓥߦ߅ߌࠆࠥࡁࡓක⑼ቇߩน⢻ᕈ
11:00㨪11:15
— ࡉ࡟ࠗࠢ —
11:15㨪11:45 ᵏ⟜㓷⊓ 㧔᧲੩ක⑼ᱤ⑼ᄢቇᄢቇ㒮 කᱤቇ✚ว⎇ⓥ⑼㧕㜞㦂⠪↪⹺⍮ᯏ⢻ᬌᩏߣ⣖ᵴേߩ㑐ଥ
11:45㨪12:15 ૒⮮ብ㆏ 㧔ᄢ㒋ᄢቇᄢቇ㒮 කቇ♽⎇ⓥ⑼㧕ࡀࠦೋᦼⷞⷡ♽ߦ߅ߌࠆฃኈ㊁๟࿐ᛥ೙ߣᣇ૏ㆬᛯᕈ
12:15㨪12:45 ධㇱ ◊ 㧔↢ℂቇ⎇ⓥᚲ㧕␹⚻ᵴേࠍ⸥㍳ߔࠆߎߣߦࠃࠅ‫ޔ‬ᄢ⣖ၮᐩᩭ∔ᖚߩ∛ᘒߦㄼࠆ
12:45㨪13:30
— ᤤ㘩 —
13:30㨪14:00 ᤊ ⧷ม 㧔᧲੩ㇺකቇ✚ว⎇ⓥᚲ㧕ᗵⷡᖱႎߦၮߠߊⴕേ೙ᓮߦ߅ߌࠆ೨㗡ㅪว㊁ߩᓎഀ
14:00㨪14:30 ᷡ᳓⾆⟤ሶ㧔᧲੩ᄢቇᄢቇ㒮 ℂቇ♽⎇ⓥ⑼㧕㐳ᦼ⸥ᙘᒻᚑߣ᭎ᣣ࡝࠭ࡓߩᯏ⢻ㅪ㑐ߦ߅ߌࠆ SCOP
ߩᓎഀ
14:30㨪15:00 ዊᨋ๺ੱ 㧔⑔ፉ⋵┙ක⑼ᄢቇ කቇㇱ㧕⣖ౝㆮવሶዉ౉ᛛⴚࠍ೑↪ߒߚⴕേߣ⣖ᯏ⢻ߩ⎇ⓥ
㧔ᢥ⽿㧦㜞↰᣽ᒾ㧕
㧙㧙
ࠕࠫࠕߩ㔤㐳㘃ߩ଻ోߣ␠ળ↢ᘒ⎇ⓥߦ㑐ߔࠆㄭᐕߩᣂߚߥዷ㐿
ᣣᤨ㧦2012 ᐕ 3 ᦬ 10 ᣣ㧔࿯㧕࡮11 ᣣ㧔ᣣ㧕
႐ᚲ㧦ᗲ⍮⋵›ጊᏒቭᨋ ੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲᄢળ⼏ቶ㧔ෳടੱᢙ㧦⚂ 50 ੱ㧕
਎⹤ੱ㧦ฎᏒ೰ผ㧘MA Huffman㧘ඨ⼱๋㇢㧘ᯅᧄජ⛗㧘ㄞᄢ๺
⃻࿷‫ࠍ࡞ࠩࡦࡎ࠾ޔ‬฽߻ᣣᧄߩ㊁↢₞㘃ߩ୘૕ᢙ▤ℂߪᄢ߈ߥᦛ߇ࠅⷺࠍㄫ߃ߡ޿ࠆ‫ޕ‬ฦ࿾ߢⵍኂ໧㗴߇᜛ᄢߒ‫ޔ‬
ߎࠇ߹ߢߩ᡽╷߇⷗⋥ߒࠍㄼࠄࠇߡ޿ࠆ߆ࠄߢ޽ࠆ‫࡞ࠩࡦࡎ࠾ޕ‬એᄖߩࠕࠫࠕߩ㔤㐳㘃ฦ⒳߽‫↢ޔ‬ᕷ࿾ߩ⎕უ߿ੱ
ߣߩゟ㄂ߥߤ‫ߥ߹ߑ߹ߐޔ‬໧㗴ࠍᛴ߃ߡ޿ࠆ‫⎇ᧄޕ‬ⓥળߢߪ‫ࠍ࡞ࠩࡦࡎ࠾ޔ‬฽߻ࠕࠫࠕߩ㔤㐳㘃ߦߟ޿ߡ‫ߎߎޔ‬
10 ᐕ⒟ᐲߩ↢ᘒቇ⊛⎇ⓥߩᣂߚߥዷ㐿ࠍ✚᜝ߒ‫⎇ޔ‬ⓥ㕙‫ో଻ޔ‬㕙ߢߩᰴ߳ߩዷᦸࠍᮨ⚝ߒߚ‫ޕ‬
㧨ࡊࡠࠣ࡜ࡓ㧪
3 ᦬ 10 ᣣ
13:00㨪13:05 ⿰ᣦ⺑᣿㧦ฎᏒ೰ผ
㧨ࠕࠫࠕߩ㔤㐳㘃㧔1㧕㧪 ᐳ㐳㧦ᷰㆻ㇌ᄦ
13:05㨪13:40 Ꮉᧄ ⧐ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕ධࠕࠫࠕߩ㔤㐳㘃ߩ♽⛔࿾ℂ⎇ⓥ
13:40㨪14:15 ự↰ ⓛ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕᧲ධࠕࠫࠕߩ㔤㐳㘃‫ߩࠢࠞࡑߦ․ޔ‬ᄙ᭽ᕈ⎇ⓥ
14:15㨪14:50 ⷫᎹජ⚓ሶ㧔᧲ർᄢቇᄢቇ㒮 ㄘቇ⎇ⓥ⑼㧕᧲ධࠕࠫࠕߦ߅ߌࠆ㊁↢࠹࠽ࠟࠩ࡞ߩ㖸ჿࠦࡒࡘ࠾ࠤ
࡯࡚ࠪࡦ⎇ⓥ
㧨ࠕࠫࠕߩ㔤㐳㘃㧔2㧕㧪 ᐳ㐳㧦Ꮉᧄ⧐
15:00㨪15:35 ඨ⼱๋㇢ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕ࡑ࡟࡯ࠪࠕ࡮࠳࠽ࡓࡃ࡟࡯ߩ࡟࠶࠼࡝࡯ࡈࡕࡦࠠ࡯ߩ㘩ᕈ
ߣࡈࠜ࡯࡞ࡃ࠶ࠢ㘩‛
15:35㨪16:10 ਃ⼱㓷⚐ 㧔౓ᐶ⋵┙ᄢቇ ⥄ὼ࡮ⅣႺ⑼ቇ⎇ⓥᚲ㧕ੑᰴᬀ↢ߣ❫⛽㘩ᕈ㔤㐳㘃ߩ㘩ᕈ㧦ᶏᵗߦ㕙
ߔࠆቅ┙ߒߚࡂࡆ࠲࡯࠻ߩᜬߟᗧ๧
16:20㨪16:55 ᧻᧛⑲৻ 㧔ጘ㒂ᄢቇ ᔕ↪↢‛⑼ቇㇱ㧕ㅴൻ↢‛ቇ߆ࠄߺߚࠬ࡜࠙ࠚࠪࡑࠞࠢ
16:55㨪17:30 ਣᯅ⃨᮸ 㧔ᱞ⬿ᄢቇ ੱᢥቇㇱ㧕࠲ࠗ࡮ࠞࠝࠢ࡜ࡊ࠶ࠢߩࡌ࠾ࠟࠝࠩ࡞ߩ␠ળߣ↢ᘒ
3 ᦬ 11 ᣣ
㧨࠾ࡎࡦࠩ࡞㧔1㧕㧪 ᐳ㐳㧦ㄞᄢ๺
9:30㨪10:05 ᳯᚑᐢ᢯ 㧔ቝㇺችᄢቇ ㄘቇㇱ㧕ੱญᷫዋ␠ળߦ߅ߌࠆ࠾ࡎࡦࠩ࡞୘૕⟲▤ℂߩ⃻⁁ߣ⺖㗴
10:05㨪10:40 ቝ㊁ჽᤐ 㧔ችၔ ㊁↢േ‛଻⼔▤ℂ࠮ࡦ࠲࡯㧕᧲ർ࿾ᣇߦ߅ߌࠆ࠾ࡎࡦࠩ࡞ߩ㐳ᦼ⊛ߥ଻⼔▤ℂ
ࠍ⋡ᜰߒߡ
10:50㨪11:25 ᧻ጟผᦶ 㧔ਅർඨፉߩࠨ࡞⺞ᩏળ㧕ਅർඨፉߩࠨ࡞ 㧙ㆊ෰࡮⃻࿷࡮ᧂ᧪㧙
11:25㨪12:00 ⿒ᐳਭ᣿ 㧔ንጊ⋵┙౎የ㜞╬ቇᩞ㧕࠾ࡎࡦࠩ࡞ߩㆮવ⊛ᄙ᭽ᕈ଻ోߣ଻⼔▤ℂ⸘↹
㧨࠾ࡎࡦࠩ࡞㧔2㧕㧪 ᐳ㐳㧦ඨ⼱๋㇢
13:00㨪13:35 ቶጊᵏਯ 㧔౓ᐶ⋵┙ᄢቇ ⥄ὼ࡮ⅣႺ⑼ቇ⎇ⓥᚲ㧕࠾ࡎࡦࠩ࡞ߩ଻ోߣ▤ℂ㧙౓ᐶ⋵ߦ߅ߌࠆข
ࠅ⚵ߺߦߟ޿ߡ
13:35㨪14:10 ᄢ੗ ᔀ 㧔᫪ᨋ✚ว⎇ⓥᚲ ↢‛ᄙ᭽ᕈ⎇ⓥࠣ࡞࡯ࡊ㧕ߤ߁⹏ଔߔࠆ࠾ࡎࡦࠩ࡞୘૕⟲ߩ⛘Ṍෂ
㒾ᐲ㧙࡟࠶࠼࠺࡯࠲ߩ⃻⁁ߣ੹ᓟ
14:10㨪14:55 ✚ว⸛⺰
14:55㨪15:00 㐽ળ᜿ᜦ
㧔ᢥ⽿㧦ฎᏒ೰ผ㧕
╙ 40 ࿁ࡎࡒ࠾࠯࡯࡚ࠪࡦ⎇ⓥળ‫ߩࡦࡒࡄ࡯࠼ޟ‬ᓎഀ㧦ㆇേᯏ⢻߆ࠄ㜞ᰴᯏ⢻߳‫ޠ‬
ᣣᤨ㧦2012 ᐕ 3 ᦬ 15 ᣣ㧔ᧁ㧕࡮16 ᣣ㧔㊄㧕
႐ᚲ㧦ᗲ⍮⋵›ጊᏒቭᨋ ੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲᄢળ⼏ቶ㧔ෳടੱᢙ㧦⚂ 30 ੱ㧕
਎⹤ੱ㧦᧻ᧄᱜᐘ㧘㜞↰᣽ᒾ㧘ਛ᧛స᮸
㧨ࡊࡠࠣ࡜ࡓ㧪
3 ᦬ 15 ᣣ㧔ᧁ㧕
࡮ᯌ ศኼ 㧔↢ℂቇ⎇ⓥᚲ㧕‫ޟ‬The ventral pallidum encodes expected reward value of future action‫ޠ‬
࡮દ⮮ ⌀ 㧔ᴒ✽⑼ቇᛛⴚᄢቇ㒮ᄢቇ㧕‫ޟ‬ႎ㈽ߦၮߠߊᗧᕁ᳿ቯߩࠕ࡞ࠧ࡝࠭ࡓߣ✢⁁૕ߩᖱႎ⴫⃻‫ޠ‬
࡮੗਄⻞৻ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕‫ޟ‬㔤㐳㘃ߦ߅ߌࠆ㤥⾰࠼࡯ࡄࡒࡦ࠾ࡘ࡯ࡠࡦㆬᛯ⊛ߥㆮવሶዉ౉‫ޠ‬
࡮⇸↰⾆ବ 㧔੩ㇺᄢቇ කቇ⎇ⓥ⑼㧕‫ޟ‬ႎ㈽࡮ᔊㆱⴕേߦ߅ߌࠆᄢ⣖ၮᐩᩭ␹⚻࿁〝ᯏ᭴‫ޠ‬
࡮ᧁ᧛ᐘᄥ㇢㧔ᄢ㒋ᄢቇ ℂቇ⎇ⓥ⑼㧕‫ߩ⯻✢ޟ‬൬޿ᔊㆱቇ⠌ࠍ೙ᓮߔࠆ࠼࡯ࡄࡒࡦߩ૞↪ࡔࠞ࠾࠭ࡓ‫ޠ‬
3 ᦬ 16 ᣣ㧔㊄㧕
࡮᧻ᧄᱜᐘ 㧔੩ㇺᄢቇ 㔤㐳㘃⎇ⓥᚲ㧕‫ޟ‬ㇱ૏․⇣⊛ߥਛ⣖࠼࡯ࡄࡒࡦ⚦⢩ߩᵴേߣߘߩᯏ⢻⊛ᓎഀ‫ޠ‬
࡮᭑ᧄ৻♿ 㧔₹Ꮉᄢቇ࡮⣖⑼ቇ⎇ⓥᚲ㧕‫࡞ࠨޟ‬ਛ⣖࠼࡯ࡄࡒࡦ⚦⢩ߦࠃࠆ‫ᤨߚߞߥ⇣ޔ‬㑆ࠬࠤ࡯࡞ߢߩ዁᧪ႎ㈽
㧙㧙
ߩଔ୯ߣ⺋Ꮕ⴫⃻‫ޠ‬
࡮᧻↰๺㇢ 㧔ṑ⾐ක⑼ᄢቇ࡮කቇㇱ㧕‫ޟ‬ਛ⣖࠼࡯ࡄࡒࡦ⚦⢩ߩᒻᘒߣ⠨ኤ㧦ࡄ࡯ࠠࡦ࠰ࡦ∛ߩ⊒∝ߦ㑐ਈߔࠆ␹
⚻ၮ⋚‫ޠ‬
࡮ධᧄᢘผ 㧔᡼኿✢කቇ✚ว⎇ⓥᚲ㧕‫ࡦ࡚ࠪ࡯ࡌ࠴ࡕߣࡦࡒࡄ࡯࠼ޟ‬㧦േᯏઃߌ೙ᓮߣߘߩ㓚ኂ߆ࠄ⷗߃ߡ߈ߚ
ߎߣ‫ޠ‬
ߘߩᰳਲ߇᭽‫∔⚻␹࡮␹♖ߥޘ‬ᖚߩේ࿃ߣߥࠆߎߣ߆ࠄ߽ࠊ߆ࠆࠃ߁ߦ‫ߩ࠻ࡅߪࡦࡒࡄ࡯࠼ޔ‬㜞ᰴ⣖ᯏ⢻ࠍታ⃻
ߔࠆߚ߼ߦ㊀ⷐߥ␹⚻વ㆐‛⾰ߢ޽ࠆ‫৻ޕ‬ᣇ‫ޔ‬᣸⯻߿㝼㘃ߥߤߩ␹⚻♽ߦ߽࠼࡯ࡄࡒࡦߪሽ࿷ߒ‫ޔ‬៨㘩߿ㅏㆱⴕേ
ߥߤ‫↢ޔ‬๮⛽ᜬߦߣߞߡᔅ㗇ߥᯏ⢻߳ߩ㑐ࠊࠅ߇ႎ๔ߐࠇߡ߅ࠅ‫ߩߘޔ‬ᯏ⢻ࠍ↹৻⊛ߦᝒ߃ࠆߎߣߪ㔍ߒ޿‫ޔߦ․ޕ‬
࠼࡯ࡄࡒࡦ↥↢⚦⢩ߩ␹⚻ㅪ⛊ߪ‫⊒߽ᦨߢ࠻ࡅޔ‬㆐ߒߚ೨㗡ㅪว㊁ߣㅴൻ⊛ߦ଻ሽߐࠇߚᄢ⣖ၮᐩᩭߦ߅޿ߡᒝ޿
ߎߣ߆ࠄ‫ߩߘޔ‬ᓎഀߩㅴൻ⊛ᄌㆫࠍ⍮ࠆߎߣߪ‫⹺ޔ‬⍮߿ᖱേߥߤ‫ޔ‬ᚒ‫ߩޘ‬㜞ᰴ⣖ᯏ⢻߇ㅴൻߩㆊ⒟ߢߤߩࠃ߁ߦ⊒
㆐ߒߡ߈ߚ߆ࠍℂ⸃ߔࠆ਄ߢ㊀ⷐߢ޽ࠆ‫੹ޕ‬࿁ߩࡎࡒ࠾࠯࡯࡚ࠪࡦ⎇ⓥળߢߪ‫ߞߍޔ‬ᱤ㘃߿㔤㐳㘃ߛߌߢߪߥߊ‫ޔ‬
✢⯻ࠍ↪޿ߡታ㛎ࠍ߅ߎߥߞߡ޿ࠆ⎇ⓥ⠪߽ෳടߒ‫ޔ‬ቇ⠌߿േᯏઃߌ‫⹺ޔ‬⍮ᯏ⢻╬߳ߩ࠼࡯ࡄࡒࡦߩᓎഀߦߟ޿ߡ
߅੕޿ߩ⍮⷗ࠍᜬߜࠃࠅ‫ߩߘޔ‬ᯏ⢻ߩᄙ᭽ᕈ߿ㅴൻ⊛ᄌㆫߦߟ޿ߡ⸛⺰ࠍ߅ߎߥߞߚ‫ޕ‬
㧔ᢥ⽿㧦᧻ᧄᱜᐘ㧕
‫߇࡞ࠨޟ‬ੑ⿷ߢᱠߊߣ߈‫ޠ‬
㐿௅ᣣ㧦2012 ᐕ 3 ᦬ 17 ᣣ㧔࿯㧕࡮18 ᣣ㧔ᣣ㧕
႐ᚲ㧦੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲ ᄢળ⼏ቶ㧔ෳടੱᢙ㧦⚂ 40 ੱ㧕
਎⹤ੱ㧦ᐔፒ㍈⍫㧘ự↰ⓛ㧘ฎᏒ೰ผ㧘ች࿾㊀ᒄ㧘Ძ೑ବ㓶㧘᫪ᄢᔒ㧔ጊญᄢ㧕
‫┙⋥ޟ‬ੑ⿷ᱠⴕߩ⿠Ḯߣㅴൻ‫ࠍޠ‬⠨߃ࠆᤨ‫ޔ‬໧㗴ߪᄢ߈ߊ 3 ߟߦಽ߆ࠇࠆ‫ޕ‬ੑ⿷ᱠⴕࠍน⢻ߦߒߚり૕᭴ㅧ߿⣖
ᯏ⢻ߪߤߩࠃ߁ߥ߽ߩ߆‫ޔ‬ੑ⿷ᕈߪ૗ࠍ߈ߞ߆ߌߦߤ߁޿ߞߚ⁁ᴫߢ₪ᓧߐࠇߚߩ߆‫ޔ‬ੑ⿷ᕈ₪ᓧᓟߩᱠⴕߪߤߩ
ࠃ߁ߦᵞ✵ߐࠇߡ޿ߞߚߩ߆‫߁޿ߣ߆ߩߚߞߎ⿠ߟ޿߇࠻ࡦࡌࠗߩࠄࠇߎޔߦࠄߐޕ‬໧޿߇ 4 ߟ⋡ߣߒߡടࠊࠆ‫ޕ‬
ߎ߁ߒߚ⇼໧ߦኻߒߡ‫ޔࠇߙࠇߘޔ‬ᯏ⢻ᒻᘒቇߣ␹⚻⑼ቇ‫ޔ‬㔤㐳㘃␠ળቇ࡮↢ᘒቇ‫ޔ‬ㅴൻࡠࡏ࠹ࠖࠢࠬ‫ޔ‬ฎੱ㘃ቇ
߆ࠄߩࠕࡊࡠ࡯࠴߇ߥߐࠇ‫ޔ‬ᄙߊߩᚑᨐࠍ޽ߍߡ߈ߚ‫ోޔߒ߆ߒޕ‬૕ࠍ⷗ᷰߔ⺰⼏‫ޔ‬੕޿ߩᚑᨐࠍෳᾖ࡮⵬ᒝߒ޽
߁⠨ኤߪᔅߕߒ߽චಽߦߥߐࠇߡ޿ࠆߣߪ⸒߃ߥ޿‫ޕ‬ੑ⿷ᱠⴕߣ޿߁౒ㅢ࠹࡯ࡑߩਅߦ⇣ߥࠆ┙႐ߢ⎇ⓥࠍⴕߞߡ
޿ࠆᄙ᭽ߥಽ㊁ߩ⎇ⓥ⠪߇㓸߹ࠅ‫ᦨޔ‬ᣂߩᚑᨐߦၮߠ޿ߡᗧ⷗ࠍ੤឵ߔࠆߎߣ߇ߎߩ⎇ⓥળߩ⋡⊛ߢ޽ߞߚ‫ޕ‬
ೋᣣߪ߹ߕ‫ޔ‬㔤㐳㘃⎇ⓥᚲߢ 40 ᐕ೨ߦᆎ߹ߞߚ㔤㐳㘃ࡠࠦࡕ࡯࡚ࠪࡦ⎇ⓥߩᵹࠇࠍ᳹߻⻠Ṷ 3 㗴߆ࠄᆎ߹ࠅ‫ޔ‬
⛯޿ߡㄭᐕ⋡ⷡߒ޿ᚑᨐࠍ޽ߍߡ޿ࠆ␹⚻⑼ቇ⊛ࠕࡊࡠ࡯࠴ߦࠃࠆᦨᣂߩᚑᨐߩ⚫੺߇ߥߐࠇߚ‫ޕ‬ᙣⷫળߢߪ‫⾰ޔ‬
⇼ᔕ╵߿⸛⺰ߩᤨ㑆ᨒߦߪ෼߹ࠅ߈ࠇߥ߆ߞߚ⾰໧߿⼏⺰߇ᵴ⊒ߦߥߐࠇߚ‫ޕ‬ੑᣣ⋡ߦߪ‫ⷰ࠼࡞࡯ࠖࡈޔ‬ኤߦࠃࠆ
ᄢဳ㘃ੱ₎ 3 ⒳ߩࡐ࡚ࠫࠪ࠽࡞ⴕേߦߟ޿ߡߩ⾆㊀ߥ⍮⷗߿‫ޔ‬Ꮏቇ⊛ᚻᴺߦࠃࠆᱠⴕ࡮ᆫ൓೙ᓮߩࠪࡒࡘ࡟࡯࡚ࠪ
ࡦ⎇ⓥ߇⚫੺ߐࠇ‫ߦࠄߐޔ‬ൻ⍹⎇ⓥࠍㅢߒߡో૕ࠍ⷗ᷰߔⷞὐ߇ឭଏߐࠇߚ‫ޕ‬ᤤ㘩ᓟߦⴕࠊࠇߚ✚ว⸛⺰ߦ߅޿ߡ
߽‫ޔ‬ዧ߈ࠆߎߣߥߊ⚂ 40 ฬߩෳട⠪ߦࠃࠆᵴ⊒ߥ⼏⺰߇ⴕࠊࠇߚ‫ࠎߛ߰ޕ‬੕޿ߩ⎇ⓥߦធߔࠆߎߣߩዋߥ޿⇣ಽ
㊁ߩ⎇ⓥ⠪߇౒ㅢߩ࠹࡯ࡑߢ⸛⺰ߔࠆ⾆㊀ߥᯏળߣߥߞߚ‫ߚߒ߁ߎޕ‬ᯏળࠍ৻ᐲ߈ࠅߢ⚳ࠊࠄߖࠆߎߣߥߊ‫੹ޔ‬ᓟ
߽ᗧ⷗ߩ੤឵ࠍ⛮⛯ߒ‫ޔ‬ᣂߚߥࡄ࡜࠳ࠗࡓࠍ᭴▽ߒߡ޿߈ߚ޿‫ޕ‬
㧨ࡊࡠࠣ࡜ࡓ㧪
3 ᦬ 17 ᣣ㧔࿯㧕
13:00㨪13:10 ⿰ᣦ⺑᣿ ᐔፒ㍈⍫
ੑ⿷ᱠⴕߩၮ⋚㨪ᯏ⢻ᒻᘒቇߩ┙႐߆ࠄ㧔ᐳ㐳㧦Ძ೑ବ㓶㧕
13:10㨪13:40 ᾢୖඳ㓶㧔ᄢ㒋ᄢ㧕૕ᐙㆇേߩ᝝ᝒᣇᴺߣߘߩᯏ⢻⊛ᗧ⟵
13:40㨪14:20 ᧁ᧛ ⾥㧔᧲੩ᄢ㧕ࠨ࡞߆ࠄࡅ࠻ߩੑ⿷ᱠⴕࠍ⠨߃ࠆ
14:20㨪14:50 ᐔፒ㍈⍫㧔੩ㇺᄢ㧕⿷࿶ߣ⿷ᒻᘒ߆ࠄߺߚࠨ࡞ߩੑ⿷ᱠⴕ
ੑ⿷ᱠⴕߩၮ⋚㨪␹⚻⑼ቇߩ┙႐߆ࠄ㧔ᐳ㐳㧦ች࿾㊀ᒄ㧕
15:00㨪15:40 㜞⨲ᧁ⮍㧔ᣩᎹකᄢ㧕ᱠⴕߩ␹⚻↢ℂቇ
15:40㨪16:20 ਛ㒲సᏆ㧔ㄭ⇰ᄢ㧕࠻࡟࠶࠼ࡒ࡞ᱠⴕߦ㑐ㅪߒߚࠨ࡞৻ᰴㆇേ㊁ߩ␹⚻⚦⢩ᵴേ
16:20㨪17:00 ᫪ ᄢᔒ㧔ጊญᄢ㧕࠾ࡎࡦࠩ࡞ߩੑ⿷ᱠⴕቇ⠌
⸛⺰㧔ᐳ㐳㧦ᐔፒ㍈⍫㧕
17:15㨪 ࠦࡔࡦ࠹࡯࠲࡯㧦⍹↰⧷ኪ㧔⡛ᴰᄢ㧕‫ޔ‬ᣣ᥵ᵏ↵㧔ᄢ㒋ᄢ㧕
ᙣⷫળ㧔㔤㐳㘃⎇ⓥᚲᄙ⋡⊛ࡎ࡯࡞㧕
18:00㨪20:00
3 ᦬ 18 ᣣ㧔ᣣ㧕
૗᡿┙ߜ਄߇ࠆߩ߆‫ߦ߁ࠃߩߤޔ‬ᱠߊߩ߆㨪↢ᘒቇߩ┙႐߆ࠄ㧔ᐳ㐳㧦ᐔፒ㍈⍫㧕
9:00㨪 9:40 ฎᏒ೰ผ㧔੩ㇺᄢ㧕࠴ࡦࡄࡦࠫ࡯ߣࡏࡁࡏߩࡠࠦࡕ࡯࡚ࠪࡦ㧦᮸਄⒖േߣ࿾਄⒖േߩ⒳㑆Ყセ
9:40㨪10:20 ਭ਎Ớሶ㧔੩ㇺᄢ㧕ࠝ࡜ࡦ࠙࡯࠲ࡦߩࡠࠦࡕ࡯࡚ࠪࡦߩᄙ᭽ᕈߣ‫ޔ‬ᄙ᭽ᕈࠍ↢ߺ಴ߔⷐ࿃
㧙㧙
ੑ⿷ᱠⴕߩ઒ᗐㅴൻ㨪ㅴൻࡠࡏ࠹ࠖࠢࠬߩ┙႐߆ࠄ㧔ᐳ㐳㧦᫪ᄢᔒ㧕
10:30㨪11:00 ㊁᧛ᵏિ㧔ᄢ㒋ᄢ㧕ᨵエ߆ߟࡠࡃࠬ࠻ߥ⋥┙ᆫ൓ߩ␹⚻೙ᓮᚢ⇛
11:00㨪11:40 ⩆ේ⋥㆏㧔ᘮᔕᄢ㧕࠾ࡎࡦࠩ࡞ੑ⿷ᱠⴕㆇേߩ㗅േജቇ࡮ㅒേജቇࠪࡒࡘ࡟࡯࡚ࠪࡦ
ൻ⍹ੱ㘃ቇߩ┙႐߆ࠄ㧔ᐳ㐳㧦ฎᏒ೰ผ㧕
11:40㨪12:20 ਛോ⌀ੱ㧔੩ㇺᄢ㧕ೋᦼੱ㘃ߩੑ⿷ᱠⴕ
✚ว⸛⺰㧔ᐳ㐳㧦ự↰ⓛ㧕
13:20㨪 ࠦࡔࡦ࠹࡯࠲࡯㧦ጟ↰቞ᒾ㧔Ꮲ੩ᐔᚑᄢ㧕‫ޔ‬ട⾐⼱⟤ᐘ㧔੩ㇺᄢ㧕
㧔ᢥ⽿㧦ᐔፒ㍈⍫㧕
‫ ╙ޟ‬7 ࿁›ጊᲧセ␠ળ⹺⍮ࠪࡦࡐࠫ࠙ࡓ‫ޠ‬
ᣣᤨ㧦2012 ᐕ 3 ᦬ 24 ᣣ㧔࿯㧕࡮25 ᣣ㧔ᣣ㧕
႐ᚲ㧦੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲᄢળ⼏ቶ㧔ෳടੱᢙ㧦⚂ 50 ੱ㧕
⎇ⓥળ਎⹤ੱ㧦෹᳗㓷Ꮖ㧘ᨋ⟤㉿㧘દ᧛⍮ሶ㧘⿷┙ᐞ⏴㧘᧼ୖᤘੑ㧔ᢥቇ⎇ⓥ⑼㧕㧘᣿๺᡽ሶ㧔ᢎ⢒ቇ⎇ⓥ⑼㧕
㧨ࡊࡠࠣ࡜ࡓ㧪
3 ᦬ 24 ᣣ㧔࿯㧕
13:00㨪18:10
࠮࠶࡚ࠪࡦ 1
᫪ ⵨♿ 㧔ᄢ㒋ᄢቇ࡮Ꮏ㧕‫߇ⷡ⸅ޟ‬ዉߊೋᦼ⢝ఽⴕേ⊒㆐᭴ᚑ⺰‫ޠ‬
⣁ဈፏᐔ 㧔ℂ⎇࡮BSI࡮ㆡᔕ⍮ᕈ㧕‫ޟ‬ઍᦧ⃻ታࠪࠬ࠹ࡓߣߘߩ⹺⍮ᔃℂታ㛎ⵝ⟎ߣߒߡߩᔕ↪‫ޠ‬
ዊᎹஜੑ 㧔㧭㨀㧾⹺⍮ᯏ᭴⎇㧕‫ⴕޟ‬ὑ⹺⼂ߦ㑐ࠊࠆࡒ࡜࡯ࠪࠬ࠹ࡓߩ␹⚻⴫⽎‫ޠ‬
ᆂየᱞᴦ 㧔਻Ꮊᄢ࡮Ꮏ࡮ቇᝄ PD㧕‫ޟ‬X with Vection‫ޠ‬
࠮࠶࡚ࠪࡦ 2
ᷰㄝഃᄥ
‐⼱ᥓ৻
ᓟ⮮๺ብ
෹᳗㓷Ꮖ
ᙣⷫળ
㧔੩ㇺᄢ࡮ᢥ㧕‫⋧ߩ࠻ࡂޟ‬ኻఝ૏ᕈ㧒࠾ࡎࡦࠩ࡞ߩ⛘ኻఝ૏ᕈ‫ޠ‬
㧔ජ⪲ᄢ࡮ᢥ㧕‫ߩࠬ࡯ࡌ࠻ࠢࠚࠫࡉࠝޟ‬ᵈᗧߩᲧセ⹺⍮⑼ቇ‫ޠ‬
㧔੩ㇺᄢ࡮↢๮⑼ቇ♽ࠠࡖ࡝ࠕࡄࠬᒻᚑ࡙࠾࠶࠻㧕‫࠻࡞࠲ࡘࠪࠥޟ‬Ყセᔃℂቇ㧦ഃ⊒ᕈ߳ߩ᜸ᚢ‫ޠ‬
㧔੩ㇺᄢ࡮㔤㐳⎇㧕‫ߚ⷗ࠄ߆ࠞ࡞ࠗޟ‬਎⇇‫ޠ‬
3 ᦬ 25 ᣣ㧔ᣣ㧕 9:00㨪12:40
࠮࠶࡚ࠪࡦ 3
⼾Ꮉ ⥶ 㧔ർᶏ㆏ᄢ࡮ᢥ㧕‫ޟ‬ญࠦࡒߩવ㆐ߪ㓸ว⍮ࠍ↢ߺ಴ߔ߆㧫‫ޠ‬
᧻రஜੑ 㧔₹Ꮉᄢ࡮⣖⎇㧕‫ޟ‬⣖ౝߩଔ୯⴫⃻ߣߘߩᄌኈ‫ޠ‬
࠮࠶࡚ࠪࡦ 4
ᐔ੗⌀ᵗ 㧔ᗲ⍮⋵ᔃり㓚ኂ⠪ࠦࡠ࠾࡯㧕‫ࡦ࡚ࠪ࡯ࡕ࡞ࠞࠫࡠࠝࠗࡃޟ‬⍮ⷡᯏ᭴ߩᯏ⢻⊛⸃೬㧦㓏ጀ⊛ಣℂ઒
⺑ߩᬌ⸽ߣ⊒㆐ߦࠃࠆᄌൻ‫ޠ‬
⍹ፉߎߩߺ㧔ᣧⒷ↰ᄢ࡮ੱ㑆⑼ቇ㧕
‫ߣఽ੃ޟ‬Უⷫߩߊߔߋࠅㆆ߮ߦ߅ߌࠆ⋧੕૞↪㧦
‫ޟ‬ᗧ࿑‫ߩޠ‬ℂ⸃ߣᢥ⣂ߩ౒
᦭‫ޠ‬
᧻↰૫ዏ 㧔JST-ERATO࡮ጟࡁ⼱ᖱേᖱႎࡊࡠࠫࠚࠢ࠻㧕‫੃ޟ‬ᐜఽᦼߦ߅ߌࠆੱ⷗⍮ࠅߩࡔࠞ࠾࠭ࡓ‫ޠ‬
᣿๺᡽ሶ 㧔੩ㇺᄢ࡮ᢎ⢒㧕‫ޟ‬๟↥ᦼ߆ࠄߩᔃߩ⊒㆐ߣߘߩ↢‛ቇ⊛ၮ⋚‫ޠ‬
‫›ޟ‬ጊᲧセ␠ળ⹺⍮ࠪࡦࡐࠫ࠙ࡓ‫੹߽ޠ‬࿁ߢ╙ 7 ࿁ࠍㄫ߃ߚ‫ ╙ޕ‬3 ࿁߹ߢߪ․ቯߩ࠹࡯ࡑࠍ⸳ቯߒߡ㐿௅ߒߡ߈
ߚ߇‫ ╙ޔ‬4 ࿁એ㒠ߪ‫ࠍࠢ࠶ࡇ࠻ޔ‬㒢ቯߔࠆߎߣߥߊ‫⹺ޔ‬⍮⑼ቇߦ㑐ߔࠆᄙ᭽ߥ⎇ⓥ㗔ၞߢߩᦨవ┵ߩ⎇ⓥߩ⃻⁁ߦ
ߟ޿ߡ‫ޔ‬ᄙߊߩ⎇ⓥ⠪߇ಽߌ㓒ߡߥߊ⼏⺰ߒว߁႐ࠍ᭴ᚑߔࠆߎߣࠍ⋡ᜰߒߡ߈ߚ‫ޕ‬੕޿ߦ㑐ㅪߔࠆߣߪ޿߃‫ޔ‬ታ
㛎߆ࠄⷰኤߦ೔ࠆᄙ⒳ᄙ᭽ߥᣇᴺ⺰߆ࠄ↢ߺ಴ߐࠇࠆᚑᨐࠍߘࠇߙࠇߩ┙႐߆ࠄ⺰ߓว߁႐ߣߒߡ‫ޔ‬ቇ⇇ߦ߅޿ߡ
߽޽ࠆ৻⒟ᐲߩ⹺⍮ࠍ₪ᓧߒߡ߈ߚߣ⥄⽶ߒߡ޿ࠆ‫੹ޕ‬࿁ߪ‫ߩ⥸⻉ޔ‬੐ᖱߢ‫ޔ‬ᓥ᧪ߩ 12 ᦬㐿௅߆ࠄ 3 ᦬ߩ㐿௅ߣ
ߥߞߚߚ߼‫ޔ‬ෳട⠪ߩᷫዋ߇ෂ߱߹ࠇߚ߇‫੍ޔ‬ᗐએ਄ߩෳട⠪ߦߏෳ㓸޿ߚߛߌߚ‫੹ޕ‬࿁߽ࡠࡏ࠶࠻߆ࠄࠗ࡞ࠞ߹
ߢᄙ᭽ߥኻ⽎ߦኻߔࠆߘࠇߙࠇߦ⥝๧ᷓ޿ᚑᨐ߇ႎ๔ߐࠇߚ‫੹ޔߚ߹ޕ‬࿁ߪ‫ޔ‬੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲᕁ⠨⸒⺆ಽ㊁
ߣߩ౒ห೑↪⎇ⓥߩᚑᨐߦߟ޿ߡ߽‫ޔ‬ᷰㄝ‫ޔ⼱‐ޔ‬ᓟ⮮ߩ 3 ฬ߆ࠄႎ๔߇޽ߞߚ‫੹ޕ‬ᓟߣ߽౒ห೑↪⎇ⓥߩᚑᨐࠍ
ߎߩ႐ߢႎ๔ߔࠆᯏળࠍჇ߿ߒߡ޿߈ߚ޿‫੹ޕ‬࿁ߪ‫ޔ‬Ḱ஻ߩㇺวߢ଀ᐕⴕߞߡ߈ߚࡐࠬ࠲࡯⊒⴫ࠍⴕࠊߥ߆ߞߚ‫ޕ‬
੹ᓟߪ߹ߚౣ㐿ߒߚ޿‫ޕ‬
㧔ᢥ⽿㧦෹᳗㓷Ꮖ㧕
㧙㧙
Ϋ㧚ㅌ⡯ߦ޽ߚߞߡ
␠ળ↢ᘒ⎇ⓥㇱ㐷↢ᘒ଻ోಽ㊁࡮ᷰㆻ㇌ᄦ
⑳ߪ 1972 ᐕߦ‫ޔ‬㔤㐳㘃⎇ⓥᚲߩᄢቇ㒮╙৻ᦼ↢ߣߒߡ›ጊߦ᧪߹ߒߚ‫ߣߞߕ߹߹ߩߘߡߒߘޕ‬㔤㐳
㘃⎇ⓥᚲߦ߅਎⹤ߦߥߞߡߒ߹޿߹ߒߚ‫ߩߘޕ‬㑆 1982 ᐕߦᐘፉ㊁ᄖⷰኤᣉ⸳ߩഥᚻߣߒߡណ↪ߐࠇ‫ޔ‬
ችፒ⋵ਠ㑆Ꮢߢ 13 ᐕࠍㆊߏߒ߹ߒߚ‫ޕ‬ᐘፉ㊁ᄖⷰኤᣉ⸳ߪ⑳ߩ⿞છᓟ 1 ᐕߢᡷ⚵ߐࠇ‫࡞ࠩࡦࡎ࠾ޔ‬㊁
ᄖⷰኤᣉ⸳ߣ޿߁࿖ౝ 5 ࠞᚲߩ⎇ⓥᨋ࡮ⷰኤࠬ࠹࡯࡚ࠪࡦࠍᜬߟᣉ⸳ߦߥࠅ߹ߒߚ‫࡞ࠩࡦࡎ࠾ߩߘޕ‬㊁
ᄖⷰኤᣉ⸳߇ 2008 ᐕߦ੩ㇺᄢቇ㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯ഃ⸳ߦߣ߽ߥߞߡᡷᑄߐࠇ‫␠ߪ⑳ޔ‬ળ↢ᘒ⎇ⓥ
ㇱ㐷↢ᘒ଻ోಽ㊁ߦ⒖ࠅ‫੹ޔ‬ᣣߦ⥋ߞߚࠊߌߢߔ‫ࠍ࡞ࠩࡦࡎ࠾ޔߪߩߚ߈ߡߞ߿߇⑳ޕ‬฽߻ࠕࠫࠕߩࠨ
࡞ߩ␠ળⴕേߣ↢ᘒ‫⎇ߩో଻ߡߒߘޔ‬ⓥߢߔ߇‫ޔ‬㐳ߊ㒝ዻᣉ⸳ߦᚲዻߒߡ޿ߚߦ߽߆߆ࠊࠄߕ‫ࠅߥ߆ޔ‬
⥄↱ߦᶏᄖ⺞ᩏߦ߽ขࠅ⚵߻ߎߣ߇ߢ߈߹ߒߚ‫࡞ࠩࡦࡎ࠾ߚ߹ޕ‬㊁ᄖⷰኤᣉ⸳ߣ޿߁ߩߪ‫৻ࠅࠃ⑳ޔ‬਎
ઍ೨ߩ㊁ᄖ⎇ⓥ⠪ߚߜߘࠇߙࠇߩᕁ޿߇ߟ߼ߎ߹ࠇߚ‫ޔ‬࿾ၞ␠ળߣߩኒធߥߟߥ߇ࠅߩ޽ࠆᣉ⸳ߛߞߚ
ߩߢ‫⎇ߩߤޔ‬ⓥᨋ࡮ⷰኤࠬ࠹࡯࡚ࠪࡦߦ߅޿ߡ߽ㅢᏱߢߪߢ߈ߘ߁ߥ޿߽ߥ޿ઃ߈ว޿ࠍߚߊߐࠎߐߖ
ߡ߽ࠄ޿߹ߒߚ‫ޕ‬
ߎ߁ߒߡᝄࠅ㄰ߞߡߺࠆߣ‫߇ߣߎߥࠈ޿ࠈ޿ޔ‬⣖ⵣߦᶋ߆ࠎߢ߈߹ߔ‫߇⑳ޕ‬㔤㐳㘃⎇ⓥᚲߦ᧪ߚ㗃ߣ
޿߁ߩߪ‫ޔ‬ᣣᧄߩ㔤㐳㘃ቇ߇᦭ฬߦߥߞߚ‫ߩߘޔ‬ၮ␆ࠍ▽޿ߚ⏋ቇߚߜ߇߹ߛᵴべߒߡ޿ߚᦨᓟߩ㗃ߦ
޽ߚࠅ߹ߔ‫ࠅ߅߈ߣޕ‬㐿௅ߐࠇࠆ⎇ⓥળߦߪ‫ߥ߹ߑ߹ߐޔ‬ಽ㊁ߩ⪺ฬߥ⎇ⓥ⠪߇಴౉ࠅߒߡ޿ߚߛߌߢ
ߥߊ‫ޔ‬࿷㊁ߩ⎇ⓥ⠪߽ᄙᢙෳടߒߡ߅ࠅ‫ޔ‬ᵴ⊒ߥ⼏⺰߇ߚߚ߆ࠊߐࠇߚ߽ߩߢߔ‫ߡߒߘޕ‬ᄢቇ⚗੎ߩߥ
ߏࠅ߇ᱷࠆਛߢ‫ޔ‬ᄢቇ߿⎇ⓥߩ␠ળ⊛ߥᓎഀߣ޿߁ߎߣߦߟ޿ߡ߽‫ߘޕߚߒ߹ࠅ޽߇⺰⼏ߥࠈ޿ࠈ޿ޔ‬
ߩ㗃ߪ‫ޔ‬㔤㐳㘃⎇ⓥᚲ߇ߢ߈ߚ߫߆ࠅߛߞߚߩߢ‫ޔ‬ᢎຬߩඨᢙ⒟ᐲߪ߶ࠎߩߜࠂߞߣᐕ㈩ߩఱ⾆ಽߣᕁ
߃ࠆ߶ߤߦ⧯ߊ‫⁛ޔ‬り⠪߽ᄙ߆ߞߚߩߢ‫ޔ‬᳇౗ߨߥߊ㈬ࠍߨߛࠅߥ߇ࠄ‫ߩ߽ߚߌ޿ߦߌ߆ߞ߰ࠍ⺰⼏ޔ‬
ߢߔ‫੹ޕ‬⠨߃ࠆߣ‫ޔ‬ᱜ⋥‫ᤨߥࠎ޽ޔ‬ઍ߽޽ߞߚߩߛߣߒ߆⸒޿ࠃ߁߇޽ࠅ߹ߖࠎ‫ޕ‬
ߘࠇ߆ࠄ 40 ᐕ‫ޔ‬㔤㐳㘃⎇ⓥᚲߪߕ޿߱ࠎߣᄌࠊࠅ߹ߒߚ‫⎇ޕ‬ⓥᚲߛߌߢߥߊᚒ‫␠ࠆ޿ߡ߈↢ߩޘ‬ળ
ߘߩ߽ߩ߇ᄢ߈ߊᄌൻߒߡ޿ࠆߩߢߔ߇‫ߩߘߪߦ⑳ޔ‬ༀߒᖡߒߩ್ᢿ߇ߟ߈߆ߨࠆߣߎࠈ߇޽ࠅ߹ߔ‫ޕ‬
ߥߦ߿ࠄ⼾߆ߦߪߥߞߚࠃ߁ߛ߇‫┆ޔ‬ዮߢߚ߹ࠄߥ޿‫ޔߣ߁⸒ߡ߃଀ޕ‬ᤄߪ᳓ߩਛࠍ᳇߹߹ߦᵒ޿ߢ޿
ߚ‫ࠎߛࠎߛ߇ࠇߘޕ‬᳖߇ᒛࠅߟ߼ߡ߈ߡേߊߦേߌߥߊߥߞߡ߈ߚ‫ߥࠎߘޕ‬ᕷ⧰ߒߐࠍᗵߓࠆߩߢߔ‫ޕ‬
⎇ⓥ⠪ߢ޽ࠆ߇ߚ߼ߩ⎇ⓥߦㅊࠊࠇߡ‫ޔ‬ᄞߩ޽ࠆ࠹࡯ࡑࠍ⠨߃ࠆࠁߣࠅ߇ߥߊߥߞߚߣ޿߁ߎߣߢߒࠂ
߁߆‫ޕ‬⠧ੱߩ➅ࠅ⸒ߢߒ߆ߥ޿ߩߢߎߩ⒟ᐲߦߒ߹ߔ߇‫ޔ‬ᣣᧄߩ␠ળ೙ᐲో૕߇ᚢᓟඨ਎♿૛ࠍ⚻ߡ޿
ߟߩ߹ߦ߆∋ᑷߒߡ߈ߚ‫⎇ޕ‬ⓥ⠪ߩ਎⇇߽ߘߩᤨઍ⊛⢛᥊߆ࠄ఺ࠇࠆߎߣߪߢ߈ߘ߁߽ߥ޿‫ߎߥࠎߘޕ‬
ߣࠍᗵߓߡ޿߹ߔ‫ޕ‬
⑳ߪ‫ޔ‬ችፒ⋵ਠ㑆Ꮢߦ޽ࠆᐘፉ㊁ᄖⷰኤᚲߦᆎ߹ࠅ‫࡞ࠩࡦࡎ࠾ޔ‬㊁ᄖⷰኤᣉ⸳ߩ੖ߟߩ⎇ⓥᨋ࡮ⷰኤ
ࠬ࠹࡯࡚ࠪࡦߩㆇ༡ߦ៤ࠊߞߡ߈߹ߒߚ‫߽ࠇߕ޿ޕ‬㐳ᦼ⛮⛯ⷰኤࠍ߁ߚ޿ᢥฏߦ‫␠࡞ࠨޔ‬ળߩ࠳ࠗ࠽ࡒ
࠶ࠢߥᄌേߩ⸃ᨆࠍ⋡⊛ߣߒߡ޿߹ߒߚ‫ߌߛࠇߤ߇ࠇߘޕ‬ᚑഞߒߡ޿ߚߩߛࠈ߁߆‫ࠍߣߎߥࠎߘߊࠃޕ‬
⠨߃߹ߒߚ‫ߦߢߔޕ‬㔤㐳⎇߆ࠄߪಽ㔌ߐࠇߚᣉ⸳ߢߔߩߢ⺰⹏ߪᏅߒប߃߹ߔ߇‫ߪߢߌߛࠆߌ⛯ߛߚޔ‬
ᗧ๧߇ߥ޿‫ޕ‬Ᏹߦᣂߒ޿૗߆ࠍ↢ߺ಴ߒߡ޿߆ߥ޿ߣ޿ߌߥ޿‫ࠎ߫ߜ޿߇ߎߘޕ‬㔍ߒ޿⢄ߥߩߛߣᕁ޿
߹ߔ‫⎇޿⧯ޕ‬ⓥ⠪ߚߜ߇‫੹ޔ‬ᓟߐࠄߦ⊒ዷߐߖߡ޿ߞߡߊࠇࠆߎߣࠍᦼᓙߒߡ޿߹ߔ‫ޕ‬
࠾ࡎࡦࠩ࡞㊁ᄖⷰኤᣉ⸳ߢߪ‫߽ߡߞߣߦ⑳ޔ‬ᱷᔨߥߎߣ߇޿ߊߟ߆޽ࠅ߹ߒߚ‫ޔߪߟ৻ޕ‬ਅർ⎇ⓥᨋ
߇ 1973 ᐕߦ⊒⿷ߒߡએ᧪‫ޔ‬30 ᐕㄭߊᛛቭᓙㆄ⡯ຬߣߒߡൕ߼ߡ޿ߚߛ޿ߚ⿷ᴛ⽵ᚑߐࠎߦஜᐽ਄ߩℂ
↱ߢㄉ߼ߡ޿ߚߛ߆ߑࠆࠍᓧߥ߆ߞߚߎߣ‫⚿ޔ‬ᨐߣߒߡ 60 ᱦࠍ೨ߦߒߡᄨ᛬ߔࠆߣ޿߁ਇᐘࠍ᜗޿ߡ
ߒ߹ߞߚߎߣߢߔ‫ޕ‬㊁↢࠾ࡎࡦࠩ࡞ߩⷰኤߪߤ߁ߒߡ߽ੱ㉿㔌ࠇߚᚲߢߩ‫ߩߢߌߛੱ৻ޔ‬ቅ⁛ߥ૞ᬺߦ
ߥࠅ߹ߔ‫ߥࠎߘޕ‬ⅣႺߢ㐳ߊਇ቟ቯߥ↢ᵴࠍ⛯ߌࠆߣ޿߁ߩߪ‫ޔ‬⍮ࠄߕ⍮ࠄߕߩ߁ߜߦᄢ߈ߥ⽶ᜂߦߥ
ߞߡ޿ߚߩ߆߽ߒࠇ߹ߖࠎ‫ߏޕ‬౵⑔ࠍ␨ࠆߣ౒ߦ‫⥄⑳ޔ‬り૗߽ߢ߈ߥ߆ߞߚ⥋ࠄߥߐࠍ߅⹟߮ߒ߹ߔ‫ޕ‬
߹ߚ‫ޔ‬਄ା⿧࡮ᧁᦥߩ⎇ⓥᨋߪੱᚻਇ⿷߆ࠄ߶ߣࠎߤ૗߽ߢ߈ߥ޿߹߹ߦᑄᱛߖߑࠆࠍᓧ߹ߖࠎߢߒߚ‫ޕ‬
㧙㧙
ߘߒߡ࠾ࡎࡦࠩ࡞㊁ᄖⷰኤᣉ⸳ߘߩ߽ߩ߽‫ޔ‬㊁↢േ‛⎇ⓥ࠮ࡦ࠲࡯ߩ⊒⿷ߦߣ߽ߥߞߡ⒖▤ߐࠇ‫ޔ‬㔤㐳
⎇ࠍ㔌ࠇࠆߎߣߦߥࠅ߹ߒߚ‫ޕ‬૗᡿߽ߞߣ㗎ᒛߞߡ㔤㐳⎇ౝߩ㐳ᦼ⛮⛯ⷰኤၮ࿾ࠍ቞ࠄߥ߆ߞߚߩ߆ߣ
޿߁ᛕ್ߪ‫ߡߓࠎ↞ޔ‬ฃߌߚ޿ߣᕁ޿߹ߔ‫⎇ߩࠇߙࠇߘᤨ⿷⊒ޔߪߦ⑳ޕ‬ⓥᨋߦᄞࠍ⸤ߒߚ⎇ⓥ⠪߇ߔ
ߴߡㅌቭߒߡߒ߹޿‫⎇ߦࠬ࡯ࡌࠍߎߘޔ‬ⓥߔࠆੱ߽ᓢ‫ߦޘ‬ᷫዋߒ‫ޔ‬႐วߦࠃߞߡߪ⺕߽޿ߥߊߥߞߡ޿
ߚ‫ޕ‬ᓟࠍ⛮޿ߛ৻ੱੑੱߩ⎇ⓥ⠪߇‫ߩߘޔ‬ㆇ༡ߛߌߦߢ߽ߌߞߎ߁ᚻࠍขࠄࠇߡߒ߹ߞߡ޿ࠆ‫޿߁ߘޕ‬
߁⁁ᘒߩਅߢߪ‫ޔ‬ᨒ⚵ߺࠍᄌ߃ߡߢ߽‫ޔ‬ᣂߒ޿ⷐ⚛ࠍ⚵ߺㄟࠎߢࠪࡖ࠶ࡈ࡞ߔߴ߈ߛߣ⠨߃ߚߩߢߔ‫ޕ‬
ߓࠅ⽺ࠍ఺ࠇࠆߚ߼ߦߪ‫ޔ‬૗ࠄ߆ߩ೎ߩࡊࡠࠫࠚࠢ࠻ࠍ┙ߜ਄ߍࠆߥࠅߒߡ‫ޔ‬ᰴߩࠬ࠹࠶ࡊ߳ᒁ߈਄ߍ
ࠆߣ޿߁ߎߣ߇ᄢ੐ߛߞߚߩߢߒࠂ߁߇‫ߩ߼ߚߩߘޔ‬㐳޿ദജߪ⚿ዪታࠄߥ߆ߞߚߩߢߔ‫ߊ߆߽ߦߣޕ‬
ߦ߽‫࡞ࠩࡦࡎ࠾ޔ‬㊁ᄖⷰኤᣉ⸳ߩᱧผ߆ࠄߪ‫ޔ‬ᣣᧄߩ㔤㐳㘃ቇో૕ߩᄌㆫࠍᒝߊᗧ⼂ߐߖࠄࠇߡ޿߹ߔ‫ޕ‬
ߎ߁ߒߚᣉ⸳ㆇ༡ߦ߆߆ࠊࠆ৻ᣇߢ‫ޔࠕࠪࡀ࠼ࡦࠗߪ⑳ޔ‬ਛ࿖‫ߢߤߥࠗ࠲ޔ‬ᶏᄖ⺞ᩏࠍ⛯ߌߡ߈߹ߒ
ߚ‫ޕ‬ᢙ߃ߡߺ߹ߔߣ‫ޔ‬ᄢቇ㒮ߩ㗃߆ࠄ߶ߣࠎߤᲤᐕ‫ޔ‬ᑧߴ 77 ࿁‫ߩࠄࠇߎޔ‬࿖ߢߩ⺞ᩏࠍⴕߞߡ߈ߚߎ
ߣߦߥࠅ߹ߔ‫ᦨޕ‬ೋߩᶏᄖ⺞ᩏߩ೨ߦ‫ޔ‬૗ੱ߆ߩవ↢߆ࠄ⸒ࠊࠇ߹ߒߚ‫ޕ‬ᶏᄖߢߩ⺞ᩏ⎇ⓥߪᦨೋߩᤨ
ߩ⚻㛎߇ᄢ੐ߢ‫ޕࠆߥߊߚ߈ⴕߚ߹ߣߊ޿ߊ߹߁߇ࠇߘޔ‬ㄆ޿‫ߣ޿ߒ⧰ޔ‬ᕁ߁ߣዥㄟߺߔࠆࠃ߁ߦߥࠆ‫ޔ‬
ߣ޿߁ߎߣߢߒߚ‫ߩߘߦ߆⏕ޕ‬ㅢࠅߢ‫ߩ⑳ޔ‬႐วߪ⇣ᢥൻ㧔ߚߛߒࠕࠫࠕߩ㒢ቯߐࠇߚ࿾ၞߢߒ߆޽ࠅ
߹ߖࠎ߇㧕ߩਛߢ᥵ࠄߔᭉߒߐ‫ޔ‬㕙⊕ߐߦ‫ࠅ߼ߩࠅ߆ߞߔޔ‬ㄟࠎߢߒ߹ߞߚࠃ߁ߢߔ‫⺞ࠇߙࠇߘޕ‬ᩏ⎇
ⓥߩኻ⽎ࠍߤ߁ᢱℂߔࠆ߆ߣ޿߁ߎߣߪᒰὼ޿ߩ৻⇟ߦ⠨߃ࠆߴ߈ߎߣߥߩߢߔ߇‫⺞ޔ‬ᩏ࿾ߢ಴ળߞߚ
ੱߚߜ߇ߤ߁޿߁ੱ↢ࠍㅍߞߡ޿ߊߩߛࠈ߁߆‫߇ߣߎߥࠎߘޔ‬⍮ࠅߚߊߡㅢ޿ߟ߼ߡ޿ߚ㕙߽޽ࠅ߹ߒ
ߚ‫ߩࠄࠇߎޕ‬࿖ߦ૑߻ᄙߊߩੱߚߜߣኅᣖหὼߩઃ߈ว޿߇ߢ߈ߚߣ޿߁ߩߪ‫੹ޔ‬ᕁ߁ߛߌߢ߽ߚ޿߳
ࠎሜߒ޿ߎߣߢߔ‫ޕ‬
ㅌቭߒߡ⪭ߜ⌕޿ߡ⠨߃ߡߺࠆߣ‫ߦߣߎߥࠎࠈ޿ޔ‬ᚻࠍ಴ߒߡ‫ࠆ߼ߣ߹ޔ‬ᣇߪਇචಽߥ߹߹ߦ‫޿ߕޔ‬
߱ࠎᢔẂߦㆊߏߒߡ߈ߚࠃ߁ߦᕁ޿߹ߔ‫ߚ޿ߛߚ޿ߡߒ⸵ࠍࠇߘޕ‬ᄙߊߩవ↢ᣇ߿੐ോࠬ࠲࠶ࡈߩᣇ‫ޔޘ‬
ห௥⻉᳁‫ޔ‬ቇ↢⻉ำߦߪ‫ޔ‬ෘߊᗵ⻢ߒ߹ߔ‫޿⧯ޕ‬㗃ߪ‫ޔ‬ቯᐕㅌቭߥߤߣ޿߁ߩߪ⠨߃ߚߎߣ߽޽ࠅ߹ߖ
ࠎߢߒߚ‫⃻ޕ‬ઍ ߩᆝᝥߡጊߺߚ޿ߥ߽ߩ߆ߣᕁߞߡ޿߹ߒߚ߇‫ ↢ੱߪߢ੹ޔ‬80 ᐕߩᤨઍߢߔ‫߁ߘޕ‬
㐳޿ᦼ㑆߇ᱷߐࠇߡ޿ࠆࠊߌߢߪ޽ࠅ߹ߖࠎ߇‫ߩߎޔ‬వ‫ޔ‬⠧޿వࠍߤ߁ᭉߒ߽߁߆‫ޔ‬⠨߃޽ߋߨߡ߅ࠅ
߹ߔ‫ޕ‬
㧙㧙
2012 ᐕ 10 ᦬ 4 ᣣ
⊒ⴕ⠪
੩ㇺᄢቇ㔤㐳㘃⎇ⓥᚲ
‫ޥ‬484-8506 ᗲ⍮⋵›ጊᏒቭᨋ 41 ⇟࿾ߩ 2
http://www.pri.kyoto-u.ac.jp
[email protected]
✬㓸 ⥄Ꮖὐᬌ࡮⹏ଔᆔຬળ
㜟੗ᱜᚑ㧘㜟↰᣽ᒾ㧘Michael A Huffman㧘
ᱜ㜞ା↵㧘ች࿾㊀ᒄ㧘ᣧᎹᢅਯ
✬㓸ᜂᒰ㧦ᣧᎹᢅਯ㧘೨↰ዊᑿ
ශ೚ᚲ
ᩣᑼળ␠ࠦ࡯ࡓ࡜
http://www.kohmura.co.jp
‫ޥ‬501-2517 ጘ㒂⋵ጘ㒂Ꮢਃベ߲ࠅࠎߣ߯޽ 3
TEL㧦058-229-5858
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